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<document id="23CB8611165CD7C0D639D636F199B2AD" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657307DFF95FF8EFCD9BC8D0B62" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Brachypelus ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="206" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="205" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="430F8A16272F1870FA91CF2DCAA0BCD4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E008233179CE267B4082DE1FE2927DBF">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="37E3654D96648442B6AEC300240ADA63" type="personal">
<mods:role id="6563E264F1869E530A8DB3C105032451">
<mods:roleTerm id="BB2315C517F2B151AB408D4CE91F9072">Author</mods:roleTerm>
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<mods:namePart id="DFFB82C4AA2C8D07039CDFB6BB0634E4">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5AFDB64A447608CFB08271ED969F7027">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
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<mods:roleTerm id="2E38BDF08B24EFBA5769F25B3650975F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0AEFC71EDCA4AF73F109D2B9C6BAAF53">Rainio, Johanna</mods:namePart>
<mods:affiliation id="67077618F0DB9EBA1C6BE81B04A45AF4">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="AFD45381A175B6F086FAAD63C011C2ED">text</mods:typeOfResource>
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<mods:title id="D3B6C3DB8F5D2C9B716D172CCBF9E0FF">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="17DC1FA98D919142B3C08CC32C55EF82">
<mods:date id="EC35960238C50C7664DBDAC9F6591266">2016</mods:date>
<mods:detail id="FEB598AF1F29F31EEB2F7DD557A59F77" type="pubDate">
<mods:number id="53A472C1B88A468FEF955A180E42197A">2016-04-29</mods:number>
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<mods:detail id="7B4F9A502913D6832395184575C1610C" type="volume">
<mods:number id="A85447DE447B6131E342F3D2A8910C8B">63</mods:number>
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<mods:number id="8813A52A46D0FA5435D7A11292D7F18E">7</mods:number>
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<mods:classification id="F76D8B0D11EB52A64FC121792D20D4E1">journal article</mods:classification>
<mods:identifier id="7924A63BD6B2341D9D6DC47ECE310D8F" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="61AA03B8DA0C0C04145ADD5F30939FB2" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="554750F639B327618E53A2F21FB7EC20" type="Zenodo-Dep">13799439</mods:identifier>
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<treatment id="0A3E2657307DFF95FF8EFCD9BC8D0B62" LSID="urn:lsid:plazi:treatment:0A3E2657307DFF95FF8EFCD9BC8D0B62" httpUri="http://treatment.plazi.org/id/0A3E2657307DFF95FF8EFCD9BC8D0B62" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<subSubSection id="CA8DC4CA307DFF94FF8EFCD9BE760E7F" box="[123,789,843,868]" pageId="4" pageNumber="205" type="nomenclature">
<paragraph id="82289741307DFF94FF8EFCD9BE760E7F" blockId="4.[123,789,843,898]" box="[123,789,843,868]" pageId="4" pageNumber="205">
<heading id="D960202D307DFF94FF8EFCD9BE760E7F" box="[123,789,843,868]" fontSize="10" level="2" pageId="4" pageNumber="205" reason="4">
<emphasis id="B0E34B53307DFF94FF8EFCD9BE760E7F" bold="true" box="[123,789,843,868]" pageId="4" pageNumber="205">
<taxonomicName id="4597ECC2307DFF94FF8EFCD9BFD20E78" ID-CoL="MYCC" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,689,843,868]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53307DFF94FF8EFCD9BCC30E78" bold="true" box="[123,416,843,867]" italics="true" pageId="4" pageNumber="205">Brachypelus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628307DFF94FD4BFCDEBE760E7F" box="[702,789,844,868]" pageId="4" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA307DFF94FF8EFCFEBFA40EB0" pageId="4" pageNumber="205" type="description">
<paragraph id="82289741307DFF94FF8EFCFEBD8E0E9A" blockId="4.[123,789,843,898]" box="[123,237,876,898]" pageId="4" pageNumber="205">
<figureCitation id="1AAC8BC4307DFF94FF8EFCFEBD8E0E9A" box="[123,237,876,898]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStartId-0="5.[155,167,1152,1171]" captionStartId-1="6.[155,167,1136,1155]" captionTargetBox-0="[133,1174,245,1135]" captionTargetBox-1="[306,980,229,1110]" captionTargetId-0="figure-997@5.[120,1174,216,1150]" captionTargetId-1="figure-647@6.[306,980,215,1110]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." captionText-1="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13799443" figureDoi-1="http://doi.org/10.5281/zenodo.13799447" httpUri-0="https://zenodo.org/record/13799443/files/figure.png" httpUri-1="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Figures 2–3</figureCitation>
</paragraph>
<paragraph id="82289741307DFF94FF8EFC07BFA40EB0" blockId="4.[123,711,917,939]" box="[123,711,917,939]" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF8EFC07BC5A0EB1" box="[123,313,917,939]" italics="true" pageId="4" pageNumber="205">
<taxonomicName id="4597ECC2307DFF94FF8EFC07BD900EB1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,243,917,938]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus" status="n. sp.">Brachypelus</taxonomicName>
<taxonomicNameLabel id="ABD0F628307DFF94FF0CFC04BC4E0EB0" box="[249,301,918,939]" pageId="4" pageNumber="205" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 31 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA307DFF95FF5EFC2DBC8D0B62" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205" type="materials_examined">
<paragraph id="82289741307DFF94FF5EFC2DBF06092C" blockId="4.[123,1174,958,1726]" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF5EFC2DBC3E0ECE" bold="true" box="[171,349,959,983]" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFC2DBD870ECE" baselines="977,976" box="[171,228,959,983]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="4" pageNumber="205">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FF05FC50BC3E0ECE" baselines="976" box="[240,349,962,981]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="4" pageNumber="205">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C307DFF94FE71FC2CBF80090C" collectingDate="2001-01-15" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="130" location="MADAGASCAR" pageId="4" pageNumber="205" specimenCount="130" typeStatus="holotype">
<typeStatus id="5D2C29E3307DFF94FE71FC2CBC840ECD" box="[388,487,958,982]" pageId="4" pageNumber="205" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4307DFF94FE0EFC2CBF180ECD" box="[507,635,958,983]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="4" pageNumber="205">Figs. 2A–B</figureCitation>
), a female, in CAS, labeled: “
<collectingCountry id="FA80D7D1307DFF94FC2DFC2DB9F50ECC" box="[984,1174,959,983]" name="Madagascar" pageId="4" pageNumber="205">MADAGASCAR</collectingCountry>
Ranomafana NP. Talatakely, X,
<specimenCount id="94915CC8307DFF94FE39FC4DBF760EED" box="[460,533,991,1015]" count="130" pageId="4" pageNumber="205" type="generic">
<quantity id="456F3AA4307DFF94FE39FC4DBF6C0EED" box="[460,527,991,1015]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="205" unit="m" value="130.0">
<elevation id="09BA7072307DFF94FE39FC4DBF6C0EED" box="[460,527,991,1015]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="205" unit="m" value="130.0">130 m</elevation>
</quantity>
.
</specimenCount>
<date id="F629B181307DFF94FDEEFC4DBFE60EEC" box="[539,645,991,1015]" pageId="4" pageNumber="205" value="2001-01-15">
<collectingDate id="E66D4869307DFF94FDEEFC4DBFE60EEC" box="[539,645,991,1015]" pageId="4" pageNumber="205" value="2001-01-15">15.1.2001</collectingDate>
</date>
”/ “
<collectingMethod id="5BD6EF56307DFF94FD5FFC4DBEF90EED" box="[682,922,991,1015]" pageId="4" pageNumber="205">collected in pitfall trap</collectingMethod>
on forest floor,
<collectorName id="2F62F297307DFF94FBB7FC4DBDC2090C" pageId="4" pageNumber="205">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F297307DFF94FF3DFB92BC4A090C" box="[200,297,1023,1047]" pageId="4" pageNumber="205">J. Rainio</collectorName>
collectors”/ “32.” [handwritten label]/ “
</materialsCitation>
<typeStatus id="5D2C29E3307DFF94FD16FC6DBE13090C" box="[739,880,1023,1047]" pageId="4" pageNumber="205" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2307DFF94FC8CFC6DBC3C092C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53307DFF94FC8CFC6DB9F6090C" box="[889,1173,1023,1047]" italics="true" pageId="4" pageNumber="205">Brachypelus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307DFF94FE93FBB2BCC3092C" box="[358,416,1056,1079]" pageId="4" pageNumber="205" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</paragraph>
<paragraph id="82289741307DFF94FF5EFBD2B950094C" blockId="4.[123,1174,958,1726]" box="[171,1075,1088,1112]" pageId="4" pageNumber="205">
<materialsCitation id="32FF9D1C307DFF94FF5EFBD2B94F0943" box="[171,1068,1088,1112]" country="Madagascar" location="Ranomafana National Park" pageId="4" pageNumber="205" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53307DFF94FF5EFBD2BC34094D" bold="true" box="[171,343,1088,1112]" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFBD2BD87094D" baselines="1106,1105" box="[171,228,1088,1112]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="4" pageNumber="205">
<typeStatus id="5D2C29E3307DFF94FF5EFBD2BD87094D" box="[171,228,1088,1112]" pageId="4" pageNumber="205">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FF19FBD1BC34094D" baselines="1105" box="[236,343,1091,1110]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="4" pageNumber="205">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1307DFF94FE8BFBD2BC9C0943" box="[382,511,1088,1112]" name="Madagascar" pageId="4" pageNumber="205">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3307DFF94FDF8FBD2BF9F0943" box="[525,764,1088,1112]" country="Madagascar" name="Fianarantsoa" pageId="4" pageNumber="205">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A307DFF94FCFFFBD2B94F0943" LSID="urn:lsid:plazi:treatment:0A3E2657307DFF95FF8EFCD9BC8D0B62:8748C19A307DFF94FCFFFBD2B94F0943" box="[778,1068,1088,1112]" country="Madagascar" name="Ranomafana National Park" pageId="4" pageNumber="205" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741307DFF94FF5EFBF2BECA0983" blockId="4.[123,1174,958,1726]" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF5EFBF2BC91096D" bold="true" box="[171,498,1120,1144]" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFBF2BC4D096D" baselines="1138,1137" box="[171,302,1120,1144]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="4" pageNumber="205">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FEC3FBF1BC31096D" baselines="1137" box="[310,338,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="4" pageNumber="205">OF</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FEACFBF1BCCF096D" baselines="1137" box="[345,428,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="4" pageNumber="205">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D307DFF94FE40FBF1BC91096D" baselines="1137" box="[437,498,1123,1142]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="4" pageNumber="205">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2307DFF94FD0CFBF2BEE40963" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1120,1144]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307DFF94FD0CFBF2BEE40963" box="[761,903,1120,1144]" italics="true" pageId="4" pageNumber="205">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E3307DFF94FD29FB13BE690982" box="[732,778,1153,1177]" pageId="4" pageNumber="205">type</typeStatus>
was collected.
</paragraph>
<paragraph id="82289741307DFF95FF5EFB33BC8008B4" blockId="4.[123,1174,958,1726]" lastBlockId="5.[123,1174,1229,1658]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
<smallCapsWord id="84CE019D307DFF94FF5EFB33BC2109AC" baselines="1203,1202" box="[171,322,1185,1209]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="4" pageNumber="205">
<emphasis id="B0E34B53307DFF94FF5EFB33BC2109AC" bold="true" box="[171,322,1185,1209]" pageId="4" pageNumber="205">RECOGNITION</emphasis>
</smallCapsWord>
.— Large for genus, SBL =
<quantity id="456F3AA4307DFF94FD99FB33BFDE09A3" box="[620,701,1185,1209]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="4" pageNumber="205" unit="mm" value="6.8">6.8 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4307DFF94FC2FFB33B94809A2" box="[986,1067,1185,1209]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="4" pageNumber="205">Fig. 2A</figureCitation>
) are most similar to those of
<taxonomicName id="4597ECC2307DFF94FEB2FB53BFA509C2" authority="Putzeys (1866)" authorityName="Putzeys" authorityYear="1866" box="[327,710,1217,1241]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FEB2FB53BF7E09C2" box="[327,541,1217,1241]" italics="true" pageId="4" pageNumber="205">Brachypelus obesus</emphasis>
<bibRefCitation id="E606EAB0307DFF94FDD1FB53BFA509C2" author="PUTZEYS, J. A. A. H." box="[548,710,1217,1241]" pageId="4" pageNumber="205" pagination="1 - 242" refId="ref32024" refString="PUTZEYS, J. A. A. H. 1866. Revision generale des clivinides. Annales de la Societe Entomologique de Belgique 10: 1 - 242." type="journal article" year="1866">Putzeys (1866)</bibRefCitation>
</taxonomicName>
, with which they share large size, only six setae on the labrum, and a single discal setiferous pore on elytral interval 3 (at apical one-fifth). They differ from
<taxonomicName id="4597ECC2307DFF94FEC1FA90BCF80802" authorityName="Putzeys" authorityYear="1866" box="[308,411,1281,1305]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FEC1FA90BCF80802" box="[308,411,1281,1305]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members in the following: transverse fronto-clypeal suture present and deeply impressed (
<figureCitation id="1AAC8BC4307DFF94FEB3FAB0BCFB0821" box="[326,408,1314,1338]" captionStart="FIGURE" captionStartId="6.[155,167,1136,1155]" captionTargetBox="[306,980,229,1110]" captionTargetId="figure-647@6.[306,980,215,1110]" captionTargetPageId="6" captionText="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799447" httpUri="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Fig. 3A</figureCitation>
) (absent from
<taxonomicName id="4597ECC2307DFF94FDCCFAB1BFC30821" authorityName="Putzeys" authorityYear="1866" box="[569,672,1314,1338]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FDCCFAB1BFC30821" box="[569,672,1314,1338]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); frons with longitudinal frontal furrows narrower and shallower than in
<taxonomicName id="4597ECC2307DFF94FDF6FAD1BF0A0841" authorityName="Putzeys" authorityYear="1866" box="[515,617,1346,1370]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FDF6FAD1BF0A0841" box="[515,617,1346,1370]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
; posterior lateral margins of the eyes broadly round- ed (more narrowly and angularly rounded in
<taxonomicName id="4597ECC2307DFF94FD9DFAF1BFB20861" authorityName="Putzeys" authorityYear="1866" box="[616,721,1378,1402]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FD9DFAF1BFB20861" box="[616,721,1378,1402]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); elytral base (
<figureCitation id="1AAC8BC4307DFF94FC17FAF0B9570861" box="[994,1076,1378,1403]" captionStart="FIGURE" captionStartId="6.[155,167,1136,1155]" captionTargetBox="[306,980,229,1110]" captionTargetId="figure-647@6.[306,980,215,1110]" captionTargetPageId="6" captionText="FIGURE 3. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Head, dorsal aspect; B. Elytral base, dorsal aspect. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799447" httpUri="https://zenodo.org/record/13799447/files/figure.png" pageId="4" pageNumber="205">Fig. 3B</figureCitation>
) straight between intervals 3 and obliquely sloped laterally from interval 4 (elytral base straight between intervals 4 and gradually rounded laterally from interval
<quantity id="456F3AA4307DFF94FD2BFA31BE6B08A0" box="[734,776,1443,1467]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="205" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4597ECC2307DFF94FCFAFA36BE1508A0" authorityName="Putzeys" authorityYear="1866" box="[783,886,1443,1467]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FCFAFA36BE1508A0" box="[783,886,1443,1467]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); elytra with lateral margin distinctly toothed at humerus (smoothly rounded at humerus, without a tooth in
<taxonomicName id="4597ECC2307DFF94FBBCFA56BDFE08E0" authorityName="Putzeys" authorityYear="1866" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FBBCFA56BDFE08E0" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members); and elytral intervals slightly convex apically (nearly flat in
<taxonomicName id="4597ECC2307DFF94FC47FA76B97808E7" authorityName="Putzeys" authorityYear="1866" box="[946,1051,1508,1532]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B0E34B53307DFF94FC47FA76B97808E7" box="[946,1051,1508,1532]" italics="true" pageId="4" pageNumber="205">B. obesus</emphasis>
</taxonomicName>
members). Others species with members of comparably large size include
<taxonomicName id="4597ECC2307DFF94FCEAF996BDA10B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FCEAF996BE9A0B07" box="[799,1017,1540,1564]" italics="true" pageId="4" pageNumber="205">Brachypelus rolandi</emphasis>
<bibRefCitation id="E606EAB0307DFF94FBF5F996BDA10B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2307DFF94FF3AF9B6BFF40B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" box="[207,663,1572,1596]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="betsileo">
<emphasis id="B0E34B53307DFF94FF3AF9B6BCCD0B27" box="[207,430,1572,1596]" italics="true" pageId="4" pageNumber="205">Brachypelus betsileo</emphasis>
<bibRefCitation id="E606EAB0307DFF94FE41F9B6BFF40B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" box="[436,663,1572,1596]" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2307DFF94FD51F9B6B90B0B27" authority="Bulirsch et al. (2005)" authorityName="Bulirsch" authorityYear="2005" box="[676,1128,1572,1596]" class="Insecta" family="Carabidae" genus="Brachypelas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="B0E34B53307DFF94FD51F9B6BEE30B27" box="[676,896,1572,1596]" italics="true" pageId="4" pageNumber="205">Brachypelas fischeri</emphasis>
<bibRefCitation id="E606EAB0307DFF94FC73F9B6B90B0B27" author="BULIRSCH, P. &amp; J. JANAK &amp; P. MORAVEC" box="[902,1128,1572,1596]" pageId="4" pageNumber="205" pagination="1 - 35" refId="ref29568" refString="BULIRSCH, P., J. JANAK, AND P. MORAVEC. 2005. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar. Studies and Reports of District Museum Prague-East, Taxonomical Series 1: 1 - 35." type="book chapter" year="2005">Bulirsch et al. (2005)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4597ECC2307DFF94FF8EF9D6BFE90B46" authority="Bulirsch and Moravec (2009)" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[123,650,1604,1629]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307DFF94FF8EF9D6BC290B47" box="[123,330,1604,1628]" italics="true" pageId="4" pageNumber="205">Brachypelus janaki</emphasis>
<bibRefCitation id="E606EAB0307DFF94FEA5F9D6BFE90B46" author="BULIRSCH, P. &amp; P. MORAVEC" box="[336,650,1604,1629]" pageId="4" pageNumber="205" pagination="27 - 36" refId="ref29614" refString="BULIRSCH, P., AND P. MORAVEC. 2009. New species and findings of Scaritinae (Coleoptera: Carabidae) from Madagascar: Part 2. Studies and Reports of District Museum Prague-East, Taxonomical Series 5: 27 - 36." type="book chapter" year="2009">Bulirsch and Moravec (2009)</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="4597ECC2307DFF94FCF0F9D7BED20B46" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[773,945,1605,1629]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307DFF94FCF0F9D7BED20B46" box="[773,945,1605,1629]" italics="true" pageId="4" pageNumber="205">B. ranomafanae</emphasis>
</taxonomicName>
members differ from those of
<taxonomicName id="4597ECC2307DFF94FF2DF9F4BC270B66" box="[216,324,1637,1661]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FF2DF9F4BC270B66" box="[216,324,1637,1661]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
in having the labrum with only six setae (seven in
<taxonomicName id="4597ECC2307DFF94FC98F9F4BEBA0B66" box="[877,985,1637,1661]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FC98F9F4BEBA0B66" box="[877,985,1637,1661]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members), elytra widest at middle (anterior to middle in
<taxonomicName id="4597ECC2307DFF94FDEAF914BFE80B86" box="[543,651,1669,1693]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FDEAF914BFE80B86" box="[543,651,1669,1693]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members), and elytral intervals moderately convex throughout (flattened apical in
<taxonomicName id="4597ECC2307DFF94FE0FF934BF040BA6" box="[506,615,1701,1725]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="rolandi">
<emphasis id="B0E34B53307DFF94FE0FF934BF040BA6" box="[506,615,1701,1725]" italics="true" pageId="4" pageNumber="205">B. rolandi</emphasis>
</taxonomicName>
members). Members of both
<taxonomicName id="4597ECC2307DFF94FC5EF934B97E0BA6" box="[939,1053,1701,1725]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="betsileo">
<emphasis id="B0E34B53307DFF94FC5EF934B97E0BA6" box="[939,1053,1701,1725]" italics="true" pageId="4" pageNumber="205">B. betsileo</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC2307DFF95FBA1F934BDD109FE" class="Insecta" family="Carabidae" genus="Brachypelas" kingdom="Animalia" lastPageId="5" lastPageNumber="206" order="Coleoptera" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="fischeri">
<emphasis id="B0E34B53307DFF95FBA1F934BDD109FE" italics="true" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">B. fischeri</emphasis>
</taxonomicName>
have seven labral setae (whereas those of
<taxonomicName id="4597ECC2307CFF95FD85FB5CBE7809FE" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[624,795,1229,1253]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FD85FB5CBE7809FE" box="[624,795,1229,1253]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
have only six) and lack discal setiferous pores (those of
<taxonomicName id="4597ECC2307CFF95FE94FB62BF6E081C" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[353,525,1263,1287]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FE94FB62BF6E081C" box="[353,525,1263,1287]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
have a single setiferous pore in the apical one-fifth on interval 3. Finally,
<taxonomicName id="4597ECC2307CFF95FEE3FA83BCA10832" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[278,450,1297,1321]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53307CFF95FEE3FA83BCA10832" box="[278,450,1297,1321]" italics="true" pageId="5" pageNumber="206">B. ranomafanae</emphasis>
</taxonomicName>
members differ from those of
<taxonomicName id="4597ECC2307CFF95FCF8FA83BE0D0832" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[781,878,1297,1321]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FCF8FA83BE0D0832" box="[781,878,1297,1321]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
in having only six setae on the labrum (seven labral setae in
<taxonomicName id="4597ECC2307CFF95FE1CFAA1BF2F0851" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[489,588,1330,1354]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FE1CFAA1BF2F0851" box="[489,588,1330,1354]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
members), the elytral margin with a distinct humeral tooth (tooth absent from
<taxonomicName id="4597ECC2307CFF95FE7BFAC7BC910877" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[398,498,1364,1388]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FE7BFAC7BC910877" box="[398,498,1364,1388]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
members), and only a single discal setiferous pore on elytral interval 3, with intervals 5 and 7 without such punctures (five or more such punctures on intervals 3, 5 and
<quantity id="456F3AA4307CFF95FF2DFA0ABC6108B4" box="[216,258,1431,1455]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="5" pageNumber="206" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4597ECC2307CFF95FEFCFA0ABC0908B4" authorityName="Bulirsch and Moravec" authorityYear="2009" box="[265,362,1431,1455]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="janaki">
<emphasis id="B0E34B53307CFF95FEFCFA0ABC0908B4" box="[265,362,1431,1455]" italics="true" pageId="5" pageNumber="206">B. janaki</emphasis>
</taxonomicName>
members).
</paragraph>
<caption id="D6E8C7C9307CFF95FF6EFB12BF6609B6" ID-DOI="http://doi.org/10.5281/zenodo.13799443" ID-Zenodo-Dep="13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="5" pageNumber="206" startId="5.[155,167,1152,1171]" targetBox="[133,1174,245,1135]" targetPageId="5" targetType="figure">
<paragraph id="82289741307CFF95FF6EFB12BF6609B6" blockId="5.[123,1171,1152,1197]" pageId="5" pageNumber="206">
<smallCapsWord id="84CE019D307CFF95FF6EFB12BDBB0989" baselines="1166,1166" box="[155,216,1152,1171]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="5" pageNumber="206">FIGURE</smallCapsWord>
2. Digital images of holotype female of
<taxonomicName id="4597ECC2307CFF95FDCBFB12BE820988" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[574,993,1152,1171]" class="Insecta" family="Carabidae" genus="Brachypelus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53307CFF95FDCBFB12BE410988" box="[574,802,1152,1171]" italics="true" pageId="5" pageNumber="206">Brachypelus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628307CFF95FC12FB12B9490988" box="[999,1066,1152,1171]" pageId="5" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741307CFF95FF5EFA2BB96908CA" blockId="5.[123,1174,1229,1658]" box="[171,1034,1465,1490]" pageId="5" pageNumber="206">
<emphasis id="B0E34B53307CFF95FF5EFA2BBC9208CB" bold="true" box="[171,497,1465,1489]" pageId="5" pageNumber="206">
<smallCapsWord id="84CE019D307CFF95FF5EFA2BBC3508CB" baselines="1483,1483" box="[171,342,1465,1489]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="5" pageNumber="206">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D307CFF95FEA8FA2FBC9208CB" baselines="1483" box="[349,497,1469,1488]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="5" pageNumber="206">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3307CFF95FC8AFA28BECE08C9" box="[895,941,1466,1490]" pageId="5" pageNumber="206">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741307CFF95FF5EFA49BC8D0B62" blockId="5.[123,1174,1229,1658]" pageId="5" pageNumber="206">
<emphasis id="B0E34B53307CFF95FF5EFA49BCC608EA" bold="true" box="[171,421,1499,1523]" pageId="5" pageNumber="206">
<smallCapsWord id="84CE019D307CFF95FF5EFA49BC6A08EA" baselines="1517,1516" box="[171,265,1499,1523]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="5" pageNumber="206">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D307CFF95FEE4FA4CBCC608EA" baselines="1516" box="[273,421,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="5" pageNumber="206">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E3307CFF95FDA4FA49BFCD08E8" box="[593,686,1499,1523]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
was collected in a pitfall trap placed in secondary montane rain forest in the Talatakely area, at an elevation of
<quantity id="456F3AA4307CFF95FCADFA6FBEFF0B0F" box="[856,924,1533,1557]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="206" unit="m" value="900.0">900 m</quantity>
, at the
<quantity id="456F3AA4307CFF95FC19FA6FB93B0B0F" box="[1004,1112,1533,1557]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="206" unit="m" value="130.0">130 meter</quantity>
mark along Trail “X”. Vegetation in that area consisted of bamboo, small young trees and some larger trees with diameters at breast-height (DBH) of
<quantity id="456F3AA4307CFF95FD88F9D2BF8A0B43" box="[637,745,1600,1624]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="206" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 cm</quantity>
. Tree trunks and branches were mostly covered with epiphytes and lianas.
</paragraph>
</subSubSection>
</treatment>
</document>
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