treatments-xml/data/3C/34/5D/3C345D7BFF89FF89FCA6FF30B1D6FD67.xml
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<document id="D14B55B0504EB8349923B1C311AE3602" ID-DOI="10.11646/phytotaxa.610.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8308556" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1693574121451" checkinUser="plazi" docAuthor="Martínez-Azorín, Mario, Crespo, Manuel B., Alonso-Vargas, María Ángeles, Pinter, Michael, Crouch, Neil R., Dold, Anthony P., Mucina, Ladislav, Pfosser, Martin &amp; Wetschnig, Wolfgang" docDate="2023" docId="3C345D7BFF89FF89FCA6FF30B1D6FD67" docLanguage="en" docName="phytotaxa.610.1.1.pdf" docOrigin="Phytotaxa 610 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.610.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Triandra Mart.-Azorín, M.B.Crespo, M.Á.Alonso, N.R.Crouch &amp; M.Pinter, N.R." docType="treatment" docVersion="7" lastPageNumber="109" masterDocId="C00D2503FFE2FFE5FC2EFFAEB453FFEB" masterDocTitle="A generic monograph of the Hyacinthaceae subfamily Urgineoideae" masterLastPageNumber="143" masterPageNumber="1" pageNumber="108" updateTime="1694254639663" updateUser="carolina">
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<mods:title id="9A717542A93D16865FD01D44C137165E">A generic monograph of the Hyacinthaceae subfamily Urgineoideae</mods:title>
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<mods:namePart id="4F2D1C0D564FEB3063056CDDDD7872D6">Martínez-Azorín, Mario</mods:namePart>
<mods:affiliation id="AA3999C29651CBA8AF7D6EA0E1002652">Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; E-mail: mmartinez @ ua. es; ORCID: https: // orcid. org / 0000 - 0002 - 2605 - 9575</mods:affiliation>
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<mods:namePart id="50BD1A450FDAE74FBF098A67A592BFD2">Crespo, Manuel B.</mods:namePart>
<mods:affiliation id="1F2495A9857ED30650CB5CF23F289101">Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; E-mail: crespo @ ua. es; ORCID: https: // orcid. org / 0000 - 0002 - 3294 - 5637</mods:affiliation>
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<mods:affiliation id="957816E06E14A07B13B6E3CA0FDF5389">Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; E-mail: ma. alonso @ ua. es; ORCID: https: // orcid. org / 0000 - 0003 - 3768 - 9203</mods:affiliation>
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<mods:namePart id="C1DF38EB793CB25AF324C28959391596">Pinter, Michael</mods:namePart>
<mods:affiliation id="E0AEB24EFF27B930B73163A209D9B474">Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; Institute of Biology, NAWI Graz, Division Plant Sciences, Karl-Franzens University Graz, Holteigasse 6, A- 8010 Graz, Austria. &amp; E-mail: michael. pinter @ uni-graz. at; ORCID: https: // orcid. org / 0000 - 0002 - 6055 - 6989</mods:affiliation>
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<mods:namePart id="1DB9E0815804075756ED08EC74FAD9FF">Crouch, Neil R.</mods:namePart>
<mods:affiliation id="FD60E34B599E032E0108DE6916C9D00B">BRAM, South African National Biodiversity Institute, P. O. Box 52099, Berea Road 4007, South Africa. &amp; School of Chemistry and Physics, University of KwaZulu-Natal, Durban 4041, South Africa. &amp; E-mail: N. Crouch @ sanbi. org. za; ORCID: https: // orcid. org / 0000 - 0002 - 4938 - 5840.</mods:affiliation>
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<mods:affiliation id="906A244F8B19B5471EF896AAF8EC550F">Selmar Schonland Herbarium, Department of Botany, Rhodes University, Makhanda, 6140, South Africa. &amp; E-mail: t. dold @ ru. ac. za; ORCID: https: // orcid. org / 0000 - 0002 - 9497 - 7503</mods:affiliation>
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<mods:namePart id="7EA112A0D221180448E2D2FF6A694348">Mucina, Ladislav</mods:namePart>
<mods:affiliation id="C5CFE613F8390C991FC78067F2464C4B">Harry Butler Institute, Murdoch University, 90 South Street, Building 390, Murdoch WA 6150, Perth, Australia. &amp; Dept. of Geography &amp; Environmental Studies, Stellenbosch University, Private Bag X 1, Matieland 7602, Stellenbosch, South Africa. &amp; E-mail: ladislav. mucina @ murdoch. edu. au; ORCID: https: // orcid. org / 0000 - 0003 - 0317 - 8886</mods:affiliation>
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<mods:namePart id="E09288D5DFCB99E1770434CE8530D90F">Pfosser, Martin</mods:namePart>
<mods:affiliation id="BA65B42659512E9D5F62C2A64072C1F3">Biocenter Linz, J. - W. - Klein-Str. 73, A- 4040 Linz, Austria. &amp; E-mail: martin. pfosser @ ooelkg. at; ORCID: https: // orcid. org / 0000 - 0003 - 2050 - 4997</mods:affiliation>
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<mods:namePart id="4D3367A57F7C4F80F5598F6E1604AFBA">Wetschnig, Wolfgang</mods:namePart>
<mods:affiliation id="DB4015FC2C75DED500FBC8AE802467E2">Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; Institute of Biology, NAWI Graz, Division Plant Sciences, Karl-Franzens University Graz, Holteigasse 6, A- 8010 Graz, Austria. &amp; Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain. &amp; E-mail: wolfgang. wetschnig @ uni-graz. at; ORCID: https: // orcid. org / 0000 - 0002 - 9245 - 029 X * Author for correspondence &amp; Depto. Ciencias Ambientales y Recursos Naturales (dCARN), Universidad de Alicante, P. O. Box 99, ES- 03080 Alicante, Spain.</mods:affiliation>
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<mods:title id="72C540D4D3CB8380EA6EB9DE11811E6B">Phytotaxa</mods:title>
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<mods:date id="8FEE51E0B361EE1012C63D97C6339B68">2023</mods:date>
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<mods:number id="EFD4D7DAC1267A294CD8AB7CD0DF7BD9">610</mods:number>
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25.
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<emphasis id="86E9307FFF89FF8EFC9DFF30B549FF52" bold="true" box="[179,282,158,185]" italics="true" pageId="107" pageNumber="108">Triandra</emphasis>
Mart.-
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Azorín, M.B.Crespo, M.Á.Alonso,
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Crouch &amp; M.Pinter
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in
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(
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).
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Typus generis:—
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<emphasis id="86E9307FFF89FF8EFD55FF6DB66BFF37" box="[379,568,194,220]" italics="true" pageId="107" pageNumber="108">T. pellabergensis</emphasis>
Mart.
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-Azorín, M.B.Crespo, M.Á.Alonso,
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Crouch &amp; M.Pinter (
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).
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<emphasis id="86E9307FFF89FF8EFCA6FEA4B541FECF" bold="true" box="[136,274,266,292]" pageId="107" pageNumber="108">Description</emphasis>
:—Small bulbous geophyte.
<emphasis id="86E9307FFF89FF8EFE78FEA4B6D8FECF" box="[598,651,266,292]" italics="true" pageId="107" pageNumber="108">Bulb</emphasis>
hypogeal, ovoid to subglobose,
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in diam., with white, fleshy, compact, inner scales and pale brown, membranous outer tunics. Roots thickened and branched.
<emphasis id="86E9307FFF89FF8EF8D7FE81B114FEA3" box="[1273,1351,303,328]" italics="true" pageId="107" pageNumber="108">Leaves</emphasis>
26 per bulb, hysteranthous,
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long, filiform, to
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in diam., subterete or very slightly flattened, green, smooth, glabrous.
<emphasis id="86E9307FFF89FF8EFCD6FED8B5DDFE7B" box="[248,398,374,400]" italics="true" pageId="107" pageNumber="108">Inflorescence</emphasis>
a lax, erect raceme,
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long, with 48 flowers; peduncle
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long, brownish, erect, terete, smooth; pedicels
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long, smooth, patent proximally and soon arching downwards distally at anthesis, curving upwards in fruit. Bracts ovate-lanceolate, small, ca.
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long, lowermost with short spur, ca.
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long, persistent; bracteoles absent.
<emphasis id="86E9307FFF89FF8EFDE0FE4CB679FE17" box="[462,554,482,508]" italics="true" pageId="107" pageNumber="108">Flowers</emphasis>
stellate, nodding, with strongly reflexed tepals at full anthesis with tips touching flower pedicel, nocturnal, opening in late evening and closing before sunrise, fragrant during night. Tepals 6, biseriate,
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long, narrowly oblong, slightly constricted along middle, obtuse, whitish to pale brown, with darker brownish longitudinal band more evident on abaxial side, almost free to very shortly connate for
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.
<emphasis id="86E9307FFF89FF8EF897FDE1B145FD82" box="[1209,1302,591,617]" italics="true" pageId="107" pageNumber="108">Stamens</emphasis>
3, only inner whorl present, outer absent with no basal rudiments, erect, straight, approaching ovary and style, with 3 anthers placed near stigma; filaments
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long, filiform, white, subterete, only slightly widened at base, connivent to style, free, not adnate to tepals; anthers oblong, ca.
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long after dehiscence, subbasifixed, yellow to orange, with yellow polen.
<emphasis id="86E9307FFF89FF8EFD07FD71B53CFD12" box="[297,367,735,761]" italics="true" pageId="107" pageNumber="108">Ovary</emphasis>
ovate,
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long, pale brown to yellowish. Style 2.0
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long, erect, white, filiform and narrow, ending in undifferentiated stigma.
<emphasis id="86E9307FFF89FF8EFEBCFCACB6BDFCF7" box="[658,750,770,796]" italics="true" pageId="107" pageNumber="108">Capsule</emphasis>
ovate, trigonous,
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long, pale brown, valves completely dehiscing from base, tepals cohering and inrolled above ovary after anthesis, circumcissile from base and persisting as a cap at the top of the developing capsules.
<emphasis id="86E9307FFF89FF8EFE44FCE4B6F9FC8F" box="[618,682,842,868]" italics="true" pageId="107" pageNumber="108">Seeds</emphasis>
ellipsoidal to broadly ellipsoid, flattened,
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long, testa black, glossy, with subpolygonal cells.
</paragraph>
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<paragraph id="B422EC6DFF89FF8EFCA6F9EDB6E1F9B3" blockId="107.[136,690,1603,1625]" box="[136,690,1603,1625]" pageId="107" pageNumber="108">
<emphasis id="86E9307FFF89FF8EFCA6F9EDB55AF9B2" bold="true" box="[136,265,1603,1625]" pageId="107" pageNumber="108">FIGURE 62.</emphasis>
<taxonomicName id="739D97EEFF89FF8EFD21F9EDB662F9B3" authority="Mart." authorityName="Mart." box="[271,561,1603,1624]" class="Magnoliopsida" family="Amaranthaceae" genus="Triandra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="species" species="pellabergensis">
<emphasis id="86E9307FFF89FF8EFD21F9EDB5A5F9B3" box="[271,502,1603,1624]" italics="true" pageId="107" pageNumber="108">Triandra pellabergensis</emphasis>
Mart.
</taxonomicName>
-Azorín et al.
</paragraph>
</caption>
<paragraph id="B422EC6DFF89FF8EFC93F92FB780F857" blockId="107.[136,1452,1665,2088]" pageId="107" pageNumber="108">
<emphasis id="86E9307FFF89FF8EFC93F92FB63FF970" bold="true" box="[189,620,1665,1691]" pageId="107" pageNumber="108">Number of species and distribution</emphasis>
:—A monotypic genus originally described from a single locality on the Pella-se-Berge, northwestern
<collectingCountry id="CC8AACFDFF89FF8EFDF8F90BB634F954" box="[470,615,1701,1727]" name="South Africa" pageId="107" pageNumber="108">South Africa</collectingCountry>
, near to the border with
<collectingCountry id="CC8AACFDFF89FF8EFF51F90BB7B1F954" box="[895,994,1701,1727]" name="Namibia" pageId="107" pageNumber="108">Namibia</collectingCountry>
, a locality included in the Karoo-Namib Region (sensu
<bibRefCitation id="D00C919CFF89FF8EFD00F967B588F908" author="Takhtajan, A." box="[302,475,1737,1763]" pageId="107" pageNumber="108" refId="ref110444" refString="Takhtajan, A. (1986) Florisitic regions of the World. University of California Press, Berkeley, 522 pp." type="book" year="1986">Takhtajan 1986</bibRefCitation>
) (
<figureCitation id="2CA6F0E8FF89FF8EFDDDF967B614F908" box="[499,583,1737,1763]" captionStart="FIGURE 49" captionStartId="85.[136,229,1841,1863]" captionTargetBox="[155,1429,199,1812]" captionTargetId="figure-19@85.[150,1435,193,1817]" captionTargetPageId="85" captionText="FIGURE 49. Approximate distribution ranges of accepted genera in Urgineoideae." figureDoi="http://doi.org/10.5281/zenodo.8308656" httpUri="https://zenodo.org/record/8308656/files/figure.png" pageId="107" pageNumber="108">Fig. 49</figureCitation>
). Recently,
<bibRefCitation id="D00C919CFF89FF8EFEE7F967B7D3F908" author="Manning, J. C." box="[713,896,1737,1763]" pageId="107" pageNumber="108" pagination="1 - 7" refId="ref104375" refString="Manning, J. C. (2022) New combinations in Drimia Jacq. ex Willd. (Hyacinthaceae: Urgineoideae) and an updated key to the southern African species. Bothalia 52 (1), a 2: 1 - 7. http: // dx. doi. org / 10.38201 / btha. abc. v 52. i 1.2" type="journal article" year="2022">Manning (2022)</bibRefCitation>
cited the collection
<emphasis id="86E9307FFF89FF8EF84CF964B143F908" box="[1122,1296,1737,1763]" italics="true" pageId="107" pageNumber="108">Harrower 6188</emphasis>
(NBG) as the second known locality for this species occurring in the Pienaarsfontein-se-Berge, Koedoesberge near Touwsrivier, representing an extension of more than
<quantity id="73654188FF89FF8EFE7FF8BFB6FBF8C0" box="[593,680,1809,1835]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="107" pageNumber="108" unit="km" value="400.0">400 km</quantity>
to the south, near the Tanqua Karoo in the
<collectingRegion id="7659228FFF89FF8EF88FF8BCB1FFF8C0" box="[1185,1452,1809,1836]" country="South Africa" name="Western Cape" pageId="107" pageNumber="108">Western Cape Province</collectingRegion>
of
<collectingCountry id="CC8AACFDFF89FF8EFC89F89BB569F8A4" box="[167,314,1845,1871]" name="South Africa" pageId="107" pageNumber="108">South Africa</collectingCountry>
. Two additional populations were found, in
<date id="C023CAADFF89FF8EFF02F89BB7B7F8BB" box="[812,996,1845,1872]" pageId="107" pageNumber="108" value="2015-09">
September
<quantity id="73654188FF89FF8EFF82F89BB7B7F8BB" box="[940,996,1845,1872]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="107" pageNumber="108" unit="in" value="2015.0">2015</quantity>
</date>
in the Naib se Berg and in
<date id="C023CAADFF89FF8EF935F89BB1F8F8A4" box="[1307,1451,1845,1871]" pageId="107" pageNumber="108" value="2022-08">
August
<quantity id="73654188FF89FF8EF95DF89BB1F8F8A4" box="[1395,1451,1845,1871]" metricMagnitude="1" metricUnit="m" metricValue="5.135879999999999" pageId="107" pageNumber="108" unit="in" value="2022.0">2022</quantity>
</date>
in the Nouroepberge near Dabenoris Mountains, both in vegetative stage, by M. Martínez-Azorín &amp; collaborators (
<emphasis id="86E9307FFF89FF8EFCBFF8D3B55AF87C" box="[145,265,1917,1943]" italics="true" pageId="107" pageNumber="108">MMA1269</emphasis>
ABH74121! and
<emphasis id="86E9307FFF89FF8EFDFBF8D3B61EF87C" box="[469,589,1917,1943]" italics="true" pageId="107" pageNumber="108">MMA2078</emphasis>
ABH74374! respectively). The phylogenetic analyses of these samples and their recent flowering in cultivation confirmed their placement in
<taxonomicName id="739D97EEFF89FF8EFF46F80FB79FF850" box="[872,972,1953,1979]" class="Magnoliopsida" family="Amaranthaceae" genus="Triandra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="genus">
<emphasis id="86E9307FFF89FF8EFF46F80FB79FF850" box="[872,972,1953,1979]" italics="true" pageId="107" pageNumber="108">Triandra</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="B422EC6DFF89FF8EFC93F86BB641F80B" blockId="107.[136,1452,1665,2088]" box="[189,530,1989,2016]" pageId="107" pageNumber="108">
<emphasis id="86E9307FFF89FF8EFC93F86BB568F834" bold="true" box="[189,315,1989,2015]" pageId="107" pageNumber="108">Karyology</emphasis>
:—
<collectingRegion id="7659228FFF89FF8EFD73F868B5DAF80B" box="[349,393,1990,2016]" country="Cameroon" name="North" pageId="107" pageNumber="108">Not</collectingRegion>
studied yet.
</paragraph>
</subSubSection>
<subSubSection id="FC87BFE6FF89FF89FC93F847B1D6FD67" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108" type="multiple">
<paragraph id="B422EC6DFF89FF89FC93F847B552FE7B" blockId="107.[136,1452,1665,2088]" lastBlockId="108.[136,1452,158,436]" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
<emphasis id="86E9307FFF89FF8EFC93F847B7DFF7E8" bold="true" box="[189,908,2025,2051]" pageId="107" pageNumber="108">History, diagnostic characters, and taxonomic relationships</emphasis>
:—Our field work in Southern Africa led to the discovery at the Pella-se-Berg (NW
<collectingCountry id="CC8AACFDFF89FF8EFE0EF7A3B6E2F7CC" box="[544,689,2061,2087]" name="South Africa" pageId="107" pageNumber="108">South Africa</collectingCountry>
) of a small species of
<taxonomicName id="739D97EEFF89FF8EFF83F7A3B010F7CC" box="[941,1091,2061,2087]" class="Liliopsida" family="Asparagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="107" pageNumber="108" phylum="Tracheophyta" rank="subFamily" subFamily="Urgineoideae">Urgineoideae</taxonomicName>
with a remarkable morphology. The plants produce nocturnal, nodding flowers with most notably only three stamens per flower, a character not previously reported in
<taxonomicName id="739D97EEFF8EFF89FDBFFF6CB669FF37" authorityName="sensu APG" authorityYear="2003" box="[401,570,194,220]" class="Liliopsida" family="Hyacinthaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="family">Hyacinthaceae</taxonomicName>
. This led
<bibRefCitation id="D00C919CFF8EFF89FE9EFF6CB05BFF37" author="Martinez-Azorin, M. &amp; Crespo, M. B. &amp; Alonso-Vargas, M. A. &amp; Crouch, N. R. &amp; Pinter, M." box="[688,1032,194,220]" pageId="108" pageNumber="109" pagination="65 - 74" refId="ref106533" refString="Martinez-Azorin, M., Crespo, M. B., Alonso-Vargas, M. A., Crouch, N. R. &amp; Pinter, M. (2021) Triandra pellabergensis (Hyacinthaceae subfam. Urgineoideae), a new genus and species from Pella se Berge, Northern Cape Province, South Africa. Phytotaxa 487 (1): 65 - 74. https: // doi. org / 10.11646 / phytotaxa. 487.1.5" type="journal article" year="2021">
Martínez-Azorín
<emphasis id="86E9307FFF8EFF89FF59FF6DB7E0FF37" box="[887,947,194,220]" italics="true" pageId="108" pageNumber="109">et al.</emphasis>
(2021)
</bibRefCitation>
to describe
<taxonomicName id="739D97EEFF8EFF89F8B5FF6CB65FFEEB" authority="Martinez-Azorin et al. (2021: 68)" authorityName="Martinez-Azorin" authorityPageNumber="68" authorityYear="2021" class="Magnoliopsida" family="Amaranthaceae" genus="Triandra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="pellabergensis">
<emphasis id="86E9307FFF8EFF89F8B5FF6CB1F8FF37" box="[1179,1451,194,220]" italics="true" pageId="108" pageNumber="109">Triandra pellabergensis</emphasis>
<bibRefCitation id="D00C919CFF8EFF89FCA6FF48B65FFEEB" author="Martinez-Azorin, M. &amp; Crespo, M. B. &amp; Alonso-Vargas, M. A. &amp; Crouch, N. R. &amp; Pinter, M." box="[136,524,230,256]" pageId="108" pageNumber="109" pagination="65 - 74" refId="ref106533" refString="Martinez-Azorin, M., Crespo, M. B., Alonso-Vargas, M. A., Crouch, N. R. &amp; Pinter, M. (2021) Triandra pellabergensis (Hyacinthaceae subfam. Urgineoideae), a new genus and species from Pella se Berge, Northern Cape Province, South Africa. Phytotaxa 487 (1): 65 - 74. https: // doi. org / 10.11646 / phytotaxa. 487.1.5" type="journal article" year="2021">
Martínez-Azorín
<emphasis id="86E9307FFF8EFF89FD61FF49B5D9FEEB" box="[335,394,230,256]" italics="true" pageId="108" pageNumber="109">et al.</emphasis>
(2021: 68)
</bibRefCitation>
</taxonomicName>
—a new genus and species. The phylogenetic studies by
<bibRefCitation id="D00C919CFF8EFF89F887FF48B48CFECF" author="Martinez-Azorin, M. &amp; Crespo, M. B. &amp; Alonso, M. A. &amp; Pinter, M. &amp; Crouch, N. R. &amp; Dold, A. P. &amp; Mucina, L. &amp; Pfosser, M. &amp; Wetschnig, W." pageId="108" pageNumber="109" pagination="42 - 63" refId="ref106677" refString="Martinez-Azorin, M., Crespo, M. B., Alonso, M. A., Pinter, M., Crouch, N. R., Dold, A. P., Mucina, L., Pfosser, M. &amp; Wetschnig, W. (2023 a) Molecular phylogenetics of subfamily Urgineoideae (Hyacinthaceae): Towards a coherent generic circumscription informed by molecular, morphological and distributional data. Journal of Systematics and Evolution 63 (1): 42 - 63. https: // doi. org / 10.1111 / jse. 12905" type="journal article" year="2023">
Martínez-Azorín
<emphasis id="86E9307FFF8EFF89F95EFF49B1FFFEEB" box="[1392,1452,230,256]" italics="true" pageId="108" pageNumber="109">et al.</emphasis>
(2023a)
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based on plastidial DNA sequences placed this species in an isolated position, sister to a clade comprising samples of
<taxonomicName id="739D97EEFF8EFF89FD29FE80B5C6FEA3" box="[263,405,302,328]" class="Liliopsida" family="Asparagaceae" genus="Rhodocodon" kingdom="Plantae" order="Asparagales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="genus">
<emphasis id="86E9307FFF8EFF89FD29FE80B5C6FEA3" box="[263,405,302,328]" italics="true" pageId="108" pageNumber="109">Rhodocodon</emphasis>
</taxonomicName>
from
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. Therefore, we accept
<taxonomicName id="739D97EEFF8EFF89FF73FE80B792FEA3" box="[861,961,302,328]" class="Magnoliopsida" family="Amaranthaceae" genus="Triandra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="genus">
<emphasis id="86E9307FFF8EFF89FF73FE80B792FEA3" box="[861,961,302,328]" italics="true" pageId="108" pageNumber="109">Triandra</emphasis>
</taxonomicName>
to accommodate this outstanding species in the subfamily, which is supported by clear morphological, biogeographical, and molecular data (
<bibRefCitation id="D00C919CFF8EFF89F8FFFEFCB4A6FE7B" author="Martinez-Azorin, M. &amp; Crespo, M. B. &amp; Alonso, M. A. &amp; Pinter, M. &amp; Crouch, N. R. &amp; Dold, A. P. &amp; Mucina, L. &amp; Pfosser, M. &amp; Wetschnig, W." pageId="108" pageNumber="109" pagination="42 - 63" refId="ref106677" refString="Martinez-Azorin, M., Crespo, M. B., Alonso, M. A., Pinter, M., Crouch, N. R., Dold, A. P., Mucina, L., Pfosser, M. &amp; Wetschnig, W. (2023 a) Molecular phylogenetics of subfamily Urgineoideae (Hyacinthaceae): Towards a coherent generic circumscription informed by molecular, morphological and distributional data. Journal of Systematics and Evolution 63 (1): 42 - 63. https: // doi. org / 10.1111 / jse. 12905" type="journal article" year="2023">
Martínez-Azorín
<emphasis id="86E9307FFF8EFF89F9B9FEFDB4F6FE7B" italics="true" pageId="108" pageNumber="109">et al.</emphasis>
2023a
</bibRefCitation>
).
</paragraph>
<paragraph id="B422EC6DFF8EFF89FC93FE34B5F0FE5F" blockId="108.[136,1452,158,436]" box="[189,419,410,436]" pageId="108" pageNumber="109">Accepted species:—</paragraph>
<paragraph id="B422EC6DFF8EFF89FCA6FE4BB1D6FD67" blockId="108.[136,1452,485,653]" pageId="108" pageNumber="109">
<treatmentCitationGroup id="948DCB43FF8EFF89FCA6FE4BB1D6FD67" pageId="108" pageNumber="109">
<taxonomicName id="739D97EEFF8EFF89FCA6FE4BB5E9FE10" authority="Mart." authorityName="Mart." box="[136,442,485,507]" class="Magnoliopsida" family="Amaranthaceae" genus="Triandra" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="pellabergensis">
<emphasis id="86E9307FFF8EFF89FCA6FE4BB52AFE10" bold="true" box="[136,377,485,507]" italics="true" pageId="108" pageNumber="109">Triandra pellabergensis</emphasis>
Mart.
</taxonomicName>
-
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<bibCitation id="61B7E401FF8EFF89FD91FE48B111FE10" author="Azorin, M. B. Crespo, M. A. Alonso, N. R. Crouch &amp; M. Pinter" box="[447,1346,486,509]" journal="Phytotaxa" page="68" pageId="108" pageNumber="109" score="9.729594737742866" volume="487" year="2021">
<collectorName id="196889BBFF8EFF89FD91FE48B66CFE10" box="[447,575,486,507]" pageId="108" pageNumber="109">Azorín, M.B</collectorName>
.
<collectorName id="196889BBFF8EFF89FE6FFE48B69BFE16" box="[577,712,486,509]" pageId="108" pageNumber="109">Crespo, M.Á</collectorName>
.
<collectorName id="196889BBFF8EFF89FEE4FE48B719FE17" box="[714,842,486,509]" pageId="108" pageNumber="109">
Alonso,
<taxonomicNameLabel id="9DDA8D04FF8EFF89FF0AFE48B719FE17" box="[804,842,486,508]" pageId="108" pageNumber="109" rank="subSpecies">N.R</taxonomicNameLabel>
</collectorName>
.Crouch &amp;
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M.
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</collectorName>
in
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<taxonomicName id="739D97EEFF8EFF89F948FE48B51BFDF4" class="Liliopsida" family="Asparagaceae" genus="Drimia" kingdom="Plantae" order="Asparagales" pageId="108" pageNumber="109" phylum="Tracheophyta" rank="species" species="pellabergensis">
<emphasis id="86E9307FFF8EFF89F948FE48B51BFDF4" italics="true" pageId="108" pageNumber="109">Drimia pellabergensis</emphasis>
</taxonomicName>
(
<collectorName id="196889BBFF8EFF89FD79FDA4B5DEFDF4" box="[343,397,522,543]" pageId="108" pageNumber="109">Mart.</collectorName>
-
<collectorName id="196889BBFF8EFF89FDBDFDA4B65CFDF4" box="[403,527,522,543]" pageId="108" pageNumber="109">Azorín, M.B</collectorName>
.
<collectorName id="196889BBFF8EFF89FE3CFDA4B6C6FDCA" box="[530,661,522,545]" pageId="108" pageNumber="109">Crespo, M.Á</collectorName>
.
<collectorName id="196889BBFF8EFF89FEB6FDA4B747FDCB" box="[664,788,522,545]" pageId="108" pageNumber="109">
Alonso,
<taxonomicNameLabel id="9DDA8D04FF8EFF89FEC0FDA4B747FDCB" box="[750,788,522,544]" pageId="108" pageNumber="109" rank="subSpecies">N.R</taxonomicNameLabel>
</collectorName>
.Crouch &amp;
<collectorName id="196889BBFF8EFF89FFAAFDA4B789FDCB" box="[900,986,522,544]" pageId="108" pageNumber="109">M.Pinter</collectorName>
)
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<bibCitation id="61B7E401FF8EFF89FFC7FDA4B4ADFDA8" author="J. C. Manning &amp; Goldblatt" issue="1" journal="Bothalia" page="3" pageId="108" pageNumber="109" score="9.729594737742866" volume="52" year="2022">
<collectorName id="196889BBFF8EFF89FFC7FDA4B036FDCB" box="[1001,1125,522,544]" pageId="108" pageNumber="109">J.C.Manning</collectorName>
&amp;
<collectingMunicipality id="54467617FF8EFF89F8A9FDA4B106FDCB" box="[1159,1365,522,544]" pageId="108" pageNumber="109">
Goldblatt in
<bibCitation id="61B7E401FF8EFF89F92AFDA4B106FDCB" box="[1284,1365,522,544]" issue="1" journal="Bothalia" page="3" pageId="108" pageNumber="109" score="9.729594737742866" volume="52" year="2022">Bothalia</bibCitation>
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52(1): 3 (2022)
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(
<figureCitation id="2CA6F0E8FF8EFF89FD25FD80B50FFDAF" box="[267,348,558,580]" captionStart="FIGURE 5" captionStartId="16.[136,229,1804,1826]" captionTargetBox="[158,1429,193,1777]" captionTargetId="figure-21@16.[158,1430,191,1779]" captionTargetPageId="16" captionText="FIGURE 5. Seed morphology in Urgineoideae. 1. Tenicroa multifolia (G.J.Lewis) Oberm.; 2. Tenicroa polyantha M.Pinter et al.; 3. Tenicroa unifolia (A.V.Duthie) M.Pinter et al.; 4. Thuranthos bakeri Mart.-Azorín et al.; 5. Thuranthos macrocarpum (Stedje) Speta; 6. Thuranthos nocturnale R.A.Dyer; 7. Thuranthos pauciflorum (Baker) Mart.-Azorín et al.; 8. Triandra pellabergensis Mart.-Azorín et al.; 9. Urginavia altissima (L.f.) Speta; 10. Urginavia capensis (Burm.f.) Mart.-Azorín et al.; 11. Urginavia echinostachya (Baker) Mart.-Azorín et al.; 12. Urginavia kniphofioides (Baker) Mart.-Azorín et al.; 13. Urginavia micrantha (A.Rich.) Speta; 14. Urginavia multisetosa (Baker) Mart.-Azorín et al.; 15. Urginavia simensis (Hochst. ex A.Rich.) Mart.-Azorín et al.; 16. Urginea fugax (Moris) Steinh.; 17. Urgineopsis arenosa (Adamson) Mart.-Azorín et al.; 18. Urgineopsis dregei (Baker) Mart.-Azorín et al.; 19. Urgineopsis exilis (Adamson) Mart.-Azorín et al.; 20. Urgineopsis gracilis (A.V.Duthie) Mart.-Azorín et al. All to the same scale in mm." figureDoi="http://doi.org/10.5281/zenodo.8308566" httpUri="https://zenodo.org/record/8308566/files/figure.png" pageId="108" pageNumber="109">Figs 5.8</figureCitation>
,
<figureCitation id="2CA6F0E8FF8EFF89FD48FD80B5D2FDAF" box="[358,385,558,580]" captionStart="FIGURE 62" captionStartId="107.[136,229,1603,1625]" captionTargetBox="[155,1430,937,1574]" captionTargetId="figure-349@107.[150,1435,932,1579]" captionTargetPageId="107" captionText="FIGURE 62. Triandra pellabergensis Mart.-Azorín et al." figureDoi="http://doi.org/10.5281/zenodo.8308685" httpUri="https://zenodo.org/record/8308685/files/figure.png" pageId="108" pageNumber="109">62</figureCitation>
).
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:—
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.
<collectingRegion id="7659228FFF8EFF89FEB5FD80B77FFDA8" box="[667,812,558,580]" country="South Africa" name="Northern Cape" pageId="108" pageNumber="109">Northern Cape</collectingRegion>
.
<location id="B142BAB6FF8EFF89FF18FD80B7DEFDAF" LSID="urn:lsid:plazi:treatment:3C345D7BFF89FF89FCA6FF30B1D6FD67:B142BAB6FF8EFF89FF18FD80B7DEFDAF" box="[822,909,558,580]" country="South Africa" county="Pinter in Phytotaxa" municipality="Goldblatt in Bothalia" name="Pofadder" pageId="108" pageNumber="109" stateProvince="Northern Cape">Pofadder</location>
(2919):
<location id="B142BAB6FF8EFF89FFCEFD80B041FDAF" LSID="urn:lsid:plazi:treatment:3C345D7BFF89FF89FCA6FF30B1D6FD67:B142BAB6FF8EFF89FFCEFD80B041FDAF" box="[992,1042,558,580]" country="South Africa" county="Pinter in Phytotaxa" municipality="Goldblatt in Bothalia" name="Pella" pageId="108" pageNumber="109" stateProvince="Northern Cape">Pella</location>
,
<location id="B142BAB6FF8EFF89F833FD80B01EFDAF" LSID="urn:lsid:plazi:treatment:3C345D7BFF89FF89FCA6FF30B1D6FD67:B142BAB6FF8EFF89F833FD80B01EFDAF" box="[1053,1101,558,580]" country="South Africa" county="Pinter in Phytotaxa" municipality="Goldblatt in Bothalia" name="Pella" pageId="108" pageNumber="109" stateProvince="Northern Cape">Pella</location>
se
<location id="B142BAB6FF8EFF89F840FD80B0FAFDA8" LSID="urn:lsid:plazi:treatment:3C345D7BFF89FF89FCA6FF30B1D6FD67:B142BAB6FF8EFF89F840FD80B0FAFDA8" box="[1134,1193,558,579]" country="South Africa" county="Pinter in Phytotaxa" municipality="Goldblatt in Bothalia" name="Berge" pageId="108" pageNumber="109" stateProvince="Northern Cape">Berge</location>
, ca.
<locationDeviation id="A590F54FFF8EFF89F8FBFD80B1FBFDAF" box="[1237,1448,558,580]" location="Pofadder" pageId="108" pageNumber="109">
<quantity id="73654188FF8EFF89F8FBFD80B140FDAF" box="[1237,1299,558,580]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="108" pageNumber="109" unit="km" value="25.0">25 km</quantity>
W of Pofadder
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, rocky slopes at base of large south facing cliffs, (AA), elev.
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</quantity>
, among quartz and mica rocks, fl. ex hort at
<location id="B142BAB6FF8EFF89F904FDFCB55DFD67" LSID="urn:lsid:plazi:treatment:3C345D7BFF89FF89FCA6FF30B1D6FD67:B142BAB6FF8EFF89F904FDFCB55DFD67" country="South Africa" county="Pinter in Phytotaxa" municipality="Goldblatt in Bothalia" name="University of Alicante" pageId="108" pageNumber="109" stateProvince="Northern Cape">University of Alicante</location>
on
<date id="C023CAADFF8EFF89FD1DFDD8B5E6FD60" box="[307,437,630,651]" pageId="108" pageNumber="109" value="2018-06-29">
<collectingDate id="D0673345FF8EFF89FD1DFDD8B5E6FD60" box="[307,437,630,651]" pageId="108" pageNumber="109" value="2018-06-29">29 June 2018</collectingDate>
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,
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<collectorName id="196889BBFF8EFF89FD91FDD8B6D3FD60" box="[447,640,630,651]" pageId="108" pageNumber="109">M. Martínez-Azorín</collectorName>
,
<collectorName id="196889BBFF8EFF89FEA7FDD8B756FD60" box="[649,773,630,651]" pageId="108" pageNumber="109">M.B. Crespo</collectorName>
,
<collectorName id="196889BBFF8EFF89FF20FDD8B7D5FD60" box="[782,902,630,653]" pageId="108" pageNumber="109">M.Á. Alonso</collectorName>
et al. MMA1671b
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(GRA! holo.; ABH!, K, PRE iso.).
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</paragraph>
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</treatment>
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