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<document id="BBE74E22CCF5C882512F07B26946B6DE" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="1FAF6C145BA45EB1A10D36DCFF31DAF9" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Glessula blanfordiana Nevill 1877" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="3F21FE6D9734ED29596548B5EE71B19F">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:namePart id="EE9204D4DC12342C963E98976B7E9693">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="B8B97EFF6D16EBAE062FD6A33EF68FBE">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="DE50A512FD96F493B1ADBD3656C88E7D">Ablett, Jonathan D.</mods:namePart>
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<mods:affiliation id="DD3042969C560F0DD467E61E4A220CB9">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:affiliation id="C8A6926B2B6E3674BAC9F24D83990A3A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:affiliation id="F139B7607693A51EBC3F313CC11223D1">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<treatment id="1FAF6C145BA45EB1A10D36DCFF31DAF9" ID-DOI="http://doi.org/10.5281/zenodo.13137110" ID-Zenodo-Dep="13137110" ID-arpha="1FAF6C14-5BA4-5EB1-A10D-36DCFF31DAF9" LSID="urn:lsid:plazi:treatment:1FAF6C145BA45EB1A10D36DCFF31DAF9" httpUri="http://treatment.plazi.org/id/1FAF6C145BA45EB1A10D36DCFF31DAF9">
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<paragraph id="51487170335A67F0322E6E6BA48A41CA">
<heading id="294C37F607FF2E0A9F081887C2278D86" reason="label">8</heading>
<taxonomicName id="6F383CBE33F97714F809B986D18760AD" authority="Nevill, 1877" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Subulinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="B37ECCBE31A72CB99568466F6BFAAC9B" italics="true">Glessula blanfordiana</emphasis>
<bibRefCitation id="6507260BECDC588F1EBC0FE2506C731E" author="Nevill G" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="14-41" refId="B68" refString="Nevill G (1877) List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46 (2): 14 - 41. https: // www. biodiversitylibrary. org / item / 110105" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." url="https://www.biodiversitylibrary.org/item/110105" volume="46" year="1877">Nevill, 1877</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph id="83191266FA61BCAD0A8B9DBF26F68C79">
<figureCitation id="4DE8DFE227666F1F5538608B6BCF1689" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">Fig. 5 A D</figureCitation>
,
<tableCitation id="9D40E60DF820BC3929F1679EDE55262B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Shell measurements of subulinid species examined in this study. SH: shell height; SW: shell width. The localities numbers are indicated in Fig. 1." httpUri="http://table.plazi.org/id/B099E9B05D409B67DECF1570A3A8359F" tableUuid="B099E9B05D409B67DECF1570A3A8359F">Table 1</tableCitation>
</paragraph>
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<paragraph id="22CF3B25F11F510271C03376BF1071B4">
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<treatmentCitation id="9CA01598304AAE6941B5860830B6187B" author="Nevill G" page="26" year="1877">
<taxonomicName id="5965266C66EB5B5D41E95FA0FF2A5C93" ID-CoL="7RFHX" authority="Nevill, 1877: 26" authorityName="Nevill" authorityPageNumber="26" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="B15AB45111FE7473054456E9DA87B1AA" italics="true">Glessula blanfordiana</emphasis>
<comment id="BC2E08E288014A1180F5FDF7361CAD22">
<bibRefCitation id="E1EDB58A81C029202DCDA1A340B6CDB5" author="Nevill G" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="14-41" refId="B68" refString="Nevill G (1877) List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46 (2): 14 - 41. https: // www. biodiversitylibrary. org / item / 110105" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." url="https://www.biodiversitylibrary.org/item/110105" volume="46" year="1877">Nevill, 1877: 26</bibRefCitation>
. Type locality: Ponsee,
<collectingRegion id="175BF29A163517E295DCFE28040C6B08" country="China" name="Yunnan">Yunnan</collectingRegion>
[Ponsee, Kahkyen Hills,
<collectingRegion id="24224563CF6B99D1696F3E618C247E0A" country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCountry id="EE36704248A4B1B2515FCC9494263E6E" name="China">China</collectingCountry>
].
<treatmentCitation id="31C16C7C1AC4758DA441A39601E33E6D" author="Beddome CR" page="171" year="1906">
<bibRefCitation id="A07A86DB4502391FF8EAD9ED4488F6EF" DOI="10.1093/oxfordjournals.mollus.a066160" author="Beddome CR" firstAuthor="Beddome" page="171" refId="B8" refString="Beddome CR (1906) Notes on Indian and Ceylonese species of Glessula. The Journal of Molluscan Studies 6 (3): 160172. https://doi.org/10.1093/oxfordjournals.mollus.a066160" year="1906">Beddome 1906: 171</bibRefCitation>
</treatmentCitation>
. Pilsbry 1909: 98, pl. 13, fig. 11.
<treatmentCitation id="2CB8583096D1988B18C67B5AF9E8C2B9" author="Gude GK" page="437" year="1914">
<bibRefCitation id="88B9974B24A9AEBDF0B06EC429003E64" DOI="10.5962/bhl.title.12891" author="Gude GK" firstAuthor="Gude" journalOrPublisher="Taylor &amp; Francis, London" page="437" refId="B42" refString="Gude GK (1914) Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, 520 pp. https://doi.org/10.5962/bhl.title.12891" title="Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma." year="1914">Gude 1914: 437</bibRefCitation>
</treatmentCitation>
, 438.
<treatmentCitation id="A1204C4413C5EB97EEE64B5B3DE8145F" author="Godwin-Austen HH" page="56" year="1920">
<bibRefCitation id="8C21A70F73E13B22065CB28AE3FCDB87" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="56" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920: 56</bibRefCitation>
</treatmentCitation>
, pl. 164, fig. 21.
</comment>
</taxonomicName>
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</paragraph>
<paragraph id="E2F088904EA44030DFBF548C3FAC6761">
<treatmentCitationGroup id="0431E3A7D0F5B0E98062CF6FD9071992">
<taxonomicName id="BD8DC621E595A6D69D54A17F97B44995" ID-CoL="7T7LX" baseAuthorityName="G. Nevill" baseAuthorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana" subGenus="Glessula">
<emphasis id="F9798CF34E01D5BEBCE0D7959A7F2C7D" italics="true">Stenogyra</emphasis>
(
<emphasis id="467C9BDAEEBAB3D30EC49FFB676D3605" italics="true">Glessula</emphasis>
)
<emphasis id="87B2A851F052746C7646B1A8E9095DA9" italics="true">blanfordiana</emphasis>
</taxonomicName>
<comment id="2CDD36D37C2C00BE0FC89DF23A91AB01">
<treatmentCitation id="13A94E6F93DBFA9343051B211A52E808" author="Nevill G" page="138" year="1881">
<bibRefCitation id="68A8D28696DF59A73E7DBE07EC1319F4" author="Nevill G" firstAuthor="Nevill" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" page="138" pagination="125-167" refId="B70" refString="Nevill G (1881) New or little-known Mollusca of the Indo-Malayan Fauna. Journal of the Asiatic Society of Bengal 50 (2): 125167. https://www.biodiversitylibrary.org/item/110180" title="New or little-known Mollusca of the Indo-Malayan Fauna." url="https://www.biodiversitylibrary.org/item/110180" volume="50" year="1881">Nevill 1881: 138</bibRefCitation>
</treatmentCitation>
, pl. 5, fig. 12.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWCCI" type="Type specimens">
<paragraph id="7E7F802985DC5D572A410F76BB5646C2">
<heading id="B6CF8F595E39F93D0D608860459F9202" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="142F32981785572014FC3C740EE601F9">
<emphasis id="6A889B701DA0223A518832E413D683BD" bold="true" italics="true">
<typeStatus id="DC9DDE4D915B521D550C74B8D4580107" type="syntype">Syntypes</typeStatus>
</emphasis>
<named-content id="BEF12DF9763D0575AC52DA30172895D4" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="836667DA00647F694B3EF96ED7E6EC74" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
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1988150 (2 shells; Fig.
<figureCitation id="C9042B80CF10A6F5A07DAB45D5E98A49" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">5 A, B</figureCitation>
) ex. Godwin-Austen collection from Ponsee,
<collectingRegion id="9BF0F17909D31DB8D1F2287FC2EC2F60" country="China" name="Yunnan">Yunnan</collectingRegion>
, W.
<collectingCountry id="7ED3A1F1D4084DFA0F8FD9C308A878F1" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EKDCI" type="material">
<paragraph id="FAA454757FFC90A224FC7BB4C95DB8F6">
<heading id="14397890FFDFC96AD702ECDBCD99CD0A" reason="title">Other material.</heading>
</paragraph>
<paragraph id="F7F7FAAC4F66FCF43BF225EF576714D2">
Aik Kham Cave, Taunggyi Township, Taunggyi District,
<collectingRegion id="F72862F543182079F90C632F08077FF5" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<collectingCountry id="8AD6C68D0284EB12FE14A6A3B170E11A" name="Myanmar">Myanmar</collectingCountry>
(
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<geoCoordinate id="0A8C0393482B98B89E684133B0CE395B" degrees="20" direction="north" minutes="49" orientation="latitude" precision="1" seconds="07.0" value="20.818611">20 ° 49 ' 07.0 &quot; N</geoCoordinate>
,
<geoCoordinate id="344FE784EF279F98855478FCDBBE7503" degrees="97" direction="east" minutes="13" orientation="longitude" precision="1" seconds="42.0" value="97.22833">97 ° 13 ' 42.0 &quot; E</geoCoordinate>
</named-content>
):
<named-content id="D11505BACFE56F573E6E06C6D64F8212" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University, Museum of Zoology">
<collectionCode id="0D4649F0A6615C3106BAA50E5291B042" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
13067 (3 shells; Fig.
<figureCitation id="194B6D2A491F365237A4821C5C9485E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">5 C, D</figureCitation>
).
</paragraph>
<caption id="63212FFE0F448276FCF19DE125B27782" ID-DOI="10.3897/zookeys.1208.116083.figure5" ID-arpha="E1FFC094-C885-5CF1-92A7-30B5241EC806" httpUri="https://binary.pensoft.net/fig/1104503" startId="F5">
<paragraph id="1B413A86C3FAE4FFF60F4B2725ECBE2D">
<label id="12B493CE6E6414649F345578A9B5DA28">Figure 5.</label>
</paragraph>
<paragraph id="499A3CC4EB8A47EA58969D5D49DB60BC">
<emphasis id="1167036E88093F11E9EBB3065010917C" bold="true">A D</emphasis>
<taxonomicName id="732DCB9CF50BF1244D8B4A7B22CAAC07" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="D4DF2110672125FB26E59D68E96A915E" italics="true">Glessula blanfordiana</emphasis>
</taxonomicName>
<emphasis id="66B23D0D8E75E46DA480CCC262977ACB" bold="true">A, B</emphasis>
syntypes
<named-content id="25B3E1724174C7A02A45DAE59853275C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
1988150 from Ponsee, Yunnan, W. China and
<emphasis id="2699A00FB72155AC17ECD6F0D2B64F48" bold="true">C, D</emphasis>
specimen
<named-content id="846BA5063E9DC618786F24AFB00F333F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University, Museum of Zoology">CUMZ</named-content>
13067 from Taunggyi, Shan State, Myanmar
<emphasis id="35C58068DC6CA0DA98F5296A98233A1F" bold="true">E, F</emphasis>
<taxonomicName id="0FFFEC855D906501969E6C36B6102BFB" baseAuthorityName="Benson" baseAuthorityYear="1836" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="crassilabris">
<emphasis id="1D640FB241A2DFD26B5AE5A7588E4433" italics="true">Glessula crassilabris</emphasis>
</taxonomicName>
<emphasis id="45DDAC391A10DFEE2A9D46B3A67BE97B" bold="true">E</emphasis>
syntype
<abbrev content-type="institution" id="ABBRID0ETFCI" xlink_title="University Museum of Zoology Cambridge, Cambridge">UMZC</abbrev>
I. 102430 from Bengal (after
<bibRefCitation id="02F9FE94198ACBA6DD8373024F77F6CF" author="Preece" etAl="et al." firstAuthor="Preece" refId="B88" refString="Preece RC, White TS, Raheem DC, Ketchum H, Ablett J, Taylor H, Naggs F (2022) William Benson and the golden age of malacology in British India: Biography, illustrated catalogue and evaluation of his molluscan types. Tropical Natural History (Supplement 6): 1434." year="2022">Preece et al. 2022</bibRefCitation>
: fig. 51 e) and
<emphasis id="519C16FE3F11DD2746C9F320FB11F0E1" bold="true">F</emphasis>
specimen
<named-content id="BBA17296FBAB6B92F8E173F0865FF715" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">NHMUK</named-content>
from Cherrapoonji (M 10604 / 2).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EIGCI" type="description">
<paragraph id="5FC6E9A36361ACA4A9C7C0BBA8A08A24">
<heading id="C1BC4F53BA7165D60053181EEC020F2F" reason="title">Description.</heading>
</paragraph>
<paragraph id="B51F7FEDB5DC55788AD2A7F357CBD892">Shell ovately conical, solid, brownish colour, and with 66 ½ whorls. Apex rounded; protoconch ~ 2 whorls with weak radial striations; subsequent whorls with equally and widely spaced ribs, more raised, and coarser near suture of last whorl. Spire regularly attenuated, slightly turreted; whorls convex; suture deep; last whorl largest. Aperture obliquely and narrowly ovate; peristome white and thickened; columella short, concave, slightly twisted and abruptly truncated. Umbilicus closed.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENGCI" type="distribution">
<paragraph id="1E4CA3A2BD5A10D617DB7FD86DBA1541">
<heading id="0967C8CC26BA1BA5592B63A266065CB4" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="939CE7F202C64EAEEE00ADFF9B41092C">
This species was originally described from
<collectingRegion id="514F2BBFA5919D11D323E45775A28B92" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<collectingCountry id="294458EFCF985122B07442C96335E8FF" name="China">China</collectingCountry>
, and subsequently reported from Bhamo,
<collectingRegion id="9E735AF899DE532A20CBEF96C2A0F810" country="Myanmar" name="Kachin">Kachin State</collectingRegion>
,
<collectingCountry id="9AEA3A9E53408C20EB3B6C19A750CD18" name="Myanmar">Myanmar</collectingCountry>
(
<bibRefCitation id="21034A39A737E867C5CD217C6B2407F4" author="Nevill" firstAuthor="Nevill" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="125-167" refId="B70" refString="Nevill G (1881) New or little-known Mollusca of the Indo-Malayan Fauna. Journal of the Asiatic Society of Bengal 50 (2): 125167. https://www.biodiversitylibrary.org/item/110180" title="New or little-known Mollusca of the Indo-Malayan Fauna." url="https://www.biodiversitylibrary.org/item/110180" volume="50" year="1881">Nevill 1881</bibRefCitation>
). In this study, new material has been collected from
<collectingRegion id="FCD9DD3822D9672383C15C63FAC10A67" country="Myanmar" name="Shan">Shan State</collectingRegion>
, which borders Yunnan to the east and Bhamo to the north.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXGCI" type="remarks">
<paragraph id="171F28D078A21628DA923F963AF442BD">
<heading id="D9AA33CC94AC0E0E70AE59825F93C396" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4D467BA58DB26D078518B49BE1145029">
When nominating this species,
<bibRefCitation id="6BE8407E7852D4954F6A761C3AB2998A" author="Nevill G" firstAuthor="Nevill" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" page="26" pagination="14-41" refId="B68" refString="Nevill G (1877) List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species. Journal of the Asiatic Society of Bengal 46 (2): 1441. https://www.biodiversitylibrary.org/item/110105" title="List of the Mollusca brought back by Dr. J. Anderson from Yunnan and Upper Burma, with descriptions of new species." url="https://www.biodiversitylibrary.org/item/110105" volume="46" year="1877">Nevill (1877: 26)</bibRefCitation>
compared it with
<taxonomicName id="53F8DDB4F995830542840EDDFE51DF04" authority="(Blanford, 1865)" baseAuthorityName="Blanford" baseAuthorityYear="1865" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="peguensis">
<emphasis id="16BA8FA9A2CBF29BC2219151DDCA3EBE" italics="true">G. peguensis</emphasis>
(
<bibRefCitation id="CA79FEC14083FFCA81FC0EBBEFAA50B5" author="Blanford WT" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="66-105" refId="B16" refString="Blanford WT (1865) Contributions to Indian Malacology, No. 5. Description of new land shells from Arakan, Pegu, and Ava; with notes on the distributions of described species. Journal of the Asiatic Society of Bengal 34 (2): 66 - 105. https: // www. biodiversitylibrary. org / item / 114408" title="Contributions to Indian Malacology, No. 5. Description of new land shells from Arakan, Pegu, and Ava; with notes on the distributions of described species." url="https://www.biodiversitylibrary.org/item/114408" volume="34" year="1865">Blanford, 1865</bibRefCitation>
)
</taxonomicName>
from
<collectingRegion id="CA96E9EE8AD83C979AB13FB6A21B3B28" country="Myanmar" name="Bago">Bago</collectingRegion>
and stated that
<taxonomicName id="6210961E528C09774B430CADEF32C771" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="E29CD665C0D25BF9CB8F165DA40B1648" italics="true">G. blanfordiana</emphasis>
</taxonomicName>
has slender (less convex) whorls, wavy or curving, and stronger radial ribs. Nevertheless, the type specimens of these two species,
<taxonomicName id="3AB3A85A098547EB0F5403FF249C9659" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="E2504271875E5A4C40E0639D4B31DF79" italics="true">G. blanfordiana</emphasis>
</taxonomicName>
(height
<quantity id="0BEFE1792B67D279C8504DF2EC15214C" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
, Fig.
<figureCitation id="E96BBD887C46E32D5A8CDD2C9D8ECD8C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A D Glessula blanfordiana A, B syntypes NHMUK 1988150 from Ponsee, Yunnan, W. China and C, D specimen CUMZ 13067 from Taunggyi, Shan State, Myanmar E, F Glessula crassilabris E syntype UMZC I. 102430 from Bengal (after Preece et al. 2022: fig. 51 e) and F specimen NHMUK from Cherrapoonji (M 10604 / 2)." figureDoi="10.3897/zookeys.1208.116083.figure5" httpUri="https://binary.pensoft.net/fig/1104503">5 A, B</figureCitation>
) and
<taxonomicName id="D47C8BC211A7ADA464CB59A41853F24E" baseAuthorityName="Blanford" baseAuthorityYear="1865" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="peguensis">
<emphasis id="838402F59A3A4629453C2CAE286D6601" italics="true">G. peguensis</emphasis>
</taxonomicName>
(height ~
<quantity id="1D0C578C74B28CDC2299D49274D4FE51" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
, Fig.
<figureCitation id="5BC590DFCC7DA61542560498E2B969A1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A Glessula orophila, specimen NHMUK ex. Blanford collection from Mahableshwar B, C Glessula peguensis, specimen NHMUK 1909.3. 15.8 from Thyet-Myo D, E Glessula perlevis, syntypes NHMUK 1906.5. 5.89 from Shan Hills, Burma F Glessula ponsiensis, syntypes NHMUK 1989042 from Ponsee." figureDoi="10.3897/zookeys.1208.116083.figure10" httpUri="https://binary.pensoft.net/fig/1104508">10 B, C</figureCitation>
), exhibit considerable similarity in terms of shell shape and shell size. Therefore, more specimens from the
<collectingRegion id="105BC06E73547D755AD733BF3B373CED" country="Myanmar" name="Bago">Bago Region</collectingRegion>
are necessary to ascertain whether the difference in shell sculptures represents variations or are taxonomically informative characters separating these two species.
</paragraph>
<paragraph id="CB790458CAD0554FDDCC8389A38B12FB">
<taxonomicName id="2165E5AA46CEF51793303F8BC328E092" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="1FDCFB6A324FC80FBD7794030249D596" italics="true">Glessula blanfordiana</emphasis>
</taxonomicName>
examined herein are noticeably distinct from their congeners recorded in
<collectingRegion id="9AF63A9CBF6EB14D953D1121569A9B60" country="Myanmar" name="Shan">Shan State</collectingRegion>
and
<collectingRegion id="18815EE7EE3446AA5A90B1942E1C072F" country="Myanmar" name="Mandalay">Mandalay Region</collectingRegion>
, namely
<taxonomicName id="E36C4EC7167E578B56A65B20CEB83430" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="feddeni">
<emphasis id="CBEEC8AD053B17EC2ABB5E62D0609E7D" italics="true">G. feddeni</emphasis>
<bibRefCitation id="3110A5ADC80ACA8C3C600EC752FD7E97" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="5A918C46E68A7F6A8509E0438E7D5655" authority="von Mollendorff, 1899" authorityName="von Mollendorff" authorityYear="1899" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="latestriata">
<emphasis id="FA0EAF582231C0A45C8FB7A15D43EB21" italics="true">G. latestriata</emphasis>
<bibRefCitation id="1F772E6023C46F96F48B37E61CBD0760" author="von Möllendorff OF" refId="B119" refString="von Mollendorff OF (1899) Neue arten aus Hinterindien. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 31: 165, 166. https: // biodiversitylibrary. org / page / 28228787" url="https://biodiversitylibrary.org/page/28228787" year="1899">von Möllendorff, 1899</bibRefCitation>
</taxonomicName>
. Specifically,
<taxonomicName id="04BB45AA6D657EA556D3935CA007C378" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="2AE5E4F2C6D4862A8AFD861867389828" italics="true">G. blanfordiana</emphasis>
</taxonomicName>
has a shorter and bluntly attenuated spire, more bulging whorls, deeper suture, and stronger and densely radial ribbed sculptures. In contrast, the other species exhibit higher and pointed spires, more flattened whorls, and smooth to radial grooves sculptures (see Fig.
<figureCitation id="59C274387C6F7C46C794AC7E1F676EA0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Schematic shell morphology of the subulinid snails A protoconch sculptures as recognised in this study: smooth and striate B crenulate suture C columella shape as recognised in this study: straight, truncate and fold D shell surface sculpture of the Glessula as recognised in this study: smooth, groove, and ribbed." figureDoi="10.3897/zookeys.1208.116083.figure2" httpUri="https://binary.pensoft.net/fig/1104500">2 D</figureCitation>
).
</paragraph>
<paragraph id="3277ED859630BEA4EC5F4D3792F16476">
<taxonomicName id="D438B2D42824B142DA2D782340672144" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="E3504AF38174C72BD4EF16DEB29C9841" italics="true">Glessula blanfordiana</emphasis>
</taxonomicName>
was described based on
<specimenCount id="7EE4E73F756D871D2DDE78BD1D2ED4DD" count="2" type="generic">two specimens</specimenCount>
collected by J. Anderson during the Expedition to Yunnan and Upper
<collectingCountry id="50A20FAB8464A0918459E4EE9A8467BD" name="Myanmar">Burma</collectingCountry>
. The original description was very brief, without shell dimensions and illustrations. However, under the note of
<taxonomicName id="DB19BC47BF4E50BE38687DC19DFA5CAC" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ponsiensis">
<emphasis id="6B2FFE47314B339CECB1BFA1DDE357AD" italics="true">G. ponsiensis</emphasis>
<bibRefCitation id="E58E8CE1EF09743DA92A8330751D7360" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
, states, The type from Ponsee, Yunnan (Plate CLXIV. fig. 20, apex) [=
<taxonomicName id="29EB46E2505ADED4D27BB9B6C52D2344" authorityName="Nevill" authorityYear="1877" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="blanfordiana">
<emphasis id="F007C3716A7F8A900F31DD9BFFB02EDB" italics="true">G. blanfordiana</emphasis>
</taxonomicName>
], has been sent me from Calcutta by the Director of the Zoological Survey of
<collectingCountry id="E303AA754BA9A0C6119FB2E09D4F937B" name="India">India</collectingCountry>
, Dr. N. Annandale … (
<bibRefCitation id="A591C07C535E7BB868A2E195F951B14D" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="56" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920: 56</bibRefCitation>
). The
<named-content id="0A5A8CDBE221E47DAE7573CDC7003AB6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="252678D3B9E0A82900E46C3B124BA746" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
type collections contain a single lot with an original label stating Type from Dr. J. Anderson, and collection locality from Ponsee. These
<specimenCount id="C82EAF4591CF27A65D9953DE73B8582B" count="2" type="generic">two specimens</specimenCount>
match well with the species description; both are figured herein for the first time. It is possible that N. Annandale may have gifted this type specimen lot to Godwin-Austen for comparison with other species, as was the case with other specimens from the Zoological Survey of
<collectingCountry id="E5BA2C4662932CA4626AC19187A2AE1D" name="India">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>