Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia Author Jirapatrasilp, Parin https://orcid.org/0000-0002-5591-6724 Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Sutcharit, Chirasak Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand jirasak4@yahoo.com Author Panha, Somsak Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand somsak.pan@chula.ac.th text ZooKeys 2022 2022-08-25 1119 1 115 http://dx.doi.org/10.3897/zookeys.1119.85400 journal article http://dx.doi.org/10.3897/zookeys.1119.85400 1313-2970-1119-1 A3BE91C6B79344E1A886A803BF104D8B F158A7C5261D52B69288A62F3C777CAF Pupina arula Benson, 1856 Figs 29A, B , 33D Pupina arula Benson, 1856: 230. Type locality: ad Yunglaw, in valle Tenasserim [Tanintharyi Region, Myanmar]. Theobald 1858 [1857]: 247. Pfeiffer 1860: 141, pl. 37, figs 7-9. Hanley and Theobald 1870: 4, pl. 7, fig. 4. Reeve 1878 : Pupinidae , pl. 1, sp. 5. Crosse 1879 : 340 (part). de Morgan 1885: 413 (part). von Moellendorff 1887 [1886]: 314 (part). Godwin-Austen 1897 : 37, 38, pl. 69, fig. 1, 1a. BEDO 2017 : 88. Sutcharit et al. 2018 : fig. 5-13e. Pupina avula [sic]-Sowerby I 1866: Pupinidae , pl. 3 (pl. 265), Pupina , fig. 3. Pupina (Tylotoechus) arula - Kobelt 1902 : 307. Gude 1921 : 193, 194 (part). Solem 1966 : 12, Doi Sutep [Doi Suthep Mountain, Chiang Mai Province, Thailand]. Hemmen and Hemmen 2001 : 39. Pupina arula arula - Maassen 2001 : 40. Type material examined. Syntype UMZC I.103025 (1 shell; Figs 29A , 33D ) from the R. McAndrew collection labelled "Bens. col., Ind". Other material examined. Specimen NHMUK 1888.12.4.109 (1 shell; Fig. 29B ) from Yunglaw, Myanmar , the W. Theobald collection. Diagnosis. Shell ovate; last whorl ca. 80% of shell height. Shell surface matt. Apertural lip thickened but not expanded; apertural lip curved when observed from lateral view. Columellar tooth fin-shaped. Differential diagnosis. Pupina arula can be distinguished from all other species in the P. arula species group from mainland Southeast Asia by a matt shell surface. Distribution. Myanmar and an uncertain record from northern Thailand ( Solem 1966 ). Remarks. No material of this species was found during this survey. The specimen of P. arula mentioned in Davison (1995 : 237) from Temengor dam, Perak, Malaysia possibly belongs to P. perakensis .