179 lines
35 KiB
XML
179 lines
35 KiB
XML
<document id="18BD1D8E37AB21B8545F4B3BAFCA7569" ID-CLB-Dataset="46931" ID-DOI="10.5281/zenodo.199891" ID-GBIF-Dataset="c1233bff-9caa-4381-8020-3c231030faff" ID-ISSN="1175-5326" ID-Zenodo-Dep="199891" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460349160855" checkinUser="plazi" docAuthor="Wu, Liang, Gao, Jian-Jun & Chen, Hong-Wei" docDate="2010" docId="DC4524176E17FFC30CA24EECFAF8FBFB" docLanguage="en" docName="zt02721p054.pdf" docOrigin="Zootaxa 2721" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Stegana (Steganina) biprotrusa" docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="207C5C6F6E17FFC20C354878FFE4FFC2" masterDocTitle="The Stegana (Steganina) biprotrusa species group from the Oriental Region (Diptera: Drosophilidae)" masterLastPageNumber="54" masterPageNumber="47" pageNumber="47" updateTime="1698293612910" updateUser="ExternalLinkService">
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<mods:titleInfo id="4EAA79C630C0A6AE3525EA37D3AE6843">
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<mods:title id="67FAFBCE6EB46DF6FFC04648E7ED6FC9">The Stegana (Steganina) biprotrusa species group from the Oriental Region (Diptera: Drosophilidae)</mods:title>
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<mods:name id="14964F6AABB835CC3CBA72A7EBBD1B92" type="personal">
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<mods:namePart id="BA1893AB6763854B913617EB5B6FECE0">Wu, Liang</mods:namePart>
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<mods:namePart id="F11F988C422ECAC093D1AC6662FD40FD">Gao, Jian-Jun</mods:namePart>
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<mods:namePart id="5705F8D9DD74433D2A8EBE71B504FA83">Chen, Hong-Wei</mods:namePart>
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<mods:typeOfResource id="AAA3F03279DBD73FA70484DFEDC8D554">text</mods:typeOfResource>
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<mods:title id="86C7489829DC980C78EBBACCA5D69F19">Zootaxa</mods:title>
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<mods:part id="E03A756A4BA2B0DAF302FB6C9512D5BC">
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<mods:date id="247FA43A954591475C5C0477C56ABA2E">2010</mods:date>
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<mods:detail id="43B0D0D20D0E5AFFD73A7E29564585D5" type="volume">
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<mods:number id="A046B142F2C8A7F7FA5D9B36CCB2ECF4">2721</mods:number>
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<mods:classification id="711D329223B751701EC8225EF5D0E50F">journal article</mods:classification>
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<mods:identifier id="28CC23B391FD379524532BC2AC872345" type="CLB-Dataset">46931</mods:identifier>
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<mods:identifier id="E96F9583F8EAE9D6A74EED3318C3EFF6" type="DOI">10.5281/zenodo.199891</mods:identifier>
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<treatment id="DC4524176E17FFC30CA24EECFAF8FBFB" ID-DOI="http://doi.org/10.5281/zenodo.6209200" ID-GBIF-Taxon="119407690" ID-Zenodo-Dep="6209200" LSID="urn:lsid:plazi:treatment:DC4524176E17FFC30CA24EECFAF8FBFB" httpUri="http://treatment.plazi.org/id/DC4524176E17FFC30CA24EECFAF8FBFB" lastPageId="1" lastPageNumber="48" pageId="0" pageNumber="47">
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<subSubSection id="1CF6C68A6E17FFC20CA24EECFD42F96F" box="[151,678,1684,1709]" pageId="0" pageNumber="47" type="nomenclature">
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<paragraph id="545395016E17FFC20CA24EECFD42F96F" blockId="0.[151,678,1684,1709]" box="[151,678,1684,1709]" pageId="0" pageNumber="47">
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<heading id="0F1B226D6E17FFC20CA24EECFD42F96F" bold="true" box="[151,678,1684,1709]" fontSize="10" level="2" pageId="0" pageNumber="47" reason="2">
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<emphasis id="669849136E17FFC20CA24EECFD42F96F" bold="true" box="[151,678,1684,1709]" pageId="0" pageNumber="47">
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<taxonomicName id="93ECEE826E17FFC20CA24EECFE1BF96F" ID-CoL="8F4S8" box="[151,511,1684,1709]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="biprotrusa" subGenus="Steganina">
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<emphasis id="669849136E17FFC20CA24EECFF17F96F" bold="true" box="[151,243,1684,1709]" italics="true" pageId="0" pageNumber="47">Stegana</emphasis>
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(
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<emphasis id="669849136E17FFC20D364EECFE91F96F" bold="true" box="[259,373,1684,1709]" italics="true" pageId="0" pageNumber="47">Steganina</emphasis>
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)
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<emphasis id="669849136E17FFC20DB24EECFE1BF96F" bold="true" box="[391,511,1684,1709]" italics="true" pageId="0" pageNumber="47">biprotrusa</emphasis>
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</taxonomicName>
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species group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1CF6C68A6E17FFC20CA24EA4FD6AF8DA" pageId="0" pageNumber="47" type="diagnosis">
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<paragraph id="545395016E17FFC20CA24EA4FD6AF8DA" blockId="0.[151,1436,1756,2032]" pageId="0" pageNumber="47">
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<emphasis id="669849136E17FFC20CA24EA4FEF5F937" bold="true" box="[151,273,1756,1781]" pageId="0" pageNumber="47">Diagnosis.</emphasis>
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Articulating-to-aedeagus plate aedeagus developed, bifurcated, separated from hypandrium and mostly sclerotized (
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<figureCitation id="CCD789846E17FFC20D2A4F78FE67F8DA" box="[287,387,1792,1816]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="0" pageNumber="47">Figs 6, 7</figureCitation>
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,
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<figureCitation id="CCD789846E17FFC20DA54F78FE3CF8DA" box="[400,472,1792,1816]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="0" pageNumber="47">12, 13</figureCitation>
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,
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<figureCitation id="CCD789846E17FFC20DD64F78FDC8F8DA" box="[483,556,1792,1816]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="0" pageNumber="47">19, 20</figureCitation>
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,
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<figureCitation id="CCD789846E17FFC20E0D4F78FD99F8DA" box="[568,637,1792,1816]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="0" pageNumber="47">25, 26</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="1CF6C68A6E17FFC30CF24F5CFC1FFC0F" lastPageId="1" lastPageNumber="48" pageId="0" pageNumber="47" type="description">
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<paragraph id="545395016E17FFC30CF24F5CFEAAFD6F" blockId="0.[151,1436,1756,2032]" lastBlockId="1.[151,1436,151,1081]" lastPageId="1" lastPageNumber="48" pageId="0" pageNumber="47">
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<emphasis id="669849136E17FFC20CF24F5CFEBCF8FF" bold="true" box="[199,344,1828,1853]" pageId="0" pageNumber="47">Description.</emphasis>
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Male: Eyes red. Ocellar triangle black, with a pair of small setae above ocellar setae. Postvertical setae slightly behind vertex ridge. Frons and face rectangular in profile. Frons black, with sporadic minute setulae. Fronto-orbital plate brownish yellow above, black below. Proclinate orbital setae distinctly nearer to ptilinal fissure than to inner vertical setae. First flagellomere black. Face brown on upper 2/3, yellow on lower margin, broadened ventrally; facial carina narrow and developed on upper part like as in
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<taxonomicName id="93ECEE826E17FFC20F834FCDFAF6F80F" box="[950,1298,1972,1997]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="47" phylum="Arthropoda" rank="species" species="nigrifrons" subGenus="Steganina">
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<emphasis id="669849136E17FFC20F834FCDFBEBF80E" box="[950,1039,1973,1996]" italics="true" pageId="0" pageNumber="47">Stegana</emphasis>
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(
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<emphasis id="669849136E17FFC208174FCDFB74F80E" box="[1058,1168,1973,1996]" italics="true" pageId="0" pageNumber="47">Steganina</emphasis>
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)
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<emphasis id="669849136E17FFC208964FCDFAF6F80F" box="[1187,1298,1973,1997]" italics="true" pageId="0" pageNumber="47">nigrifrons</emphasis>
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</taxonomicName>
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de Meijere, 1991 (
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<bibRefCitation id="307DE8F06E17FFC20CE84FA0FE32F832" author="Okada" box="[221,470,2008,2032]" pageId="0" pageNumber="47" refString="Okada T. (1971) A revision and taxometric analysis of the genus Stegana Meigen of Japan and adjacent countries (Diptera, Drosophilidae). Mushi, 45, 81 - 99." type="journal article" year="1971">
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Okada 1971,
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<figureCitation id="CCD789846E17FFC20D404FA0FE32F832" box="[373,470,2008,2032]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="0" pageNumber="47">Figs 7, 8</figureCitation>
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</bibRefCitation>
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). Clypeus dark brown medially, yellow laterally. Palpus yellow, with 4–5 longer setae distally and several shorter setae basally. Gena yellow, broad (ch/o ≥ 0.15). Vibrissa prominent; other orals small. Occiput brown. Scutum flat, brownish yellow, white-yellow on prothorax. Postpronotal lobe white-yellow, with 1– 2 longer and a few small setae. Prescutellar setae 1 pair. Acrostichal setulae in approximately 10 irregular rows. Pleura with 1 black longitudinal stripe above (running from propleuron to base of halter). Katepisternum usually entirely white. Basisternum brown to black. Scutellum usually black, yellow at tip; basal scutellar setae divergent; apical scutellars cruciate. Wing brown anteriorly, paler posteriorly, not paler at tip. Basal medial-cubital crossvein present; C1 setae 2, less differentiated. Costal vein with 6–8 minute spinules on ventral surface between R2+3 and R4+5. R2+3 markedly curved to costa at tip; R4+5 and M1 convergent distally. Halter grey-brown basally, white distally. Legs white to yellowish, dark brown at fore knee, distal 1/2 of mid and hind femora and basal 1/2 of mid and hind tibiae. Fore femur with 1 row of 4, 5 suberect setae on posterior surface and 1 row of 5 suberect setae on ventral surface. Apical seta present on mid tibia. Preapical dorsal setae present on all tibiae. Mid tibia with 5 strong, suberect setae on basal part of dorsal surface. Mid and hind tarsomeres with 2 and 1 rows of minute cuneiform setulae on anteroventral surface, respectively; fore and hind first tarsomeres slightly shorter than the rest combined; mid first tarsomere longer than the rest combined. Abdominal tergites nearly entirely black; sternites broadened, wider than long.
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</paragraph>
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<paragraph id="545395016E16FFC30CF24AC1FC1FFC0F" blockId="1.[151,1436,151,1081]" pageId="1" pageNumber="48">
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<emphasis id="669849136E16FFC30CF24AC1FE99FD13" box="[199,381,697,721]" italics="true" pageId="1" pageNumber="48">Male terminalia</emphasis>
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: Epandrium pubescent except for anterior and ventral margins, with dense setae on dorsal to posterolateral portion of each body-side (
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<figureCitation id="CCD789846E16FFC30E524AA5FD54FD37" box="[615,688,733,757]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 4</figureCitation>
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,
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<figureCitation id="CCD789846E16FFC30E884AA5FD3FFD37" box="[701,731,733,757]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">10</figureCitation>
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,
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<figureCitation id="CCD789846E16FFC30EDD4AA5FCE2FD37" box="[744,774,733,757]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">16</figureCitation>
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,
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<figureCitation id="CCD789846E16FFC30F274AA5FCCBFD37" box="[786,815,733,757]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">23</figureCitation>
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). Cercus setigerous and pubescent (
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<figureCitation id="CCD789846E16FFC308F14AA5FAEAFD37" box="[1220,1294,733,757]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 4</figureCitation>
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,
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<figureCitation id="CCD789846E16FFC3092F4AA5FADCFD37" box="[1306,1336,733,757]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">10</figureCitation>
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,
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<figureCitation id="CCD789846E16FFC309704AA5FA87FD37" box="[1349,1379,733,757]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">16</figureCitation>
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,
|
||
<figureCitation id="CCD789846E16FFC309454AA5FA6BFD37" box="[1392,1423,733,757]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">23</figureCitation>
|
||
). Surstylus separated from epandrium, with numerous serrate processes on basal 3/4, one small prensiseta (but absent in
|
||
<taxonomicName id="93ECEE826E16FFC30D304B5EFE5FFCFC" box="[261,443,806,830]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="otocondyloda" status="sp. nov.">
|
||
<emphasis id="669849136E16FFC30D304B5EFEF7FCFF" box="[261,275,806,829]" italics="true" pageId="1" pageNumber="48">S</emphasis>
|
||
.
|
||
<emphasis id="669849136E16FFC30D164B5EFE5FFCFC" box="[291,443,806,830]" italics="true" pageId="1" pageNumber="48">otocondyloda</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7DABF4686E16FFC30DF14B5EFDFAFCFF" box="[452,542,805,829]" pageId="1" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and numerous setae on distal 1/4, lacking pubescence (
|
||
<figureCitation id="CCD789846E16FFC308974B5DFAEEFCFF" box="[1186,1290,805,829]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 4, 9</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC3092D4B5DFA86FCFF" box="[1304,1378,805,829]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">16, 17</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC309454B5DFA69FCFF" box="[1392,1421,805,829]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">23</figureCitation>
|
||
). Tenth sternite connected with surstyli basolaterally. Hypandrium broadened on anterior margin, sometimes concaved medially (
|
||
<figureCitation id="CCD789846E16FFC30D4B4B15FE2DFC47" box="[382,457,877,901]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30DED4B15FE12FC47" box="[472,502,877,901]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">12</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30E304B15FDC7FC47" box="[517,547,877,901]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">19</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30E074B15FDB6FC47" box="[562,594,877,901]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">25</figureCitation>
|
||
). Paramere absent. Gonopods strongly sclerotized, with serrate processes distally (
|
||
<figureCitation id="CCD789846E16FFC30CCF4BE9FEBBFC6B" box="[250,351,913,937]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 8, 9</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30D594BE9FE51FC6B" box="[364,437,913,937]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">14, 15</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30DF74BE9FDEEFC6B" box="[450,522,913,937]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">21, 22</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30E224BE9FDBAFC6B" box="[535,606,913,937]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">27, 28</figureCitation>
|
||
). Aedeagus developed, basally fused to aedeagal apodeme (
|
||
<figureCitation id="CCD789846E16FFC309334BE9FA88FC6B" box="[1286,1388,913,937]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 6, 7</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC3094C4BE9FF51FC0F" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">12, 13</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30CF44BCDFEEDFC0F" box="[193,265,949,973]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">19, 20</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30D204BCDFEBBFC0F" box="[277,351,949,973]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">25, 26</figureCitation>
|
||
); apodeme usually strong (
|
||
<figureCitation id="CCD789846E16FFC30EBE4BCDFD14FC0F" box="[651,752,949,973]" captionStart="FIGURES 4 – 9" captionStartId="2.[151,269,1855,1879]" captionTargetBox="[160,1412,743,1771]" captionTargetId="figure@2.[151,1423,725,1783]" captionTargetPageId="2" captionText="FIGURES 4 – 9. Stegana (Steganina) acantha sp. nov., 3: 4. Epandrium (epand), cercus (cerc) and surstylus (sur) (lateral view); 5. 10 th sternite (ventral view); 6, 7. hypandrium (hypd), aedeagus (aed), articulating-to-aedeagus plate (ap aed) and aedeagal apodeme (aed a) (ventral and lateral views); 8, 9. gonopods (ventral and lateral views) (scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199892/files/figure.png" pageId="1" pageNumber="48">Figs 6, 7</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30EC94BCDFCA0FC0F" box="[764,836,949,973]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,269,1290,1314]" captionTargetBox="[159,1428,494,1257]" captionTargetId="figure@4.[151,1436,483,1288]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Stegana (Steganina) ancistrophylla sp. nov., 3: 10. Epandrium, cercus and surstylus; 11. 10 th sternite; 12, 13. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 14, 15. gonopods (for orientation and abbreviations see figs 4 – 9; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199893/files/figure.png" pageId="1" pageNumber="48">12, 13</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30F654BCDFC7DFC0F" box="[848,921,949,973]" captionStart="FIGURES 16 – 22" captionStartId="5.[151,269,1274,1298]" captionTargetBox="[160,1425,482,1244]" captionTargetId="figure@5.[150,1436,469,1279]" captionTargetPageId="5" captionText="FIGURES 16 – 22. Stegana (Steganina) langufoliacea sp. nov., 3: 16. Epandrium, cercus and surstylus; 17. surstylus (ventral view); 18. 10 th sternite; 19, 20. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 21, 22. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199894/files/figure.png" pageId="1" pageNumber="48">19, 20</figureCitation>
|
||
,
|
||
<figureCitation id="CCD789846E16FFC30F914BCDFC0AFC0F" box="[932,1006,949,973]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,269,986,1010]" captionTargetBox="[161,1415,196,945]" captionTargetId="figure@6.[150,1436,168,964]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Stegana (Steganina) otocondyloda sp. nov., 3: 23. Epandrium, cercus and surstylus; 24. 10 th sternite; 25, 26. hypandrium, aedeagus, articulating-to-aedeagus plate (ap aed) and aedeagal apodeme; 27, 28. gonopods (for orientation and abbreviations see figs 1 – 6; scale bars: 0.1 mm)." httpUri="https://zenodo.org/record/199895/files/figure.png" pageId="1" pageNumber="48">25, 26</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CF6C68A6E16FFC30CF24BA2FAF8FBFB" pageId="1" pageNumber="48" type="discussion">
|
||
<paragraph id="545395016E16FFC30CF24BA2FAF8FBFB" blockId="1.[151,1436,151,1081]" pageId="1" pageNumber="48">
|
||
<emphasis id="669849136E16FFC30CF24BA2FEDFFC30" bold="true" box="[199,315,986,1010]" pageId="1" pageNumber="48">Remarks.</emphasis>
|
||
In the subgenus
|
||
<taxonomicName id="93ECEE826E16FFC30E394BA2FD9EFC33" box="[524,634,986,1009]" class="Insecta" family="Drosophilidae" genus="Steganina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="669849136E16FFC30E394BA2FD9EFC33" box="[524,634,986,1009]" italics="true" pageId="1" pageNumber="48">Steganina</emphasis>
|
||
</taxonomicName>
|
||
, usually paramere is small, and fused with hypandrium; sometimes undistinguishable or absent. The paramere of
|
||
<taxonomicName id="93ECEE826E16FFC30EAF4B86FCCFFBD7" box="[666,811,1021,1045]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="48" phylum="Arthropoda" rank="species" species="biprotrusa">
|
||
<emphasis id="669849136E16FFC30EAF4B86FD4CFBD7" box="[666,680,1022,1045]" italics="true" pageId="1" pageNumber="48">S</emphasis>
|
||
.
|
||
<emphasis id="669849136E16FFC30E834B85FCCFFBD7" box="[694,811,1021,1045]" italics="true" pageId="1" pageNumber="48">biprotrusa</emphasis>
|
||
</taxonomicName>
|
||
is changed to “articulating plate of aedeagus” based on the following two characters: lack setulae; separated from hypandrium, but fused to aedeagal apodeme.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |