treatments-xml/data/62/16/B3/6216B3248C70FFAE58B9FA97FC0491BA.xml

<document id="56BB9FDDA2EB3123ADFF1427DFBFB5FA" ID-DOI="10.5281/zenodo.187941" ID-GBIF-Dataset="075ae537-5ba7-4f84-a494-03dc91deaeed" ID-ISSN="1175-5326" ID-Zenodo-Dep="187941" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461077613966" checkinUser="plazi" docAuthor="Ota, Yuzo &amp; Hirose, Euichi" docDate="2009" docId="6216B3248C70FFAE58B9FA97FC0491BA" docLanguage="en" docName="zt02114p060.pdf" docOrigin="Zootaxa 2114" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gnathia maculosa Ota &amp; Hirose, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="9E2FCB5C8C71FFA9582EFFD5FFDC922A" masterDocTitle="Description of Gnathia maculosa and a new record of Gnathia trimaculata (Crustacea, Isopoda, Gnathiidae), ectoparasites of elasmobranchs from Okinawan coastal waters" masterLastPageNumber="60" masterPageNumber="50" pageNumber="51" updateTime="1698586747131" updateUser="plazi">
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<mods:title id="746F83185A45A7296DD4C754DF3188F8">Description of Gnathia maculosa and a new record of Gnathia trimaculata (Crustacea, Isopoda, Gnathiidae), ectoparasites of elasmobranchs from Okinawan coastal waters</mods:title>
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<mods:namePart id="C8DF0D2A6314746B0C8E22334981AC53">Ota, Yuzo</mods:namePart>
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<mods:namePart id="EAE799A3A91CD20E0A38564BD10D809D">Hirose, Euichi</mods:namePart>
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<mods:typeOfResource id="8FBC619BDD7E195741F94B55FB87B512">text</mods:typeOfResource>
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<mods:date id="F938F4A65F5D3B1523F9ADFCEEDC65E3">2009</mods:date>
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<treatment id="6216B3248C70FFAE58B9FA97FC0491BA" ID-DOI="http://doi.org/10.5281/zenodo.6214527" ID-GBIF-Taxon="119618003" ID-Zenodo-Dep="6214527" LSID="urn:lsid:plazi:treatment:6216B3248C70FFAE58B9FA97FC0491BA" httpUri="http://treatment.plazi.org/id/6216B3248C70FFAE58B9FA97FC0491BA" lastPageId="7" lastPageNumber="57" pageId="1" pageNumber="51">
<subSubSection id="A2A551B98C70FFA858B9FA97FE9F97A9" pageId="1" pageNumber="51" type="nomenclature">
<paragraph id="EA0002328C70FFA858B9FA97FE059776" blockId="1.[151,473,1346,1411]" box="[151,473,1346,1372]" pageId="1" pageNumber="51">
<heading id="B148B55E8C70FFA858B9FA97FE059776" bold="true" box="[151,473,1346,1372]" fontSize="11" level="1" pageId="1" pageNumber="51" reason="1">
<emphasis id="D8CBDE208C70FFA858B9FA97FE059776" bold="true" box="[151,473,1346,1372]" pageId="1" pageNumber="51">
<taxonomicName id="2DBF79B18C70FFA858B9FA97FEA99776" ID-CoL="3GLCL" box="[151,373,1346,1372]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="51" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D8CBDE208C70FFA858B9FA97FEA99776" bold="true" box="[151,373,1346,1372]" italics="true" pageId="1" pageNumber="51">Gnathia maculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C3F8635B8C70FFA85952FA97FE059776" box="[380,473,1346,1372]" pageId="1" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EA0002328C70FFA858B9FABCFE9F97A9" blockId="1.[151,473,1346,1411]" box="[151,323,1385,1411]" pageId="1" pageNumber="51">
(
<figureCitation id="72841EB78C70FFA858B1FABCFED497A9" box="[159,264,1385,1411]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,1936,1960]" captionStartId-1="3.[151,258,1941,1965]" captionStartId-2="4.[151,255,1916,1940]" captionTargetBox-0="[188,1407,187,1910]" captionTargetBox-1="[183,1388,194,1907]" captionTargetBox-2="[197,1399,194,1887]" captionTargetId-0="figure@2.[184,1408,187,1916]" captionTargetId-1="figure@3.[183,1388,194,1913]" captionTargetId-2="figure@4.[194,1399,194,1887]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." captionText-1="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." captionText-2="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri-0="https://zenodo.org/record/187942/files/figure.png" httpUri-1="https://zenodo.org/record/187943/files/figure.png" httpUri-2="https://zenodo.org/record/187944/files/figure.png" pageId="1" pageNumber="51">Figs 1–3</figureCitation>
, 
<figureCitation id="72841EB78C70FFA8593BFABCFEFF97A9" box="[277,291,1385,1411]" captionStart="FIGURE 5" captionStartId="9.[151,258,1218,1242]" captionTargetBox="[153,1422,302,1217]" captionTargetId="figure@9.[151,1436,263,1223]" captionTargetPageId="9" captionText="FIGURE 5. Live specimens of praniza larvae after feeding. A, paratype of Gnathia maculosa sp. nov. (NSMT-Cr 20426). B and C, Gnathia trimaculata (NSMT-Cr 20427). Scale bars = 1 mm." httpUri="https://zenodo.org/record/187946/files/figure.png" pageId="1" pageNumber="51">5</figureCitation>
A)
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C70FFA858B9FA63FCC394F5" pageId="1" pageNumber="51" type="materials_examined">
<paragraph id="EA0002328C70FFA858B9FA63FD679434" blockId="1.[151,1437,1462,2030]" pageId="1" pageNumber="51">
<emphasis id="D8CBDE208C70FFA858B9FA63FDDA97FA" bold="true" box="[151,518,1462,1488]" pageId="1" pageNumber="51">
Material examined. 
<typeStatus id="3504BC908C70FFA859BAFA63FDDA97FA" box="[404,518,1462,1488]" pageId="1" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
. Male, 
<quantity id="2D47AFD78C70FFA85A4FFA63FD6097FA" box="[609,700,1462,1488]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="51" unit="mm" value="5.7">5.7 mm</quantity>
(NSMT-Cr 20425), from gill chambers of blotched fantail ray 
<taxonomicName id="2DBF79B18C70FFA858B9FA08FDAF97DD" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[151,627,1501,1527]" class="Elasmobranchii" family="Dasyatidae" genus="Taeniura" kingdom="Animalia" order="Myliobatiformes" pageId="1" pageNumber="51" phylum="Chordata" rank="species" species="meyeni">
<emphasis id="D8CBDE208C70FFA858B9FA08FE8297DC" box="[151,350,1501,1526]" italics="true" pageId="1" pageNumber="51">Taeniura meyeni</emphasis>
Müller &amp; Henle, 1841
</taxonomicName>
, gill net, Nakagusuku Bay (26ºN, 127ºE), Okinawa Island, Ryukyu Archipelago, 
<collectingCountry id="92A842A28C70FFA85918F9D1FEA09434" box="[310,380,1540,1566]" name="Japan" pageId="1" pageNumber="51">Japan</collectingCountry>
, 
<date id="9E0124F28C70FFA859A6F9D1FDFB9434" box="[392,551,1540,1566]" pageId="1" pageNumber="51" value="2007-05-25">25 May 2007</date>
, coll. Y. O t a.
</paragraph>
<paragraph id="EA0002328C70FFA858EBF9FFFCC394F5" blockId="1.[151,1437,1462,2030]" pageId="1" pageNumber="51">
<emphasis id="D8CBDE208C70FFA858EBF9FFFE94946E" bold="true" box="[197,328,1578,1604]" pageId="1" pageNumber="51">
<typeStatus id="3504BC908C70FFA858EBF9FFFE94946E" box="[197,328,1578,1604]" pageId="1" pageNumber="51" type="paratype">Paratypes</typeStatus>
</emphasis>
. 
<specimenCount id="FCB9C9BB8C70FFA85977F9FFFE60946E" box="[345,444,1578,1604]" pageId="1" pageNumber="51" type="male">3 males</specimenCount>
, 
<specimenCount id="FCB9C9BB8C70FFA859E7F9FFFD9D946E" box="[457,577,1578,1604]" pageId="1" pageNumber="51" type="female">3 females</specimenCount>
and 5 praniza larvae from the same locality and host as the 
<typeStatus id="3504BC908C70FFA85D1FF9FFFA41946E" box="[1329,1437,1578,1604]" pageId="1" pageNumber="51" type="holotype">holotype</typeStatus>
(NSMT-Cr 20426). 
<specimenCount id="FCB9C9BB8C70FFA859ACF984FE3D9441" box="[386,481,1617,1643]" pageId="1" pageNumber="51" type="male">9 males</specimenCount>
, 
<specimenCount id="FCB9C9BB8C70FFA859C0F984FDB89441" box="[494,612,1617,1643]" pageId="1" pageNumber="51" type="female">2 females</specimenCount>
, and 2 praniza larvae from the same locality and host as the 
<typeStatus id="3504BC908C70FFA85D1BF984FA409441" box="[1333,1436,1617,1643]" pageId="1" pageNumber="51" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CAE9AF78C70FFA858B1F9ADFF3A94B8" LSID="urn:lsid:biocol.org:col:34978" box="[159,230,1656,1682]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="1" pageNumber="51">NMV</collectionCode>
<accessionNumber id="F5EC9FD18C70FFA858C0F9ADFE9694B8" box="[238,330,1656,1682]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J46718" pageId="1" pageNumber="51" type="EnaNcbi">J46718</accessionNumber>
). 2 praniza larvae from the same locality and host as the 
<typeStatus id="3504BC908C70FFA85BD6F9ADFBBC94B8" box="[1016,1120,1656,1682]" pageId="1" pageNumber="51" type="holotype">holotype</typeStatus>
(
<collectionCode id="8CAE9AF78C70FFA85C5FF9ADFB1194B8" box="[1137,1229,1656,1682]" pageId="1" pageNumber="51">KMNH</collectionCode>
IvR 500, 414). 
<specimenCount id="FCB9C9BB8C70FFA85DA3F9ADFF009492" pageId="1" pageNumber="51" type="male">9 males</specimenCount>
and 
<specimenCount id="FCB9C9BB8C70FFA85938F94BFE4A9492" box="[278,406,1694,1720]" pageId="1" pageNumber="51" type="female">12 females</specimenCount>
from a Javanese cownose ray 
<taxonomicName id="2DBF79B18C70FFA85B2DF94AFC299492" box="[771,1013,1695,1720]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="1" pageNumber="51" phylum="Chordata" rank="species" species="javanica">
<emphasis id="D8CBDE208C70FFA85B2DF94AFC299492" box="[771,1013,1695,1720]" italics="true" pageId="1" pageNumber="51">Rhinoptera javanica</emphasis>
</taxonomicName>
caught by a gill net in Nakagusuku Bay, 
<date id="9E0124F28C70FFA858FDF910FEAD94F5" box="[211,369,1733,1759]" pageId="1" pageNumber="51" value="2007-05-16">16 May 2007</date>
, coll. Y. Ota (
<collectionCode id="8CAE9AF78C70FFA85A3CF910FDB194F5" box="[530,621,1733,1759]" pageId="1" pageNumber="51">KMNH</collectionCode>
IvR 500, 413).
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C70FFAF58E8F93EFD2D9496" lastPageId="6" lastPageNumber="56" pageId="1" pageNumber="51" type="description">
<paragraph id="EA0002328C70FFA858E8F93EFE82952F" blockId="1.[151,1437,1462,2030]" box="[198,350,1771,1797]" pageId="1" pageNumber="51">
<emphasis id="D8CBDE208C70FFA858E8F93EFE82952F" bold="true" box="[198,350,1771,1797]" pageId="1" pageNumber="51">Description.</emphasis>
</paragraph>
<paragraph id="EA0002328C70FFA858B9F8C7FD1095C4" blockId="1.[151,1437,1462,2030]" pageId="1" pageNumber="51">
<emphasis id="D8CBDE208C70FFA858B9F8C7FF049506" bold="true" box="[151,216,1810,1836]" pageId="1" pageNumber="51">Male</emphasis>
(
<figureCitation id="72841EB78C70FFA858C7F8C7FEEC9506" box="[233,304,1810,1836]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="1" pageNumber="51">Fig. 1</figureCitation>
). 
<emphasis id="D8CBDE208C70FFA85962F8C6FE569506" box="[332,394,1811,1836]" italics="true" pageId="1" pageNumber="51">Body</emphasis>
<quantity id="2D47AFD78C70FFA859BDF8C7FDF89506" box="[403,548,1810,1836]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.8" metricValueMin="3.9" pageId="1" pageNumber="51" unit="mm" value="4.85" valueMax="5.8" valueMin="3.9">3.9–5.8 mm</quantity>
(mean ± SD; 5.0 ± 
<quantity id="2D47AFD78C70FFA85B36F8C7FCA59506" box="[792,889,1810,1836]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="51" unit="mm" value="0.6">0.6 mm</quantity>
, n = 21) long, color of live specimens white. 
<emphasis id="D8CBDE208C70FFA858B9F8ECFEE29578" box="[151,318,1849,1874]" italics="true" pageId="1" pageNumber="51">Cephalosome</emphasis>
(
<figureCitation id="72841EB78C70FFA8597EF8ECFE499579" box="[336,405,1849,1875]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="1" pageNumber="51">Fig. 1</figureCitation>
A–D) almost square, with posterior margin convex, about one-fifth of total length. 
<emphasis id="D8CBDE208C70FFA858B9F8B5FE959553" box="[151,329,1888,1913]" italics="true" pageId="1" pageNumber="51">Frontal border</emphasis>
(
<figureCitation id="72841EB78C70FFA85974F8B5FE419550" box="[346,413,1888,1914]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="1" pageNumber="51">Fig. 1</figureCitation>
B) with several setae, medially notched by 2 frontolateral processes; notch anteriorly opened. Apex of frontolateral process serrate. Eyes well developed, lateral and sessile, about one-fifth length of cephalosome. Supraocular lobe slightly prominent. Paraocular ornamentation present as several tubercles around the eyes. Dorsal sulcus deep and narrow.
</paragraph>
<caption id="BEC052BA8C73FFAB58B9F845FC1E95C6" httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="2" pageNumber="52" targetBox="[188,1407,187,1910]" targetPageId="2">
<paragraph id="EA0002328C73FFAB58B9F845FC1E95C6" blockId="2.[151,1436,1936,2028]" pageId="2" pageNumber="52">
<emphasis id="D8CBDE208C73FFAB58B9F845FEC59582" bold="true" box="[151,281,1936,1960]" pageId="2" pageNumber="52">FIGURE 1.</emphasis>
<taxonomicName id="2DBF79B18C73FFAB590EF845FE38958D" box="[288,484,1936,1959]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D8CBDE208C73FFAB590EF845FE38958D" box="[288,484,1936,1959]" italics="true" pageId="2" pageNumber="52">Gnathia maculosa</emphasis>
</taxonomicName>
<emphasis id="D8CBDE208C73FFAB59C5F845FD9C9582" bold="true" box="[491,576,1936,1960]" pageId="2" pageNumber="52">
<taxonomicNameLabel id="C3F8635B8C73FFAB59C5F845FD9C9582" box="[491,576,1936,1960]" pageId="2" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2.
</paragraph>
</caption>
<caption id="BEC052BA8C72FFAA58B9F840FE4495D8" httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="3" pageNumber="53" targetBox="[183,1388,194,1907]" targetPageId="3">
<paragraph id="EA0002328C72FFAA58B9F840FE4495D8" blockId="3.[151,1437,1941,2034]" pageId="3" pageNumber="53">
<emphasis id="D8CBDE208C72FFAA58B9F840FEC39587" bold="true" box="[151,287,1941,1965]" pageId="3" pageNumber="53">FIGURE 2.</emphasis>
<taxonomicName id="2DBF79B18C72FFAA5908F843FE2F9587" box="[294,499,1942,1965]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="53" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D8CBDE208C72FFAA5908F843FE2F9587" box="[294,499,1942,1965]" italics="true" pageId="3" pageNumber="53">Gnathia maculosa</emphasis>
</taxonomicName>
<emphasis id="D8CBDE208C72FFAA59D5F840FD899587" bold="true" box="[507,597,1941,1965]" pageId="3" pageNumber="53">
<taxonomicNameLabel id="C3F8635B8C72FFAA59D5F840FD899587" box="[507,597,1941,1965]" pageId="3" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2.
</paragraph>
</caption>
<caption id="BEC052BA8C75FFAD58B9F8A9FB8E95F3" httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="4" pageNumber="54" targetBox="[197,1399,194,1887]" targetPageId="4">
<paragraph id="EA0002328C75FFAD58B9F8A9FB8E95F3" blockId="4.[151,1436,1916,2009]" pageId="4" pageNumber="54">
<emphasis id="D8CBDE208C75FFAD58B9F8A9FEC595BE" bold="true" box="[151,281,1916,1940]" pageId="4" pageNumber="54">FIGURE 3.</emphasis>
<taxonomicName id="2DBF79B18C75FFAD590EF8A8FE3895BE" box="[288,484,1917,1940]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="54" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D8CBDE208C75FFAD590EF8A8FE3895BE" box="[288,484,1917,1940]" italics="true" pageId="4" pageNumber="54">Gnathia maculosa</emphasis>
</taxonomicName>
<emphasis id="D8CBDE208C75FFAD59C5F8A9FD9D95BE" bold="true" box="[491,577,1916,1940]" pageId="4" pageNumber="54">
<taxonomicNameLabel id="C3F8635B8C75FFAD59C5F8A9FD9D95BE" box="[491,577,1916,1940]" pageId="4" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2.
</paragraph>
</caption>
<paragraph id="EA0002328C74FFAC58EBFF4DFDA59359" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFF4DFE98929B" box="[197,324,152,177]" italics="true" pageId="5" pageNumber="55">Pereonites</emphasis>
(
<figureCitation id="72841EB78C74FFAC597BFF4DFE459298" box="[341,409,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
A) sparsely covered with setae. Pereonites 1–7 about half of total length. Pereonite 1 very short, not fused, concave anteriorly. Pereonites 2, 3, and 4 of same width. Pereonite 2 twice as long as pereonite 1; lateral margins extend anteriorly. Pereonite 3 slightly longer than pereonite 2. Pereonite 4 with constriction anteriorly, twice as long as pereonite 3. Pereonites 5 and 6 together subequal in length to pereonites 2–4 combined. Pereonites 5 and 6 with areae laterales and lobi laterales, respectively. Pereonite 7 short and narrow, overlapping pleonite 1.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFE55FC4593EA" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFE55FEEA93B3" box="[197,310,384,409]" italics="true" pageId="5" pageNumber="55">Pleonites</emphasis>
(
<figureCitation id="72841EB78C74FFAC5966FE55FE5193B0" box="[328,397,384,410]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
A) with several setae on lateral margins subequal in length. Pleonites and pleotelson about one-fourth of total length. Pleonal epimera not pronounced.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFE18FDDC9024" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFE18FE9893CC" box="[197,324,461,486]" italics="true" pageId="5" pageNumber="55">Pleotelson</emphasis>
(
<figureCitation id="72841EB78C74FFAC597BFE18FE4693CD" box="[341,410,461,487]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
K) triangular, with middle of outer margins convex; 3 pairs of small setae on center, lateral margin, and distal apex.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFDCFFD2D90E5" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFDCFFE9C9019" box="[197,320,538,563]" italics="true" pageId="5" pageNumber="55">Antenna 1</emphasis>
(
<figureCitation id="72841EB78C74FFAC597CFDCFFE4A901E" box="[338,406,538,564]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
E) composed of 3 peduncular and 5 flagellar articles; 2 and 3 feather-like bristles on external margins of peduncles 1 and 2, respectively. Internal margins of peduncles 1 and 2 with 1 seta. Several setae on external and distal margins of peduncle 3. Length ratio of peduncular articles about 1:1:2. Featherlike bristles on flagellar articles 1 and 5. Aesthetascs on articles 3 and 5. Article 5 terminates in 3 long setae. Length ratio of 5 flagellar articles about 1:12:2:1:2.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFD09FE3A9140" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFD09FE9F90DF" box="[197,323,732,757]" italics="true" pageId="5" pageNumber="55">Antenna 2</emphasis>
(
<figureCitation id="72841EB78C74FFAC597AFD09FE4690DC" box="[340,410,732,758]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
F) slightly longer than antenna 1; 2 and 3 feather-like bristles and several setae on peduncle articles 3 and 4, respectively. Length ratio of peduncular articles about 1:1:2:3. Few setae on distal margins of flagellar articles 1–6. Article 7 terminates in 4 long setae. Length ratio of flagellar articles approximately 2:2:2:2:2:2:1.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFCA2FD7D91F4" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFCA2FE9891BA" box="[197,324,887,912]" italics="true" pageId="5" pageNumber="55">Mandibles</emphasis>
(
<figureCitation id="72841EB78C74FFAC597BFCA3FE4491BA" box="[341,408,886,912]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
B–D) three-fourths length of cephalosome. Carina unarmed. Apex curved internally. Dentate blade occupies about two-fifths length of mandible. Basal neck and erisma prominent. One mandibular seta on middle of dorsal surface.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFC3FFBFC9678" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFC3FFEF59629" box="[197,297,1002,1027]" italics="true" pageId="5" pageNumber="55">Pylopod</emphasis>
(
<figureCitation id="72841EB78C74FFAC5914FC3FFEA1962E" box="[314,381,1002,1028]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
D, G) with 3 articles, ventral surfaces of article 1 with 19 setae, article 2 with 18 setae. Article 1 with 3 areolae, large and elliptical, fringed by fine setae on external margin, and 47 plumose setae on internal margin. Article 2 circular and fringed by fine setae. Article 3 minute.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFB8AFB0F96B5" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFB8AFE989652" box="[197,324,1119,1144]" italics="true" pageId="5" pageNumber="55">Maxilliped</emphasis>
(
<figureCitation id="72841EB78C74FFAC597DFB8BFE499652" box="[339,405,1118,1144]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
H) with fine setae on lateral margin. Endite reaches palp article 2. Palp articles 1–4 with 6, 8, 5 and 8 plumose setae on external margins, respectively. Article 4 terminates in 4 setae.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFB79FDE297AD" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFB79FE9D96EF" box="[197,321,1196,1221]" italics="true" pageId="5" pageNumber="55">Pereopods</emphasis>
(
<figureCitation id="72841EB78C74FFAC597FFB79FE4F96EC" box="[337,403,1196,1222]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
J) subequal in shape and bearing many setae; longer on outer margin than inner margin. Basis oblong; with 2 feather-like bristles on outer margin. Ischium about three-fourths length of basis, becoming larger distally. Merus about half length of ischium. Carpus elliptical and about four-fifths length of merus. Propodus rectangular, about 1.2 times longer than carpus; with 2 spines on inner-mid and inner-distal margins. Dactylus with few setae. Length of dactylus and unguis combined about half that of propodus. All pereopods similar in shape and size.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFA41FC0C9784" blockId="5.[151,1437,152,2034]" box="[197,976,1428,1454]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFA41FED79787" box="[197,267,1428,1453]" italics="true" pageId="5" pageNumber="55">Penes</emphasis>
(
<figureCitation id="72841EB78C74FFAC5935FA41FE829784" box="[283,350,1428,1454]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">
Fig. 
<date id="9E0124F28C74FFAC597CFA41FE829784" box="[338,350,1428,1454]" pageId="5" pageNumber="55">1</date>
</figureCitation>
I) composed of 2 contiguous papillae; not prominent.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBFA6EFDB89445" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBFA6EFE7297FE" box="[197,430,1467,1492]" italics="true" pageId="5" pageNumber="55">Pleopodal peduncle</emphasis>
(
<figureCitation id="72841EB78C74FFAC5990FA6EFE2397FF" box="[446,511,1467,1493]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
L) fringed with fine setae, 1 seta on outer distal corner; coupling hook on inner margin. Both pleopodal rami oval-shaped and equal in length. All pleopods subequal in shape. Pleopod 1 exopod with 7 plumose setae, 6 plumose setae on endopod; pleopod 2 exopod with 9 plumose setae, endopod with 7 plumose setae. Appendix masculina extends beyond half of endopod of pleopod 2. Exopod of pleopods 3–5 with 9 setae, endopod with 8 setae.
</paragraph>
<paragraph id="EA0002328C74FFAC58EBF9A9FB4F9497" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBF9A9FEA994BF" box="[197,373,1660,1685]" italics="true" pageId="5" pageNumber="55">Uropodal rami</emphasis>
(
<figureCitation id="72841EB78C74FFAC59A8F9A9FE1B94BC" box="[390,455,1660,1686]" captionStart="FIGURE 1" captionStartId="2.[151,255,1936,1960]" captionTargetBox="[188,1407,187,1910]" captionTargetId="figure@2.[184,1408,187,1916]" captionTargetPageId="2" captionText="FIGURE 1. Gnathia maculosa sp. nov. Male holotype, 5.7 mm (NSMT-Cr 20425). A, body in dorsal view. B, mandible and frontal border. C, cephalosome in dorsolateral view. D, cephalosome in ventral view. E, antenna 1. F, antenna 2. G, pylopod. H, maxilliped. I, penes. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187942/files/figure.png" pageId="5" pageNumber="55">Fig. 1</figureCitation>
K) subequal in length, extending beyond apex of pleotelson. Exopod with 15 setae on margin. Endopod withng 12 setae on margin and 2 coupling setae on dorsal surface.
</paragraph>
<paragraph id="EA0002328C74FFAC58E8F91DFA4094C9" blockId="5.[151,1437,152,2034]" box="[198,1436,1736,1763]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58E8F91DFEFE94C8" bold="true" box="[198,290,1736,1762]" pageId="5" pageNumber="55">Female</emphasis>
(
<figureCitation id="72841EB78C74FFAC591FF91CFEA594C9" box="[305,377,1737,1763]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
). 
<emphasis id="D8CBDE208C74FFAC59A0F91CFE1694C8" box="[398,458,1737,1762]" italics="true" pageId="5" pageNumber="55">Body</emphasis>
<quantity id="2D47AFD78C74FFAC59FCF91CFD8294C9" box="[466,606,1737,1763]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.5" metricValueMin="4.1" pageId="5" pageNumber="55" unit="mm" value="4.8" valueMax="5.5" valueMin="4.1">4.1–5.5 mm</quantity>
(4.5 ± 
<quantity id="2D47AFD78C74FFAC5A9CF91CFCD394C9" box="[690,783,1737,1763]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="55" unit="mm" value="0.3">0.3 mm</quantity>
, n = 17), color of live specimens light orange or yellow.
</paragraph>
<paragraph id="EA0002328C74FFAC58E8F925FF37951A" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58E8F925FEBA9523" box="[198,358,1776,1801]" italics="true" pageId="5" pageNumber="55">Cephalosome</emphasis>
(
<figureCitation id="72841EB78C74FFAC5953F925FE159520" box="[381,457,1776,1802]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
A) elliptical. Frontal margin slightly convex. Eyes well developed, lateral and sessile.
</paragraph>
<paragraph id="EA0002328C74FFAC58E8F8E8FDF895E1" blockId="5.[151,1437,152,2034]" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58E8F8E8FE9E957C" box="[198,322,1853,1878]" italics="true" pageId="5" pageNumber="55">Pereonites</emphasis>
(
<figureCitation id="72841EB78C74FFAC5977F8E8FE7A957D" box="[345,422,1853,1879]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
A, B) with several setae on lateral margins and dorsal surface, swollen between pereonites 3 and 6. Pereonite 1 very short, not fused. Pereonites 2 and 3 with lateral margins that extend anteriorly. Pereonites 3–6 oval, with 3 sutures. Lateral shields of pereopods 4–6 visible dorsally. Pereopod 7 visible dorsally, short and narrow.
</paragraph>
<paragraph id="EA0002328C74FFAF58EBF80DFCF69298" blockId="5.[151,1437,152,2034]" lastBlockId="6.[151,1437,152,2034]" lastPageId="6" lastPageNumber="56" pageId="5" pageNumber="55">
<emphasis id="D8CBDE208C74FFAC58EBF80DFEEE95DB" box="[197,306,2008,2033]" italics="true" pageId="5" pageNumber="55">Pleonites</emphasis>
(
<figureCitation id="72841EB78C74FFAC596CF80DFE5495D8" box="[322,392,2008,2034]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
A, B) with several setae subequal in length. 
<emphasis id="D8CBDE208C74FFAC5BA0F80DFBD695DB" box="[910,1034,2008,2033]" italics="true" pageId="5" pageNumber="55">Pleotelson</emphasis>
(
<figureCitation id="72841EB78C74FFAC5C34F80DFBBC95D8" box="[1050,1120,2008,2034]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="5" pageNumber="55">Fig. 2</figureCitation>
G) triangular with an acute apex. 5 pairs of simple setae on dorsal surface and apex.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFF6AFA8A930C" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFF6AFEE392F2" box="[197,319,191,216]" italics="true" pageId="6" pageNumber="56">Antenna 1</emphasis>
(
<figureCitation id="72841EB78C77FFAF597FFF6BFE4492F2" box="[337,408,190,216]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
C) composed of 3 peduncular and 4 flagellar articles. Pedunclar articles covered with scales and fine setae. Aesthetasc on articles 2–4. Article 4 terminates in 3 setae. 
<emphasis id="D8CBDE208C77FFAF5CB2FF30FAC292D4" box="[1180,1310,229,254]" italics="true" pageId="6" pageNumber="56">Antenna 2</emphasis>
(
<figureCitation id="72841EB78C77FFAF5D1EFF30FAA092D5" box="[1328,1404,229,255]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
D) composed of 4 peduncular and 7 flagellar articles. Pedunclar articles covered with scales and fine setae.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFEE7FBD29366" blockId="6.[151,1437,152,2034]" box="[197,1038,306,332]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFEE7FEFB9361" box="[197,295,306,331]" italics="true" pageId="6" pageNumber="56">Pylopod</emphasis>
(
<figureCitation id="72841EB78C77FFAF5918FEE7FEA09366" box="[310,380,306,332]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
E) with 3 articles; article 2 rectangular; article 3 minute.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFE8CFCB993B0" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFE8CFE999358" box="[197,325,345,370]" italics="true" pageId="6" pageNumber="56">Maxilliped</emphasis>
(
<figureCitation id="72841EB78C77FFAF597BFE8CFE409359" box="[341,412,345,371]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
F) endite sparsely covered with fine setae, reaching half of article 1 of palp. Basis and articles 1–4 with 2, 7, 9, 8, and 9 plumose setae, respectively.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFE72FE489024" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFE72FE9493EA" box="[197,328,423,448]" italics="true" pageId="6" pageNumber="56">Pereopods</emphasis>
(
<figureCitation id="72841EB78C77FFAF5975FE73FE7B93EA" box="[347,423,422,448]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
G) bearing fewer setae than those of male, covered with scales and fine setae on ischium, merus, and carpus. Each article more slender in shape than those of male. Pereopods 5 and 6 larger than other pereopods.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFDCFFEDC9082" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFDCFFE739019" box="[197,431,538,563]" italics="true" pageId="6" pageNumber="56">Pleopodal peduncle</emphasis>
(
<figureCitation id="72841EB78C77FFAF59EEFDCFFDDB901E" box="[448,519,538,564]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">
Fig. 
<date id="9E0124F28C77FFAF59D6FDCFFDDB901E" box="[504,519,538,564]" pageId="6" pageNumber="56">2</date>
</figureCitation>
I) has a few setae on lateral margin. Exopods fan-shaped and with 8 (pleopod 3) or 9 (pleopods 1, 2, 4, and 5) plumose setae on distal margins. Endopods subequal in shape and size to exopods; 7 (pleopod 3) or 8 (pleopods 1, 2, 4, and 5) plumose setae on distal margins. All pleopods subequal in shape.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFD60FE2A9136" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFD60FDBF90E4" box="[197,611,693,718]" italics="true" pageId="6" pageNumber="56">Uropodal peduncle and both rami</emphasis>
(
<figureCitation id="72841EB78C77FFAF5A5BFD60FD6190E5" box="[629,701,693,719]" captionStart="FIGURE 2" captionStartId="3.[151,258,1941,1965]" captionTargetBox="[183,1388,194,1907]" captionTargetId="figure@3.[183,1388,194,1913]" captionTargetPageId="3" captionText="FIGURE 2. Gnathia maculosa sp. nov. Female paratype, 4.6 mm (NSMT-Cr 20246). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, pylopod. F, maxilliped. G, pereopod 2. H, pleotelson. I, pleopod 2." httpUri="https://zenodo.org/record/187943/files/figure.png" pageId="6" pageNumber="56">Fig. 2</figureCitation>
H) subequal in shape. Uropodal rami extend beyond apex of pleotelson. Exopod bears 12 long setae on margin. Endopod bears 9 setae on margin and 2 pairs and 7 featherlike bristles on dorsal surface.
</paragraph>
<paragraph id="EA0002328C77FFAF58E8FCFDFB969140" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58E8FCFDFEAC9168" bold="true" box="[198,368,808,834]" pageId="6" pageNumber="56">Praniza larva</emphasis>
(
<figureCitation id="72841EB78C77FFAF5951FCFCFE0D9169" box="[383,465,809,835]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Figs. 3</figureCitation>
, 
<figureCitation id="72841EB78C77FFAF59F0FCFCFE319169" box="[478,493,809,835]" captionStart="FIGURE 5" captionStartId="9.[151,258,1218,1242]" captionTargetBox="[153,1422,302,1217]" captionTargetId="figure@9.[151,1436,263,1223]" captionTargetPageId="9" captionText="FIGURE 5. Live specimens of praniza larvae after feeding. A, paratype of Gnathia maculosa sp. nov. (NSMT-Cr 20426). B and C, Gnathia trimaculata (NSMT-Cr 20427). Scale bars = 1 mm." httpUri="https://zenodo.org/record/187946/files/figure.png" pageId="6" pageNumber="56">5</figureCitation>
A). 
<emphasis id="D8CBDE208C77FFAF5A34FCFCFD8A9168" box="[538,598,809,834]" italics="true" pageId="6" pageNumber="56">Body</emphasis>
<quantity id="2D47AFD78C77FFAF5A73FCFCFD369169" box="[605,746,809,835]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.8" metricValueMin="4.2" pageId="6" pageNumber="56" unit="mm" value="5.0" valueMax="5.8" valueMin="4.2">4.2–5.8 mm</quantity>
(5.1 ± 
<quantity id="2D47AFD78C77FFAF5B13FCFCFC469169" box="[829,922,809,835]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="56" unit="mm" value="0.5">0.5 mm</quantity>
, n = 7). Thorax of live specimen dark red or black with white speckles or dapples (
<figureCitation id="72841EB78C77FFAF5A7BFC85FD479140" box="[597,667,848,874]" captionStart="FIGURE 5" captionStartId="9.[151,258,1218,1242]" captionTargetBox="[153,1422,302,1217]" captionTargetId="figure@9.[151,1436,263,1223]" captionTargetPageId="9" captionText="FIGURE 5. Live specimens of praniza larvae after feeding. A, paratype of Gnathia maculosa sp. nov. (NSMT-Cr 20426). B and C, Gnathia trimaculata (NSMT-Cr 20427). Scale bars = 1 mm." httpUri="https://zenodo.org/record/187946/files/figure.png" pageId="6" pageNumber="56">Fig. 5</figureCitation>
A). Other parts light brown or white.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFCA2FBB491BA" blockId="6.[151,1437,152,2034]" box="[197,1128,886,912]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFCA2FEB991BA" box="[197,357,887,912]" italics="true" pageId="6" pageNumber="56">Cephalosome</emphasis>
(
<figureCitation id="72841EB78C77FFAF595AFCA3FE6491BA" box="[372,440,886,912]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
A) triangular. Eyes well developed, and lateral and sessile.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFC48FDA5962E" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFC48FEE7919C" box="[197,315,925,950]" italics="true" pageId="6" pageNumber="56">Pereonite</emphasis>
1 (
<figureCitation id="72841EB78C77FFAF594BFC48FE72919D" box="[357,430,925,951]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
A, B) very short, reaching lateral margin. Anterior margins of pereonites 2 and 3 concave. Lateral shields of pereopods 4–6 visible in dorsal view, elliptical. Pereonite 7 visible in dorsal view; short and narrow, overlapping pleonite 1.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFBC4FADA9678" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFBC4FEEE9600" box="[197,306,1041,1066]" italics="true" pageId="6" pageNumber="56">Pleonites</emphasis>
1–4 subequal in length. Epimera not prominent in dorsal view. 
<emphasis id="D8CBDE208C77FFAF5C06FBC4FB789600" box="[1064,1188,1041,1066]" italics="true" pageId="6" pageNumber="56">Pleotelson</emphasis>
(
<figureCitation id="72841EB78C77FFAF5C98FBC4FB279601" box="[1206,1275,1041,1067]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
K) triangular, with dentate margin at apex, bearing 2 pairs of simple setae on posterior lateral margin and apex.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFB8AFAAC96EC" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFB8AFEE39652" box="[197,319,1119,1144]" italics="true" pageId="6" pageNumber="56">Antenna 1</emphasis>
(
<figureCitation id="72841EB78C77FFAF597EFB8BFE4B9652" box="[336,407,1118,1144]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
C) composed of 3 peduncular and 4 flagellar articles. Internal margins of peduncles 2 and 3 fringed by fine setae. An aesthetasc each on articles 2–4. 
<emphasis id="D8CBDE208C77FFAF5B59FB50FC3296B4" box="[887,1006,1157,1182]" italics="true" pageId="6" pageNumber="56">Antenna 2</emphasis>
(
<figureCitation id="72841EB78C77FFAF5BD3FB50FB9C96B5" box="[1021,1088,1157,1183]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
D) composed of 4 peduncular and 7 flagellar articles; longer than antenna 1. Internal margins of peduncles 3 and 4 fringed by fine setae.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFB06FC699784" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFB06FE9596C6" box="[197,329,1235,1260]" italics="true" pageId="6" pageNumber="56">Gnathopod</emphasis>
(
<figureCitation id="72841EB78C77FFAF5976FB07FE4796C6" box="[344,411,1234,1260]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
E) with pereopodal shape. Ischium 2 times as long as basis. Merus subequal in length to ischium. Carpus reduced. Propodus with distal bulbous protrusion. 
<emphasis id="D8CBDE208C77FFAF5B97FB2CFBF69738" box="[953,1066,1273,1298]" italics="true" pageId="6" pageNumber="56">Mandible</emphasis>
(
<figureCitation id="72841EB78C77FFAF5C15FB2CFB5D9739" box="[1083,1153,1273,1299]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
F) with 9 teeth on inner margin. 
<emphasis id="D8CBDE208C77FFAF58D1FAF5FEAA9713" box="[255,374,1312,1337]" italics="true" pageId="6" pageNumber="56">Maxillule</emphasis>
(
<figureCitation id="72841EB78C77FFAF59A6FAF5FE0D9710" box="[392,465,1312,1338]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
G) with slender article; 6 teeth on apex. 
<emphasis id="D8CBDE208C77FFAF5BE1FAF5FB8F9713" box="[975,1107,1312,1337]" italics="true" pageId="6" pageNumber="56">Paragnath</emphasis>
(
<figureCitation id="72841EB78C77FFAF5C4BFAF5FB739710" box="[1125,1199,1312,1338]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
H) slightly curved. 
<emphasis id="D8CBDE208C77FFAF58B9FA92FECA974A" box="[151,278,1351,1376]" italics="true" pageId="6" pageNumber="56">Maxilliped</emphasis>
(
<figureCitation id="72841EB78C77FFAF5908FA93FEB6974A" box="[294,362,1350,1376]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">
Fig. 
<date id="9E0124F28C77FFAF5972FA93FEB6974A" box="[348,362,1350,1376]" pageId="6" pageNumber="56">3</date>
</figureCitation>
I) composed of a basis and two-articled palp. Basis with coupling hook on inner margin and an endite with one seta. Apex of palp 1 has 4 fine setae, 2 stout setae, 2 teeth, and endite; style-like with 3 teeth facing ventrally. 5 setae and 1 spine on distal margin of palp 2.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBFA6EFDD797D1" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBFA6EFE9997FE" box="[197,325,1467,1492]" italics="true" pageId="6" pageNumber="56">Pereopods</emphasis>
(
<figureCitation id="72841EB78C77FFAF5979FA6FFE4397FE" box="[343,415,1466,1492]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
J) more slender shape and more scarce setae than those of male. Pereopods 5 and 6 larger than the other pereopods.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBF9DDFB579445" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBF9DDFE66940B" box="[197,442,1544,1569]" italics="true" pageId="6" pageNumber="56">Pleopodal peduncles</emphasis>
(
<figureCitation id="72841EB78C77FFAF59E4F9DDFDC69408" box="[458,538,1544,1570]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Figs. 3</figureCitation>
L) with coupling hook on inner margin. Exopods fan-shaped and 8 (pleopod 1) or 9 (pleopod 2–5) plumose setae on distal margins. Endopods subequal in shape and size to exopods, and eight (pleopods 1–5) plumose setae on distal margins. All pleopods subequal in shape.
</paragraph>
<paragraph id="EA0002328C77FFAF58EBF9A9FD2D9496" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58EBF9A9FD8394BF" box="[197,607,1660,1685]" italics="true" pageId="6" pageNumber="56">Uropodal peduncle and both rami</emphasis>
(
<figureCitation id="72841EB78C77FFAF5A41F9A9FD6A94BC" box="[623,694,1660,1686]" captionStart="FIGURE 3" captionStartId="4.[151,255,1916,1940]" captionTargetBox="[197,1399,194,1887]" captionTargetId="figure@4.[194,1399,194,1887]" captionTargetPageId="4" captionText="FIGURE 3. Gnathia maculosa sp. nov. Praniza larva paratype, 5.5 mm (NSMT-Cr 20426). A, body, left eye and frontal border, dorsal view. B, body in lateral view. C, antenna 1. D, antenna 2. E, gnathopod. F, mandible in lateral view. G, maxillule. H, pair of paragnaths. I, maxilliped. J, pereopod 2. K, pleotelson. L, pleopod 2." httpUri="https://zenodo.org/record/187944/files/figure.png" pageId="6" pageNumber="56">Fig. 3</figureCitation>
K) subequal in shape to those of male. Uropodal rami fringed with fine setae and do not reach apex of pleotelson.
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C77FFAF58E8F91DFD229520" pageId="6" pageNumber="56" type="etymology">
<paragraph id="EA0002328C77FFAF58E8F91DFD229520" blockId="6.[151,1437,152,2034]" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58E8F91DFE8F94C8" bold="true" box="[198,339,1736,1762]" pageId="6" pageNumber="56">Etymology.</emphasis>
The specific name 
<taxonomicName id="2DBF79B18C77FFAF5A6AF91CFD6894C8" box="[580,692,1737,1762]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="D8CBDE208C77FFAF5A6AF91CFD6894C8" box="[580,692,1737,1762]" italics="true" pageId="6" pageNumber="56">maculosa</emphasis>
</taxonomicName>
is derived from Latin, meaning “the color disposed in broad, irregular blotches” and referring to the larval thorax.
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C77FFAE58E8F8C3FBFA9366" lastPageId="7" lastPageNumber="57" pageId="6" pageNumber="56" type="discussion">
<paragraph id="EA0002328C77FFAE58E8F8C3FBFA9366" blockId="6.[151,1437,152,2034]" lastBlockId="7.[151,1436,152,912]" lastPageId="7" lastPageNumber="57" pageId="6" pageNumber="56">
<emphasis id="D8CBDE208C77FFAF58E8F8C3FEE3951A" bold="true" box="[198,319,1814,1840]" pageId="6" pageNumber="56">Remarks.</emphasis>
<taxonomicName id="2DBF79B18C77FFAF5964F8C2FDF8951A" box="[330,548,1815,1840]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="D8CBDE208C77FFAF5964F8C2FDF8951A" box="[330,548,1815,1840]" italics="true" pageId="6" pageNumber="56">Gnathia maculosa</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="2DBF79B18C77FFAF5B22F8C2FC6A951A" box="[780,950,1815,1840]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C77FFAF5B22F8C2FC6A951A" box="[780,950,1815,1840]" italics="true" pageId="6" pageNumber="56">G. trimaculata</emphasis>
</taxonomicName>
. However, 
<emphasis id="D8CBDE208C77FFAF5C6AF8C2FB08951A" box="[1092,1236,1815,1840]" italics="true" pageId="6" pageNumber="56">G. m a c u l o s a</emphasis>
is distinguished from 
<taxonomicName id="2DBF79B18C77FFAF58F9F8E8FE5C957C" box="[215,384,1853,1878]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C77FFAF58F9F8E8FE5C957C" box="[215,384,1853,1878]" italics="true" pageId="6" pageNumber="56">G. trimaculata</emphasis>
</taxonomicName>
by deeper and narrower dorsal sulcus on cephalosome, a narrower body, and the shape of the pylopod (slender pylopod in 
<taxonomicName id="2DBF79B18C77FFAF5A3DF8B1FD609557" box="[531,700,1892,1917]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C77FFAF5A3DF8B1FD609557" box="[531,700,1892,1917]" italics="true" pageId="6" pageNumber="56">G. trimaculata</emphasis>
</taxonomicName>
) (
<bibRefCitation id="8E2E7FC38C77FFAF5AFAF8B1FC189554" author="Coetzee" box="[724,964,1892,1918]" pageId="6" pageNumber="60" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2009) Gnathia trimaculata n. sp. (Crustacea: Isopoda: Gnathiidae), an ectoparasite found parasitising requiem sharks from Lizard Island, Great Barrier Reef, Australia. Systematic Parasitology, 72, 97 - 112." type="journal article" year="2009">
Coetzee 
<emphasis id="D8CBDE208C77FFAF5B12F8B1FCB39557" box="[828,879,1892,1917]" italics="true" pageId="6" pageNumber="56">et al</emphasis>
., 2009
</bibRefCitation>
). The overall proportions and mandibles of 
<emphasis id="D8CBDE208C77FFAF5897F85EFE96958E" box="[185,330,1931,1956]" italics="true" pageId="6" pageNumber="56">G. m a c u l o s a</emphasis>
are also similar to 
<taxonomicName id="2DBF79B18C77FFAF5A15F85EFBFF958E" authority="Cohen &amp; Poore, 1994" authorityName="Cohen &amp; Poore" authorityYear="1994" box="[571,1059,1930,1956]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="odontomachus">
<emphasis id="D8CBDE208C77FFAF5A15F85EFCD1958E" box="[571,781,1931,1956]" italics="true" pageId="6" pageNumber="56">G. odontomachus</emphasis>
<bibRefCitation id="8E2E7FC38C77FFAF5B3BF85FFBFF958E" author="Cohen" box="[789,1059,1930,1956]" pageId="6" pageNumber="60" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore, 1994</bibRefCitation>
</taxonomicName>
. However, 
<taxonomicName id="2DBF79B18C77FFAF5C83F85EFAA3958E" box="[1197,1407,1931,1956]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="odontomachus">
<emphasis id="D8CBDE208C77FFAF5C83F85EFAA3958E" box="[1197,1407,1931,1956]" italics="true" pageId="6" pageNumber="56">G. odontomachus</emphasis>
</taxonomicName>
is smaller in its total length (
<quantity id="2D47AFD78C77FFAF59E4F864FDFA95E1" box="[458,550,1969,1995]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="56" unit="mm" value="2.9">2.9 mm</quantity>
) than 
<emphasis id="D8CBDE208C77FFAF5A5EF864FD2095E0" box="[624,764,1969,1994]" italics="true" pageId="6" pageNumber="56">G. m a c u l o s a</emphasis>
and has a mediofrontal process (
<bibRefCitation id="8E2E7FC38C77FFAF5CAFF864FA5B95E1" author="Cohen" box="[1153,1415,1969,1995]" pageId="6" pageNumber="60" refString="Cohen, B. F. &amp; Poore, G. C. B. (1994) Phylogeny and biogeography of the Gnathiidae (Crustacea: Isopoda) with descriptions of new genera and species, most from south-eastern Australia. Memoirs of Museum Victoria, 54 (2), 271 - 397." type="journal article" year="1994">Cohen &amp; Poore, 1994</bibRefCitation>
). 
<taxonomicName id="2DBF79B18C77FFAF58B9F80DFEB595DB" box="[151,361,2008,2033]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="capillata">
<emphasis id="D8CBDE208C77FFAF58B9F80DFEB595DB" box="[151,361,2008,2033]" italics="true" pageId="6" pageNumber="56">Gnathia capillata</emphasis>
</taxonomicName>
, 
<emphasis id="D8CBDE208C77FFAF5957F80DFCDB95DB" box="[377,775,2008,2033]" italics="true" pageId="6" pageNumber="56">
<taxonomicName id="2DBF79B18C77FFAF5957F80DFDFE95DB" box="[377,546,2008,2033]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="grandilaris">G. grandilaris</taxonomicName>
, and 
<taxonomicName id="2DBF79B18C77FFAF5A4DF80DFCDB95DB" box="[611,775,2008,2033]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="56" phylum="Arthropoda" rank="species" species="pantherina">G. pantherina</taxonomicName>
</emphasis>
have also recorded from elasmobranchs. These species can be distinguished from the following features. 
<taxonomicName id="2DBF79B18C76FFAE5ACEFF4DFC6C929B" box="[736,944,152,177]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="capillata">
<emphasis id="D8CBDE208C76FFAE5ACEFF4DFC6C929B" box="[736,944,152,177]" italics="true" pageId="7" pageNumber="57">Gnathia capillata</emphasis>
</taxonomicName>
has a shallow and wide dorsal sulcus and a notch on article 1 of the pylopod (
<bibRefCitation id="8E2E7FC38C76FFAE5A6DFF6BFC4E92F2" author="Nunomura" box="[579,914,190,216]" pageId="7" pageNumber="60" refString="Nunomura, N. &amp; Honma, Y. (2004) Gnathia capillata, a new species of the genus Gnathia (Crustacea, Isopoda) from Sado Island, the Sea of Japan. Contribution from the Biological Laboratory, Kyoto University, 29, 343 - 349." type="journal article" year="2004">Nunomura &amp; Honma, 2004</bibRefCitation>
). The total length of 
<taxonomicName id="2DBF79B18C76FFAE5CA2FF6AFAEE92F2" box="[1164,1330,191,216]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="grandilaris">
<emphasis id="D8CBDE208C76FFAE5CA2FF6AFAEE92F2" box="[1164,1330,191,216]" italics="true" pageId="7" pageNumber="57">G. grandilaris</emphasis>
</taxonomicName>
is larger (
<quantity id="2D47AFD78C76FFAE58B1FF30FEEE92D5" box="[159,306,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.3" metricValueMin="5.7" pageId="7" pageNumber="57" unit="mm" value="7.0" valueMax="8.3" valueMin="5.7">5.7–8.3 mm</quantity>
), and has a medio-frontal process on frontal boarder and more slender pylopod with two areolae (
<bibRefCitation id="8E2E7FC38C76FFAE58B1FED9FE41930C" author="Coetzee" box="[159,413,268,294]" lastPageId="7" lastPageNumber="60" pageId="7" pageNumber="59" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2008) A new gnathiid (Crustacea, Isopoda) parasitising two species of Requiem sharks from Lizard Island, Great Barrier Reef, Australia. Journal of Parasitology 94, 608 - 615." type="journal article" year="2008">
Coetzee 
<emphasis id="D8CBDE208C76FFAE5922FED9FE9F930F" box="[268,323,268,293]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2008
</bibRefCitation>
). 
<taxonomicName id="2DBF79B18C76FFAE599CFED9FD78930F" box="[434,676,268,293]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="pantherina">
<emphasis id="D8CBDE208C76FFAE599CFED9FD78930F" box="[434,676,268,293]" italics="true" pageId="7" pageNumber="57">Gnathia pantherina</emphasis>
</taxonomicName>
has pectinate scales on the pleotelson, fine setae on peduncle articles of the antennae, and no areola on the pylopod (Smit &amp; Basson, 2002).
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C76FFAE58EBFE8CFB0790E5" pageId="7" pageNumber="57" type="reference_group">
<paragraph id="EA0002328C76FFAE58EBFE8CFB5893CD" blockId="7.[151,1436,152,912]" pageId="7" pageNumber="57">
<taxonomicName id="2DBF79B18C76FFAE58EBFE8CFE1B9358" ID-CoL="6KNCC" box="[197,455,345,370]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="camuripenis">
<emphasis id="D8CBDE208C76FFAE58EBFE8CFE1B9358" box="[197,455,345,370]" italics="true" pageId="7" pageNumber="57">Gnathia camuripenis</emphasis>
</taxonomicName>
and 
<emphasis id="D8CBDE208C76FFAE5A2AFE8CFD599358" box="[516,645,345,370]" italics="true" pageId="7" pageNumber="57">G. l i m i c o l a</emphasis>
might be found with 
<emphasis id="D8CBDE208C76FFAE5BBAFE8CFBFA9358" box="[916,1062,345,370]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
, as their locales are near each other. However, 
<taxonomicName id="2DBF79B18C76FFAE5977FE55FDD693B3" ID-CoL="6KNCC" box="[345,522,384,409]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="camuripenis">
<emphasis id="D8CBDE208C76FFAE5977FE55FDD693B3" box="[345,522,384,409]" italics="true" pageId="7" pageNumber="57">G. camuripenis</emphasis>
</taxonomicName>
and 
<taxonomicName id="2DBF79B18C76FFAE5A6DFE55FD1F93B3" ID-CoL="3GLCJ" box="[579,707,384,409]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="limicola">
<emphasis id="D8CBDE208C76FFAE5A6DFE55FD1F93B3" box="[579,707,384,409]" italics="true" pageId="7" pageNumber="57">G. limicola</emphasis>
</taxonomicName>
are smaller (the former is longest 
<quantity id="2D47AFD78C76FFAE5C7BFE55FB6E93B0" box="[1109,1202,384,410]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="57" unit="mm" value="2.8">2.8 mm</quantity>
; the latter is longest 
<quantity id="2D47AFD78C76FFAE58B9FE73FF2893EA" box="[151,244,422,448]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="57" unit="mm" value="2.6">2.6 mm</quantity>
) and have prominent penes fused into a laterally compressed blade (
<bibRefCitation id="8E2E7FC38C76FFAE5C08FE73FB1793EA" author="Tanaka" box="[1062,1227,422,448]" pageId="7" pageNumber="60" refString="Tanaka, K. (2004) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Ishigaki Island, the Ryukyus, southwestern Japan. Crustacean Research, 33, 51 - 60." type="journal article" year="2004">Tanaka, 2004</bibRefCitation>
; 
<bibRefCitation id="8E2E7FC38C76FFAE5CF6FE73FA5393EA" author="Ota" box="[1240,1423,422,448]" pageId="7" pageNumber="60" refString="Ota, Y., Tanaka, K. &amp; Hirose, E. (2007) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Okinawajima Island, Ryukyu Archipelago, southwestern Japan. Zoological Science, 24, 1266 - 1277." type="journal article" year="2007">
Ota 
<emphasis id="D8CBDE208C76FFAE5D25FE72FA9493EA" box="[1291,1352,423,448]" italics="true" pageId="7" pageNumber="57">et al.</emphasis>
2007
</bibRefCitation>
). Thus, adult males of these species are easily distinguished from those of 
<emphasis id="D8CBDE208C76FFAE5BC2FE18FBA593CC" box="[1004,1145,461,486]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
.
</paragraph>
<paragraph id="EA0002328C76FFAE58EBFE21FB0790E5" blockId="7.[151,1436,152,912]" pageId="7" pageNumber="57">
Only a few detailed descriptions of adult females are available for gnathiids. In the genus 
<taxonomicName id="2DBF79B18C76FFAE5CF0FE21FAE29027" ID-CoL="7P3YN" box="[1246,1342,500,525]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="D8CBDE208C76FFAE5CF0FE21FAE29027" box="[1246,1342,500,525]" italics="true" pageId="7" pageNumber="57">Gnathia</emphasis>
</taxonomicName>
, female mouthparts have been described in detail only for the following six species, i.e. 
<emphasis id="D8CBDE208C76FFAE5C19FDCFFB389019" box="[1079,1252,538,563]" italics="true" pageId="7" pageNumber="57">G. c a m u r i p e n i s</emphasis>
, 
<taxonomicName id="2DBF79B18C76FFAE5CDBFDCFFAA99019" ID-CoL="3GLCJ" box="[1269,1397,538,563]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="limicola">
<emphasis id="D8CBDE208C76FFAE5CDBFDCFFAA99019" box="[1269,1397,538,563]" italics="true" pageId="7" pageNumber="57">G. limicola</emphasis>
</taxonomicName>
, 
<taxonomicName id="2DBF79B18C76FFAE5DADFDCFFF279070" ID-CoL="6KNQD" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="D8CBDE208C76FFAE5DADFDCFFF279070" italics="true" pageId="7" pageNumber="57">G. africana</emphasis>
</taxonomicName>
, 
<taxonomicName id="2DBF79B18C76FFAE5924FD94FE779070" ID-CoL="3GLDD" box="[266,427,577,602]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="pantherina">
<emphasis id="D8CBDE208C76FFAE5924FD94FE779070" box="[266,427,577,602]" italics="true" pageId="7" pageNumber="57">G. pantherina</emphasis>
</taxonomicName>
, 
<taxonomicName id="2DBF79B18C76FFAE5994FD94FDBD9070" ID-CoL="3GLBZ" box="[442,609,577,602]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="gurjanovae">
<emphasis id="D8CBDE208C76FFAE5994FD94FDBD9070" box="[442,609,577,602]" italics="true" pageId="7" pageNumber="57">G. gurjanovae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="2DBF79B18C76FFAE5A8DFD94FC909070" ID-CoL="6KND7" box="[675,844,577,602]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C76FFAE5A8DFD94FC909070" box="[675,844,577,602]" italics="true" pageId="7" pageNumber="57">G. trimaculata</emphasis>
</taxonomicName>
(Smit &amp; Basson, 2002; Smit 
<emphasis id="D8CBDE208C76FFAE5C85FD94FB039070" box="[1195,1247,577,602]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2002; 
<bibRefCitation id="8E2E7FC38C76FFAE5D6EFD94FF0A90A8" author="Tanaka" pageId="7" pageNumber="60" refString="Tanaka, K. (2004) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Ishigaki Island, the Ryukyus, southwestern Japan. Crustacean Research, 33, 51 - 60." type="journal article" year="2004">Tanaka, 2004</bibRefCitation>
; 
<bibRefCitation id="8E2E7FC38C76FFAE58CCFDBDFE4A90A8" author="Golovan" box="[226,406,616,642]" pageId="7" pageNumber="60" refString="Golovan, O. A, (2006) Gnathia gurjanovae sp. n., a new species of Gnathiids (Isopoda: Gnathiidae) from Peter the Great Bay, Sea of Japan. Russian Journal of Marine Biology, 32, 28 - 36." type="journal article" year="2006">Golovan, 2006</bibRefCitation>
; 
<bibRefCitation id="8E2E7FC38C76FFAE598CFDBDFD8290A8" author="Ota" box="[418,606,616,642]" pageId="7" pageNumber="60" refString="Ota, Y., Tanaka, K. &amp; Hirose, E. (2007) A new species of Gnathia (Isopoda: Cymothoida: Gnathiidae) from Okinawajima Island, Ryukyu Archipelago, southwestern Japan. Zoological Science, 24, 1266 - 1277." type="journal article" year="2007">
Ota 
<emphasis id="D8CBDE208C76FFAE59FAFDBDFDD590AB" box="[468,521,616,641]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2007
</bibRefCitation>
; 
<bibRefCitation id="8E2E7FC38C76FFAE5A45FDBDFC8190A8" author="Coetzee" box="[619,861,616,642]" pageId="7" pageNumber="60" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2009) Gnathia trimaculata n. sp. (Crustacea: Isopoda: Gnathiidae), an ectoparasite found parasitising requiem sharks from Lizard Island, Great Barrier Reef, Australia. Systematic Parasitology, 72, 97 - 112." type="journal article" year="2009">
Coetzee 
<emphasis id="D8CBDE208C76FFAE5AFCFDBDFCDA90AB" box="[722,774,616,641]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2009
</bibRefCitation>
). Of these species, the females of 
<taxonomicName id="2DBF79B18C76FFAE5CDFFDBDFA4790AB" ID-CoL="6KND7" box="[1265,1435,616,641]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C76FFAE5CDFFDBDFA4790AB" box="[1265,1435,616,641]" italics="true" pageId="7" pageNumber="57">G. trimaculata</emphasis>
</taxonomicName>
are most similar to the females of 
<emphasis id="D8CBDE208C76FFAE5A0AFD5AFD6C9082" box="[548,688,655,680]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
. However, 
<taxonomicName id="2DBF79B18C76FFAE5B15FD5AFC3F9082" ID-CoL="6KND7" box="[827,995,655,680]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C76FFAE5B15FD5AFC3F9082" box="[827,995,655,680]" italics="true" pageId="7" pageNumber="57">G. trimaculata</emphasis>
</taxonomicName>
is distinguished from 
<emphasis id="D8CBDE208C76FFAE5CC6FD5AFAA89082" box="[1256,1396,655,680]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
by a wider pleotelson and the simple pappose setae on the frontal boarder (
<bibRefCitation id="8E2E7FC38C76FFAE5BF0FD60FB1390E5" author="Coetzee" box="[990,1231,693,719]" pageId="7" pageNumber="60" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2009) Gnathia trimaculata n. sp. (Crustacea: Isopoda: Gnathiidae), an ectoparasite found parasitising requiem sharks from Lizard Island, Great Barrier Reef, Australia. Systematic Parasitology, 72, 97 - 112." type="journal article" year="2009">
Coetzee 
<emphasis id="D8CBDE208C76FFAE5C68FD60FBA590E4" box="[1094,1145,693,718]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A2A551B98C76FFAE58EBFD09FC0491BA" pageId="7" pageNumber="57" type="description">
<paragraph id="EA0002328C76FFAE58EBFD09FC0491BA" blockId="7.[151,1436,152,912]" pageId="7" pageNumber="57">
The praniza larvae of 
<taxonomicName id="2DBF79B18C76FFAE59E0FD09FDA890DF" box="[462,628,732,757]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="pantherina">
<emphasis id="D8CBDE208C76FFAE59E0FD09FDA890DF" box="[462,628,732,757]" italics="true" pageId="7" pageNumber="57">G. pantherina</emphasis>
</taxonomicName>
, 
<taxonomicName id="2DBF79B18C76FFAE5AADFD09FCCC90DF" box="[643,784,732,757]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="capillata">
<emphasis id="D8CBDE208C76FFAE5AADFD09FCCC90DF" box="[643,784,732,757]" italics="true" pageId="7" pageNumber="57">G. capillata</emphasis>
</taxonomicName>
, 
<taxonomicName id="2DBF79B18C76FFAE5B31FD09FC1490DF" box="[799,968,732,757]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="grandilaris">
<emphasis id="D8CBDE208C76FFAE5B31FD09FC1490DF" box="[799,968,732,757]" italics="true" pageId="7" pageNumber="57">G. grandilaris</emphasis>
</taxonomicName>
and 
<taxonomicName id="2DBF79B18C76FFAE5C2DFD09FB6D90DF" box="[1027,1201,732,757]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="D8CBDE208C76FFAE5C2DFD09FB6D90DF" box="[1027,1201,732,757]" italics="true" pageId="7" pageNumber="57">G. trimaculata</emphasis>
</taxonomicName>
may be found with those of 
<emphasis id="D8CBDE208C76FFAE58D2FCD7FE559131" box="[252,393,770,795]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
, as they are ectoparasites on elasmobranchs. The apex of the pleotelson does not extend beyond the uropodal rami in these 3 species, except for 
<taxonomicName id="2DBF79B18C76FFAE5B6AFCFCFC0E9168" box="[836,978,809,834]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="capillata">
<emphasis id="D8CBDE208C76FFAE5B6AFCFCFC0E9168" box="[836,978,809,834]" italics="true" pageId="7" pageNumber="57">G. capillata</emphasis>
</taxonomicName>
(Smit &amp; Basson, 2002; 
<bibRefCitation id="8E2E7FC38C76FFAE5CD2FCFCFEE39140" author="Nunomura" pageId="7" pageNumber="60" refString="Nunomura, N. &amp; Honma, Y. (2004) Gnathia capillata, a new species of the genus Gnathia (Crustacea, Isopoda) from Sado Island, the Sea of Japan. Contribution from the Biological Laboratory, Kyoto University, 29, 343 - 349." type="journal article" year="2004">Nunomura &amp; Honma, 2004</bibRefCitation>
; 
<bibRefCitation id="8E2E7FC38C76FFAE5965FC85FD9C9140" author="Coetzee" box="[331,576,848,874]" lastPageId="7" lastPageNumber="60" pageId="7" pageNumber="59" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2008) A new gnathiid (Crustacea, Isopoda) parasitising two species of Requiem sharks from Lizard Island, Great Barrier Reef, Australia. Journal of Parasitology 94, 608 - 615." type="journal article" year="2008">
Coetzee 
<emphasis id="D8CBDE208C76FFAE599AFC85FE359143" box="[436,489,848,873]" italics="true" pageId="7" pageNumber="57">et al</emphasis>
., 2008
</bibRefCitation>
, 
<bibRefCitation id="8E2E7FC38C76FFAE5A62FC85FD519140" author="Coetzee" box="[588,653,848,874]" pageId="7" pageNumber="60" refString="Coetzee, M. L., Smit, N. J., Grutter, A. S. &amp; Davis A. J. (2009) Gnathia trimaculata n. sp. (Crustacea: Isopoda: Gnathiidae), an ectoparasite found parasitising requiem sharks from Lizard Island, Great Barrier Reef, Australia. Systematic Parasitology, 72, 97 - 112." type="journal article" year="2009">2009</bibRefCitation>
). 
<taxonomicName id="2DBF79B18C76FFAE5A8DFC85FCA49143" box="[675,888,848,873]" class="Malacostraca" family="Gnathiidae" genus="Gnathia" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="capillata">
<emphasis id="D8CBDE208C76FFAE5A8DFC85FCA49143" box="[675,888,848,873]" italics="true" pageId="7" pageNumber="57">Gnathia capillata</emphasis>
</taxonomicName>
is distinguished from 
<emphasis id="D8CBDE208C76FFAE5CA6FC85FAC49143" box="[1160,1304,848,873]" italics="true" pageId="7" pageNumber="57">G. m a c u l o s a</emphasis>
by a large plumose seta on peduncle 4 of antenna 2 (
<bibRefCitation id="8E2E7FC38C76FFAE5AACFCA3FC1791BA" author="Nunomura" box="[642,971,886,912]" pageId="7" pageNumber="60" refString="Nunomura, N. &amp; Honma, Y. (2004) Gnathia capillata, a new species of the genus Gnathia (Crustacea, Isopoda) from Sado Island, the Sea of Japan. Contribution from the Biological Laboratory, Kyoto University, 29, 343 - 349." type="journal article" year="2004">Nunomura &amp; Honma, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>