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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="13132970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E782FFFECFF013A44AB17E839" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Flagellobolus Wesener &amp; Enghoff &amp; Sierwald 2009, gen. n." docType="treatment" docUuid="30252D40-A093-48F8-9756-96A56C848B70" docUuidSource="ZooBank" docVersion="5" lastPageNumber="43" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="41" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791409" ID-GBIF-Taxon="164184225" ID-Zenodo-Dep="3791409" LSID="urn:lsid:zoobank.org:act:30252D40-A093-48F8-9756-96A56C848B70" httpUri="http://treatment.plazi.org/id/1772122E782FFFECFF013A44AB17E839" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<subSubSection box="[140,496,1328,1355]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph blockId="40.[140,953,1328,1390]" box="[140,496,1328,1355]" pageId="40" pageNumber="41">
<heading box="[140,496,1328,1355]" fontSize="11" level="2" pageId="40" pageNumber="41" reason="3">
<emphasis bold="true" box="[140,496,1328,1355]" pageId="40" pageNumber="41">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,403,1328,1355]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis bold="true" box="[140,295,1328,1355]" italics="true" pageId="40" pageNumber="41">Flagellobolus</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[415,496,1328,1354]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,953,1363,1390]" pageId="40" pageNumber="41" type="description">
<paragraph blockId="40.[140,953,1328,1390]" box="[140,953,1363,1390]" pageId="40" pageNumber="41">
<uri box="[140,953,1363,1390]" pageId="40" pageNumber="41">
urn:lsid:zoobank.org:act:
<uuid box="[420,953,1363,1390]" pageId="40" pageNumber="41">30252D40-A093-48F8-9756-96A56C848B70</uuid>
</uri>
</paragraph>
</subSubSection>
<subSection lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="multiple">
<subSubSection box="[140,745,1433,1460]" pageId="40" pageNumber="41" type="type_taxon">
<paragraph blockId="40.[140,745,1433,1460]" box="[140,745,1433,1460]" pageId="40" pageNumber="41">
<emphasis bold="true" box="[140,287,1433,1460]" pageId="40" pageNumber="41">
<typeStatus box="[140,198,1434,1460]" pageId="40" pageNumber="41">Type</typeStatus>
species
</emphasis>
:
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[302,539,1434,1460]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pauliani" status="sp. n.">
<emphasis box="[302,539,1434,1460]" italics="true" pageId="40" pageNumber="41">Flagellobolus pauliani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[546,610,1434,1460]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
, monotypic
</paragraph>
</subSubSection>
<subSubSection lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="diagnosis">
<paragraph blockId="40.[140,1108,1504,1672]" lastBlockId="42.[140,1108,167,616]" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
<emphasis bold="true" box="[140,259,1504,1530]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Diagnosis</emphasis>
: anterior and posterior gonopods of unique shape (
<figureCitation box="[849,996,1504,1531]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="40" pageNumber="41">Figs 23C, D</figureCitation>
): anterior gonopod with a telopodite process, retrorse part slender, located at midpoint of process instead of apically, a feature not shared with any other genus from
<collectingCountry box="[975,1108,1575,1601]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
(
<figureCitation box="[148,260,1610,1636]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="40" pageNumber="41">Fig. 23D</figureCitation>
). Most unusual feature of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[578,716,1610,1636]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[578,716,1610,1636]" italics="true" pageId="40" pageNumber="41">Flagellobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[728,809,1610,1636]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
is the unique elongation of the posterior gonopod telopodite into a simple, strongly elongated, thin tip (
<figureCitation captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">Figs 23E, F</figureCitation>
). This genus shares a small body size and elongated antennae with
<taxonomicName authorityName="de Saussure &amp; Zehntner" authorityYear="1901" box="[970,1100,167,193]" class="Diplopoda" family="Pachybolidae" genus="Spiromimus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[970,1100,167,193]" italics="true" pageId="42" pageNumber="43">Spiromimus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,306,202,228]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[140,306,202,228]" italics="true" pageId="42" pageNumber="43">Zehntnerobolus</emphasis>
</taxonomicName>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[323,457,202,228]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[323,457,202,228]" italics="true" pageId="42" pageNumber="43">Alluviobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[465,544,202,228]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
and
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[605,732,202,228]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[605,732,202,228]" italics="true" pageId="42" pageNumber="43">Ostinobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[742,821,202,228]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
Gnathochilarium with a subdivided mentum and a single sclerotized ledge on each stipites, like in
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[964,1107,237,263]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[964,1107,237,263]" italics="true" pageId="42" pageNumber="43">Riotintobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,216,273,299]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[230,423,272,298]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[230,423,272,298]" italics="true" pageId="42" pageNumber="43">Pseudocentrobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[429,505,273,299]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[519,659,272,298]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[519,659,272,298]" italics="true" pageId="42" pageNumber="43">Granitobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[666,742,273,299]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[756,878,272,298]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[756,878,272,298]" italics="true" pageId="42" pageNumber="43">Caprobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[884,960,273,299]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[974,1108,272,298]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[974,1108,272,298]" italics="true" pageId="42" pageNumber="43">Alluviobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,219,308,334]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
,
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[235,362,307,333]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[235,362,307,333]" italics="true" pageId="42" pageNumber="43">Ostinobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[371,451,307,334]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
Although the posterior gonopods of
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[881,1019,307,333]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" status="gen. n.">
<emphasis box="[881,1019,307,333]" italics="true" pageId="42" pageNumber="43">Flagellobolus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1028,1107,308,334]" pageId="42" pageNumber="43" rank="genus">gen. n.</taxonomicNameLabel>
are connected by a large sternite, the telopodites do more closely resemble those of species of the
<taxonomicName authorityName="Cook" authorityYear="1895" box="[301,443,378,404]" order="null" pageId="42" pageNumber="125" rank="subOrder" subOrder="Spirobolidea">Spirobolidea</taxonomicName>
families
<taxonomicName box="[547,715,378,404]" class="Diplopoda" family="Spirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Spirobolellidae</taxonomicName>
and
<taxonomicName authorityName="Brolemann" authorityYear="1913" box="[775,1019,378,404]" class="Diplopoda" family="Pseudospirobolellidae" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Pseudospirobolellidae</taxonomicName>
instead of other
<taxonomicName authorityName="Brölemann" authorityYear="1913" box="[236,391,413,439]" order="null" pageId="42" pageNumber="4" rank="subOrder" subOrder="Trigoniulidea">Trigoniulidea</taxonomicName>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767300" ID-Zenodo-Dep="3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="41" pageNumber="42" startId="41.[141,213,1526,1548]" targetBox="[140,1109,204,1471]" targetPageId="41">
<paragraph blockId="41.[141,1109,1526,1676]" pageId="41" pageNumber="42">
<emphasis bold="true" box="[141,251,1526,1548]" pageId="41" pageNumber="42">Figure 23.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[256,449,1526,1547]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="pauliani" status="sp. n.">
<emphasis box="[256,449,1526,1547]" italics="true" pageId="41" pageNumber="42">Flagellobolus pauliani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[455,506,1527,1548]" pageId="41" pageNumber="42" rank="species">sp. n.</taxonomicNameLabel>
, male holotype,
<emphasis bold="true" box="[657,676,1526,1548]" pageId="41" pageNumber="42">A</emphasis>
telson
<emphasis bold="true" box="[741,758,1526,1548]" pageId="41" pageNumber="42">B</emphasis>
left midbody leg
<emphasis bold="true" box="[918,937,1526,1548]" pageId="41" pageNumber="42">C</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[271,290,1558,1580]" pageId="41" pageNumber="42">D</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[613,628,1558,1580]" pageId="41" pageNumber="42">E</emphasis>
posterior gonopods, posterior view, slightly turned
<emphasis bold="true" box="[141,155,1590,1612]" pageId="41" pageNumber="42">F</emphasis>
posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection pageId="42" pageNumber="43" type="distribution">
<paragraph blockId="42.[140,1108,167,616]" pageId="42" pageNumber="43">
<emphasis bold="true" box="[185,487,448,474]" pageId="42" pageNumber="43">
<emphasis box="[185,336,448,474]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Distribution</emphasis>
and ecology
</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[502,640,448,474]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[502,640,448,474]" italics="true" pageId="42" pageNumber="43">Flagellobolus</emphasis>
</taxonomicName>
was collected on
<collectingCountry box="[841,974,448,474]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
in the eastern rainforest of Périnet (Andasibe) and the montane rainforest of Andranomay. Its ecology is entirely unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="42" pageNumber="43" type="etymology">
<paragraph blockId="42.[140,1108,167,616]" pageId="42" pageNumber="43">
<emphasis bold="true" box="[185,315,554,580]" inLineHeading="true" pageId="42" pageNumber="43" reason="1">Etymology</emphasis>
:
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[329,467,554,580]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis box="[329,467,554,580]" italics="true" pageId="42" pageNumber="43">Flagellobolus</emphasis>
</taxonomicName>
, masculine, is composed of
<emphasis box="[790,865,554,580]" italics="true" pageId="42" pageNumber="43">flagello</emphasis>
, after
<emphasis box="[943,1042,554,580]" italics="true" pageId="42" pageNumber="43">flagellum</emphasis>
, latin for whip, which refers to the characteristically shaped posterior gonopods, and -
<emphasis box="[1040,1095,589,615]" italics="true" pageId="42" pageNumber="43">bolus</emphasis>
.
</paragraph>
</subSubSection>
</subSection>
</treatment>
</document>