185 lines
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185 lines
25 KiB
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<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="6F8DB999B63E5743A58ADE18F878CB59" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina arula Benson 1856" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:titleInfo id="04E2EA0802AF3ACD982D0378956E5D49">
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<mods:title id="D5E0D300978AD027C00E27B847499FCC">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name id="ABBD80E4BA0524EC517998D8867CF5D8" type="personal">
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<mods:roleTerm id="784F03BD9227C559171B7ABCB8CB3884">Author</mods:roleTerm>
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<mods:namePart id="C496DA4CC64C7AA8215E1F181FB47FED">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="296741B371E9A8CDC74957DD39388715" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="66A8BAE380C8171EE36B62F94038BEF1">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="C446E91D413F83784BE7CC76044D735D" type="personal">
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<mods:roleTerm id="BABCBE56BAC679AE223F0375AFF21A3A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="494C9A3E247CE1768A856E0E55B9F896">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="157854A5FEAB315E7627582117A68FA4">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="A94C5E541965781D300F6807BC77EA86" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="3E8AD1CEBF605F455E26E7530A44D7B4">Panha, Somsak</mods:namePart>
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<mods:affiliation id="4FDB8A26C36A9C4EA2AC9EA0DB9146BE">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="35D040D27BEBE49D22379A1E616EA5B7" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="74CD6459C501F698E1D5C39E97A57E7A">text</mods:typeOfResource>
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<mods:title id="070CEA4DBB50F9C36404587ACCB4282B">ZooKeys</mods:title>
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<mods:date id="754C0FAB763C5BE834EDBFA0F2A2D107">2022</mods:date>
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<mods:number id="6C726E02BF941545026EA710EA39BB24">2022-08-25</mods:number>
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<mods:detail id="F966122E6DFC9D9D791203059996E43A" type="volume">
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<mods:number id="DF945BA03F602D4E006BACFD84C895E1">1119</mods:number>
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<mods:start id="58CAF0F54ACE0B2D60F2F96AB71BB8FA">1</mods:start>
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<mods:end id="1EB8E37B434EBD2BFF5A232576012082">115</mods:end>
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<mods:classification id="B4865F8A001AA52005593771C0F77E0E">journal article</mods:classification>
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<mods:identifier id="1B5A81A817A1747E787D0CF8AD495CD4" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier id="84B96D8A0A952A374584817CCEB387B1" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<treatment id="6F8DB999B63E5743A58ADE18F878CB59" ID-GBIF-Taxon="199838979" LSID="urn:lsid:plazi:treatment:6F8DB999B63E5743A58ADE18F878CB59" httpUri="http://treatment.plazi.org/id/6F8DB999B63E5743A58ADE18F878CB59" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="618C302329F91C58F468F32A01BF627F" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="E9F5817F5BB8A5381AF10370DCD816F9" pageId="0" pageNumber="1">
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<taxonomicName id="F41D708F4364F5ACB7C6565C88A7ACA1" ID-CoL="7SWXC" LSID="6F8DB999-B63E-5743-A58A-DE18F878CB59" authority="Benson, 1856" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">Pupina arula Benson, 1856</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="1C7FB43C5D8824FDB6E5B6D0284DD693" pageId="0" pageNumber="1" type="description">
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<paragraph id="0F6667FD1868670D4359B61B75E12014" pageId="0" pageNumber="1">
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<figureCitation id="B5D804AF3211B938116504180F775353" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">Figs 29A, B</figureCitation>
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<figureCitation id="F1707DAC13C7A9A770F8B92EC5F6BFE4" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D)." figureDoi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1">, 33D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="68D0966EFCEBE3CC6CD4E6A4CBEE63A3" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="7D81A844A76E110FB904AFF7FB7EAA12" pageId="0" pageNumber="1">
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<taxonomicName id="1F0760744033D0A063A0013B8F792125" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">Pupina arula</taxonomicName>
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Benson, 1856: 230. Type locality: ad Yunglaw, in valle Tenasserim [Tanintharyi Region, Myanmar].
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<bibRefCitation id="B14B921DB21A4D6BFA05590FB1303E19" author="Theobald, W" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="245 - 254" refId="B167" refString="Theobald, W, 1858. Notes on the distribution of some of the land and freshwater shells of India. Journal of the Asiatic Society of Bengal 26 (1857): 245 - 254" title="Notes on the distribution of some of the land and freshwater shells of India." volume="26" year="1858">Theobald 1858</bibRefCitation>
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[1857]: 247. Pfeiffer 1860: 141, pl. 37, figs 7-9. Hanley and Theobald 1870: 4, pl. 7, fig. 4.
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<bibRefCitation id="65A6AADAD07412964AB7ACC76061DBCA" author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve & Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve & Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
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:
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<taxonomicName id="5B6779FB4495FAE1FB26D81756B0E553" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
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, pl. 1, sp. 5.
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<bibRefCitation id="2D82D4072D54FF540C89FEA00B15567B" author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="336 - 340" refId="B26" refString="Crosse, H, 1879. Faune malacologique de Perak (Indo-Chine). Journal de Conchyliologie 27: 336 - 340" title="Faune malacologique de Perak (Indo-Chine)." volume="27" year="1879">Crosse 1879</bibRefCitation>
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: 340 (part). de Morgan 1885: 413 (part).
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<bibRefCitation id="A2CADD40C1B83CA0B3EEC51C982BCFDC" author="von Moellendorff, OF" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="0" pageNumber="1" pagination="299 - 316" refId="B190" refString="von Moellendorff, OF, 1887. The land shells of Perak. Journal of the Asiatic Society of Bengal 55 (1886): 299 - 316" title="The land shells of Perak." volume="55" year="1887">
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von
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<normalizedToken id="462BD8FB9D0377139DB7D5C959A655DC" originalValue="Möllendorff">Moellendorff</normalizedToken>
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1887
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</bibRefCitation>
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[1886]: 314 (part).
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<bibRefCitation id="07696908B4AC0CD51AA7E6A2A3B28506" author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
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: 37, 38, pl. 69, fig. 1, 1a.
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<bibRefCitation id="2BB463AFAD2B7848D618464FA54EF4BB" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
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: 88.
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<bibRefCitation id="6D1AA2026B4160FC70E74345DCED335F" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
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: fig. 5-13e.
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</paragraph>
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<paragraph id="5AFAD82AFC0DD46C8C4815DB57D2A48E" pageId="0" pageNumber="1">
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<taxonomicName id="3917B5E37FB19BF5AD458A607A7FE975" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina avula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="avula">Pupina avula</taxonomicName>
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[sic]-Sowerby I 1866:
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<taxonomicName id="E9E85D76E3FABA4EEDE8A333E93C544D" authorityName="L. Pfeiffer" authorityYear="1853" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="family">Pupinidae</taxonomicName>
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, pl. 3 (pl. 265),
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<taxonomicName id="1EE052306762265A5EA2064D3C70B2A1" authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
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, fig. 3.
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</paragraph>
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<paragraph id="532F0B8B3AC724C50269E6341A211E61" pageId="0" pageNumber="1">
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<taxonomicName id="4AB613DDDCD9FFAC9BCC6A2ADC16B178" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula" subGenus="Tylotoechus">Pupina (Tylotoechus) arula</taxonomicName>
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-
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<bibRefCitation id="456BD91BD8D1991D3A9C86309B12C857" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 307.
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<bibRefCitation id="1E39EBBBD9E964809A5DE1A25B9F8644" author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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: 193, 194 (part).
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<bibRefCitation id="D84DB9122ABAD7CCECFD4CF917BC3874" author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
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: 12, Doi Sutep [Doi Suthep Mountain, Chiang Mai Province, Thailand].
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<bibRefCitation id="B20894864698D44478E99ACEF0E0CFBD" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
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: 39.
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</paragraph>
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<paragraph id="E645A647E97827F0F88FF380FF8EDD4A" pageId="0" pageNumber="1">
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<taxonomicName id="848BC54F3C04BB4B9DA3C919F1A98307" authority="arula" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina arula subsp. arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="arula" subSpecies="arula">Pupina arula arula</taxonomicName>
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-
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<bibRefCitation id="2D8F6B67EFCC38F562145CA081892CB6" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
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: 40.
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</paragraph>
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</subSubSection>
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<subSubSection id="2C2E5CEC006E1010D76131C37CA3E567" pageId="0" pageNumber="1" type="type material">
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<paragraph id="88EE298086BB7E61964B4108181B3918" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="A6D2CDA109E98E097605469577A1F3DE" pageId="0" pageNumber="1">
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<emphasis id="C27C0733690D826F440FA08711D11DF9" bold="true" italics="true" pageId="0" pageNumber="1">Syntype</emphasis>
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UMZC I.103025 (1 shell; Figs
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<figureCitation id="294A40060397EC56B12EFFAF5C2C4660" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">29A</figureCitation>
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,
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<figureCitation id="F5A0D8903D511DE677927CF7AA0A455A" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. A Pupina bilabiata sp. nov., specimen CUMZ 12086 from Na San Temple, Surat Thani B, C Pupina godwinausteni sp. nov. B holotype CUMZ 12090 / 1 and C paratype CUMZ 12090 / 2 from Khao Wong Cave, Uthai Thani D Pupina arula, syntype UMZC I. 103025 " Ind " E Pupina mouhoti, possible syntype NHMUK ex. Cuming coll. from Camboja F Pupina vescoi, syntype NHMUK 1893.2.4.767 from Cochin China. Photo: J. Ablett, H. Taylor, NHM (D)." figureDoi="10.3897/zookeys.1119.85400.figure33" httpUri="https://binary.pensoft.net/fig/735394" pageId="0" pageNumber="1">33D</figureCitation>
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) from the R. McAndrew collection labelled "Bens. col., Ind".
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</paragraph>
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</subSubSection>
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<subSubSection id="D5C8CDDAFE4C86FE47B8E1A5BB4C6232" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="EE8EC8AB3C00CE0226698D9F3A8ED2E0" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="DE2DBDFBC8F4E9A9570F70E0CB9BF923" pageId="0" pageNumber="1">
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Specimen NHMUK 1888.12.4.109 (1 shell; Fig.
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<figureCitation id="5BBA96D11AB74ADBEA882A035AEDCF66" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Shells of Pupina arula species group from mainland Southeast Asia A, B Pupina arula A syntype UMZC I. 103025 and B specimen NHMUK 1888.12.4.109. C, D Pupina mouhoti, possible syntypes NHMUK ex. Cuming coll. E, F Pupina vescoi E syntype NHMUK 1893.2.4.767 and F specimen SMF 109956 / 1 G Pupina perakensis, lectotype SMF 109969 / 1 H Pupina excisa, lectotype SMF 110778 / 1 I-K Pupina exclamationis I, J syntypes MNHN-IM- 2000 - 35840 and K specimen NHMUK 1901.12.23.205 " forma minor ". Photo: J. Ablett, H. Taylor, NHM (A), A. Lardeur, P. Maestrati, MNHN (I, J)." figureDoi="10.3897/zookeys.1119.85400.figure29" httpUri="https://binary.pensoft.net/fig/735390" pageId="0" pageNumber="1">29B</figureCitation>
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) from Yunglaw,
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<collectingCountry id="41A97A84BCD186378D57194BCC89DD6C" name="Myanmar">Myanmar</collectingCountry>
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, the W. Theobald collection.
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</paragraph>
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</subSubSection>
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<subSubSection id="B46E3238FD8FD63D83D51027006BE997" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="410807F53BC3A06F9677E81DCDD3EAF7" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="3AD517006D76FE3243A46CD5C4839460" pageId="0" pageNumber="1">Shell ovate; last whorl ca. 80% of shell height. Shell surface matt. Apertural lip thickened but not expanded; apertural lip curved when observed from lateral view. Columellar tooth fin-shaped.</paragraph>
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</subSubSection>
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<subSubSection id="0CFB76E6FD68A00FC496036A3975D88E" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="5C1ED784EE261F2201A5EB6321ADB65D" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="838476D8126D97298EF7FBEF1C76F30E" pageId="0" pageNumber="1">
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<taxonomicName id="37E6596151DC7F2B33914D0DF935499A" authorityName="Benson" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
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<emphasis id="343698D43ACDA802C8ED873594243106" italics="true" pageId="0" pageNumber="1">Pupina arula</emphasis>
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</taxonomicName>
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can be distinguished from all other species in the
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<taxonomicName id="20A7A9536A409B1E2E0DC68EA54DD26E" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
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<emphasis id="FEB5EF7E630887E58E89646F76C7F090" italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
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</taxonomicName>
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species group from mainland Southeast Asia by a matt shell surface.
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</paragraph>
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</subSubSection>
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<subSubSection id="50E1AA86A2D8F6E606162006D4B328A7" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="410B95BF6001FBDB58FAC9E0D09C27F0" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="8C01878C5E0F6690BFD99A71EECFD06B" pageId="0" pageNumber="1">
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|
Myanmar and an uncertain record from northern Thailand (
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|
<bibRefCitation id="4D24778EF59065660A105EE974BE8807" author="Solem, A" journalOrPublisher="Spolia Zoologica Musei Hauniensis" pageId="0" pageNumber="1" pagination="1 - 110" refId="B146" refString="Solem, A, 1966. Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae. Spolia Zoologica Musei Hauniensis 24: 1 - 110" title="Some non-marine mollusks from Thailand, with notes on classification of the Helicarionidae." volume="24" year="1966">Solem 1966</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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|
<subSubSection id="8B2E4320D0DC84CE148443E53C0C07C9" pageId="0" pageNumber="1" type="remarks">
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|
<paragraph id="ED7AEC0E82520B1EC23B1ADB22E0DAE8" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="D138BEBCADA533C1C55686DCC13CD3ED" pageId="0" pageNumber="1">
|
|
No material of this species was found during this survey. The specimen of
|
|
<taxonomicName id="44AC2366D9EDDFA77075D0B197C332E5" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="arula">
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|
<emphasis id="11C6EB8E2AFA89B4454D65722CC97605" italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
|
|
</taxonomicName>
|
|
mentioned in
|
|
<bibRefCitation id="79A1211867E9D107E55B5E1B6AC2D401" author="Davison, GWH" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="233 - 248" refId="B31" refString="Davison, GWH, 1995. The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993-1994. Malayan Nature Journal 48: 233 - 248" title="The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia. In: Belum Expedition, Temengor, 1993 - 1994." volume="48" year="1995">Davison (1995</bibRefCitation>
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|
: 237) from Temengor dam, Perak, Malaysia possibly belongs to
|
|
<taxonomicName id="01F6A99408A75096C6A47B101DC02013" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="perakensis">
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|
<emphasis id="BCC52E83AA66A43F3593D71C2381DC00" italics="true" pageId="0" pageNumber="1">P. perakensis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
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</subSubSection>
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</treatment>
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</document> |