214 lines
31 KiB
XML
214 lines
31 KiB
XML
<document id="4689C74648E39979487507CD7CD2C8F4" ID-DOI="10.50826/bnmnszool.48.3_119" ID-ISSN="2434-091X" ID-Zenodo-Dep="12760044" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721224709816" checkinUser="felipe" docAuthor="Karaoğlan, Beliz Bahar, Yekedüz, Emre, Yazgan, Satı Coşkun, Mocan, Eda Eylemer, Köksoy, Elif Berna, Yaşar, Hatime Arzu, Şenler, Filiz Çay, Utkan, Güngör, Demirkazık, Ahmet, Akbulut, Hakan & Ürün, Yüksel" docDate="2022" docId="A95B8783390AFFE7FD47FD72FB0EC3BF" docLanguage="en" docName="BullNatlMusNatSciSerA.48.3.119-138.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology (Zoology) 48 (3)" docSource="http://dx.doi.org/10.1080/1750743X.2024.2370231" docStyle="DocumentStyle:EEA6C589EB70DFF4CCECB2264A05AF4C.2:BullNatlMusNatSciSerA.2021-.journal_article.english.noindex" docStyleId="EEA6C589EB70DFF4CCECB2264A05AF4C" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.noindex" docStyleVersion="2" docTitle="Nipponaphaenops (Tanakaphaenops) Karaoğlan, Yekedüz, Yazgan, Mocan, Köksoy, Yaşar, Şenler, Utkan, Demirkazık, Akbulut & Ürün, 2022, subgen. nov." docType="treatment" docVersion="2" lastPageNumber="132" masterDocId="5562FFFB3906FFEAFFB7FFD5FFDBC705" masterDocTitle="Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan" masterLastPageNumber="138" masterPageNumber="119" pageNumber="131" updateTime="1721225289731" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F5A7114312FF57FC12C8A712C62D119D">Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan</mods:title>
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<mods:namePart id="AF0CBEF086CB3CB91CCA9A376A392ABB">Karaoğlan, Beliz Bahar</mods:namePart>
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<mods:namePart id="0C901C8CDE6CD0AD826323982233032B">Yekedüz, Emre</mods:namePart>
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<mods:namePart id="73D1285666EA6D656A3C754F36D0306B">Yazgan, Satı Coşkun</mods:namePart>
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<mods:namePart id="0ABC531F169B8A4F12304473C0196726">Mocan, Eda Eylemer</mods:namePart>
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<mods:namePart id="83568DE0B3BA54E45526D652631E2C00">Köksoy, Elif Berna</mods:namePart>
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<mods:namePart id="F07B2C57ABF4A5CFC9FA32ADD0A9B163">Yaşar, Hatime Arzu</mods:namePart>
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<mods:namePart id="23A31C9E1CFFD31BFB5AFAEE84EC6243">Şenler, Filiz Çay</mods:namePart>
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<mods:namePart id="139FF3CB8E27DF5A38AD6665483899F0">Utkan, Güngör</mods:namePart>
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<mods:namePart id="CC614F83DAE685D27FEFE384F0AD08D6">Demirkazık, Ahmet</mods:namePart>
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<mods:namePart id="67B70C98CE6EA3A30098DE12E4EB4651">Akbulut, Hakan</mods:namePart>
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<mods:namePart id="8F15D2A2F0447DF4B77C2A6CDFC30F70">Ürün, Yüksel</mods:namePart>
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<mods:title id="0650749901CA651EA3AD95BC74EFB2C3">Bulletin of the National Museum of Nature and Science. Series A, Zoology</mods:title>
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<mods:date id="3847C9FAFA97486043AB310C9F9DE692">2022</mods:date>
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<mods:title id="BF63A670ACD734D1145C1B32D6BB2921">Zoology</mods:title>
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<mods:number id="5D70457D28668C1545A9FEB4121487B9">2022-08-22</mods:number>
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<mods:number id="F73487ADE34531438FBD3E425CBF5B78">48</mods:number>
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<mods:start id="DB4E83046F84C159D2EA09D94B8D1590">119</mods:start>
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<mods:identifier id="7BD5E8272AA5F07805F4008453115059" type="ISSN">2434-091X</mods:identifier>
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<treatment id="A95B8783390AFFE7FD47FD72FB0EC3BF" LSID="urn:lsid:plazi:treatment:A95B8783390AFFE7FD47FD72FB0EC3BF" httpUri="http://treatment.plazi.org/id/A95B8783390AFFE7FD47FD72FB0EC3BF" lastPageId="13" lastPageNumber="132" pageId="12" pageNumber="131">
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<paragraph id="214D3695390AFFE6FD47FD72FB7CC5BA" blockId="12.[752,1191,679,703]" box="[752,1191,679,703]" pageId="12" pageNumber="131">
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<heading id="7A0581F9390AFFE6FD47FD72FB7CC5BA" box="[752,1191,679,703]" centered="true" fontSize="36" level="2" pageId="12" pageNumber="131" reason="3">
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Subgenus
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<taxonomicName id="E6F24D16390AFFE6FCE8FD72FBC1C5BA" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[863,1050,679,703]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Tanakaphaenops">
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<emphasis id="1386EA87390AFFE6FCE8FD72FBC1C5BA" bold="true" box="[863,1050,679,703]" italics="true" pageId="12" pageNumber="131">Tanakaphaenops</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="08B557FC390AFFE6FB96FD72FB7CC5BA" box="[1057,1191,679,703]" pageId="12" pageNumber="131" rank="subGenus">subgen. nov.</taxonomicNameLabel>
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<paragraph id="214D3695390AFFE6FD74FD03FC5BC40F" blockId="12.[707,1236,726,778]" pageId="12" pageNumber="131">
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urn:lsid:zoobank.org:act:
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<subSubSection id="69E8651E390AFFE7FD6AFC92FB0EC3BF" lastPageId="13" lastPageNumber="132" pageId="12" pageNumber="131" type="description">
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<paragraph id="214D3695390AFFE6FD6AFC92FB13C45A" blockId="12.[707,1237,839,1799]" box="[733,1224,839,863]" pageId="12" pageNumber="131">
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Type-species:
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<taxonomicName id="E6F24D16390AFFE6FCC1FC92FBB5C45A" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[886,1134,839,863]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="kojii" status="sp. nov.">
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<emphasis id="1386EA87390AFFE6FCC1FC92FBB5C45A" box="[886,1134,839,863]" italics="true" pageId="12" pageNumber="131">Nipponaphaenops kojii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="08B557FC390AFFE6FBC2FC9DFB13C45A" box="[1141,1224,840,863]" pageId="12" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="214D3695390AFFE6FD6AFCBEFC87C178" blockId="12.[707,1237,839,1799]" pageId="12" pageNumber="131">
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<emphasis id="1386EA87390AFFE6FD6AFCBEFC05C487" box="[733,990,874,898]" italics="true" pageId="12" pageNumber="131">Diagnostic characters.</emphasis>
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Discriminated from the nominate subgenus by the following character states; body size smaller (BL:
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<quantity id="E60A9B70390AFFE6FB9FFC7AFB52C4C2" box="[1064,1161,943,967]" metricMagnitude="-3" metricUnit="m" metricValue="5.39" pageId="12" pageNumber="131" unit="mm" value="5.39">5.39 mm</quantity>
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); head smaller, shorter (PW/HW 1.30, PL/HL 1.50, HW/ HL 0.98); frontal furrows complete; tempora to genae with a temporal seta in addition to a genal seta; penultimate labial palpomere about 1.05 times longer than ultimate one, with two subapical setae on the external face of the former usual in size (
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<figureCitation id="B9C92A10390AFFE6FC97FB77FCA8C3BF" box="[800,883,1186,1210]" captionStart="Fig" captionStartId="4.[163,196,919,941]" captionTargetBox="[177,1199,214,885]" captionTargetId="figure-331@4.[176,1200,213,886]" captionTargetPageId="4" captionText="Fig. 1. Right mandible in dorsal view (a–i) and labial (right) and maxillary (left) palpi in ventral view (j–p).—a, Chaetotrechiama procerus S. Uéno; b, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); c & l, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d & m, R. (Uozumitrechus) sp.; e, R. (Pilosotrechiama) mirabilis S. Uéno; f & k, R. (Izushites) sp.; g & j, R. (Rakantrechus) kawasawai S. Uéno; h & o, Nipponaphaenops (N.) erraticus S. Uéno; i & n, N. (Tanakaphaenops) kojii subgen. et sp. nov.; p, Trechiama pluto S. Uéno (this species is unique in having trisetose penultimate labial palpomere in the Trechiama Series)." figureDoi="http://doi.org/10.5281/zenodo.12760048" httpUri="https://zenodo.org/record/12760048/files/figure.png" pageId="12" pageNumber="131">Fig. 1n</figureCitation>
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); antennae shorter, stouter, less than a half longer than elytra; pronotum with hind angles obtuse at the tip; posterior latero-marginal setae of pronotum present; elytra shorter (EL/EW 1.39), with prehumeral borders shorter and more straight; pore 1 of the marginal umbilicate series of elytra more distant from marginal gutter and closer to elytral base (U1 15.8), pore 4 of the series closer to elytral base (U4 29.2); elytral discal seta of the external series more externally (in stria 5), more posteriorly (basal 11/17) positioned; legs shorter and stouter; ventrites 4–6 each with a pair of paramedian setae (
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<figureCitation id="B9C92A10390AFFE6FCB0F9B0FC8EC178" box="[775,853,1637,1661]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="12" pageNumber="131">Fig. 2k</figureCitation>
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)
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</paragraph>
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<paragraph id="214D3695390AFFE6FD6AF95DFB1BC1A5" blockId="12.[707,1237,839,1799]" box="[733,1216,1671,1696]" pageId="12" pageNumber="131">
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<emphasis id="1386EA87390AFFE6FD6AF95DFCB8C19A" box="[733,867,1672,1695]" italics="true" pageId="12" pageNumber="131">Description.</emphasis>
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Body size small for the genus.
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</paragraph>
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<paragraph id="214D3695390AFFE7FD6AF97FFEA8C6E7" blockId="12.[707,1237,839,1799]" lastBlockId="13.[140,669,215,1245]" lastPageId="13" lastPageNumber="132" pageId="12" pageNumber="131">
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Chaetotaxy of head as in the nominate subgenus except that genae with a temporal seta; both two pairs of latero-marginal setae of pronotum present; elytral discal and preapical setae as in the nominate subgenus; pore 1 of the humeral set of marginal umbilicate series of elytra postero-dorsally translocated, positioned on stria 7 and close to the level of the pore 2 of the series, and pore 4 of the series widely distant from pore 3 (
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<figureCitation id="B9C92A10390BFFE7FF23FE72FF3DC6BA" box="[148,230,423,447]" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="13" pageNumber="132">Fig. 3d</figureCitation>
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). Ventrite 7 of female with two pairs of apical setae (
|
||
<figureCitation id="B9C92A10390BFFE7FEA1FE1FFEBDC6E7" box="[278,358,458,482]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="13" pageNumber="132">Fig. 2k</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="214D3695390BFFE7FF10FE38FEFAC45A" blockId="13.[140,669,215,1245]" pageId="13" pageNumber="132">
|
||
Head elongate subquadrate; frontal furrows complete, deep in most part, though becoming superficial behind the level of posterior supraorbital pores; labial suture nearly completely vanished; mentum tooth relatively long and slender, sharply bifid at apex (
|
||
<figureCitation id="B9C92A10390BFFE7FE3BFD4FFE3BC5B7" box="[396,480,666,690]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="13" pageNumber="132">Fig. 2e</figureCitation>
|
||
); right mandible (
|
||
<figureCitation id="B9C92A10390BFFE7FF23FD68FF01C5D0" box="[148,218,701,725]" captionStart="Fig" captionStartId="4.[163,196,919,941]" captionTargetBox="[177,1199,214,885]" captionTargetId="figure-331@4.[176,1200,213,886]" captionTargetPageId="4" captionText="Fig. 1. Right mandible in dorsal view (a–i) and labial (right) and maxillary (left) palpi in ventral view (j–p).—a, Chaetotrechiama procerus S. Uéno; b, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); c & l, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d & m, R. (Uozumitrechus) sp.; e, R. (Pilosotrechiama) mirabilis S. Uéno; f & k, R. (Izushites) sp.; g & j, R. (Rakantrechus) kawasawai S. Uéno; h & o, Nipponaphaenops (N.) erraticus S. Uéno; i & n, N. (Tanakaphaenops) kojii subgen. et sp. nov.; p, Trechiama pluto S. Uéno (this species is unique in having trisetose penultimate labial palpomere in the Trechiama Series)." figureDoi="http://doi.org/10.5281/zenodo.12760048" httpUri="https://zenodo.org/record/12760048/files/figure.png" pageId="13" pageNumber="132">Fig. 1i</figureCitation>
|
||
) with a tricuspid retinacular tooth (which is supposedly formed by the union of premolar and primarily bicuspid retinaculum) whose median cusp is rather reduced (cf.
|
||
<bibRefCitation id="45634B64390BFFE7FDB8FCF0FF1FC45A" author="Deuve, Th. & D. H. Kavanaugh & H. Liang" pageId="13" pageNumber="132" pagination="341 - 455" refId="ref12726" refString="Deuve, Th., D. H. Kavanaugh and H. Liang 2016. Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Trechini (Coleoptera, Caraboidea), with descriptions of four new genera, one new subgenus and 19 new species. Proceedings of the California academy of sciences, Series 4, 63: 341 - 455, 48 figs." type="journal article" year="2016">
|
||
Deuve
|
||
<emphasis id="1386EA87390BFFE7FDE9FCF0FD4CC438" box="[606,663,805,829]" italics="true" pageId="13" pageNumber="132">et al.</emphasis>
|
||
, 2016
|
||
</bibRefCitation>
|
||
, p. 380).
|
||
</paragraph>
|
||
<paragraph id="214D3695390BFFE7FF10FCBFFD50C4EF" blockId="13.[140,669,215,1245]" pageId="13" pageNumber="132">Pronotum elongate subcordate, wider than head, obviously longer than wide, with front angles rounded but advanced, with hind angles as a whole subrectangular though obtuse at the tip.</paragraph>
|
||
<paragraph id="214D3695390BFFE7FF10FC20FC1CC522" blockId="13.[140,669,215,1245]" lastBlockId="13.[707,1237,215,1210]" pageId="13" pageNumber="132">
|
||
Elytra large, oval, with basal parts strongly rectilinearly constricted basad, with prehumeral marginations becoming evanescent anteriad a short way before shoulders; baso-lateral areas of elytral disc just before shoulders nearly vertically deflexed ventrad, forming oblique longitudinal carinae consisted of basal parts of intervals 6–7, with the outlines of prehumeral portion of elytra barely visible from above beneath these carinae (
|
||
<figureCitation id="B9C92A10390BFFE7FD7CFEC8FCC0C630" box="[715,795,285,309]" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="13" pageNumber="132">Fig. 3d</figureCitation>
|
||
); basal portions of intervals 2–5 more or less depressed, while basal parts of interval 1 including scutellar area left undepressed as in the nominate subgenus; inclined face of basal peduncle rather narrowly flat, embraced in gentle slope, which is bordered posteriorly by the obtuse transverse carinae formed by the basalmost portion of elytral disc (
|
||
<figureCitation id="B9C92A10390BFFE7FCF0FDDAFC60C522" box="[839,955,527,551]" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="13" pageNumber="132">Figs. 3c–d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="214D3695390BFFE7FD6AFDE7FBFFC54C" blockId="13.[707,1237,215,1210]" box="[733,1060,562,586]" pageId="13" pageNumber="132">Legs long, but relatively stout.</paragraph>
|
||
<paragraph id="214D3695390BFFE7FD6AFD80FBDEC568" blockId="13.[707,1237,215,1210]" box="[733,1029,597,621]" pageId="13" pageNumber="132">Male characters. Unknown.</paragraph>
|
||
<paragraph id="214D3695390BFFE7FD6AFDADFCB2C4EF" blockId="13.[707,1237,215,1210]" pageId="13" pageNumber="132">
|
||
<emphasis id="1386EA87390BFFE7FD6AFDADFC41C58A" box="[733,922,632,655]" italics="true" pageId="13" pageNumber="132">Taxonomic notes.</emphasis>
|
||
This subgenus is presently monotypic. In this subgenus (as far as known from the
|
||
<typeStatus id="FE498837390BFFE7FC99FD68FC87C5D0" box="[814,860,701,725]" pageId="13" pageNumber="132">type</typeStatus>
|
||
species), the position of the first pore of the marginal umbilicate series of elytra is most strongly postero-dorsally translocated within the
|
||
<taxonomicName id="E6F24D16390BFFE7FC80FCF0FBDCC438" authority="Phyletic" authorityName="Phyletic" box="[823,1031,805,829]" class="Insecta" family="Carabidae" genus="Trechiama" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA87390BFFE7FC80FCF0FC73C438" box="[823,936,805,829]" italics="true" pageId="13" pageNumber="132">Trechiama</emphasis>
|
||
Phyletic
|
||
</taxonomicName>
|
||
Series in a current sense (cf.
|
||
<figureCitation id="B9C92A10390BFFE7FC81FC92FC15C45A" box="[822,974,839,863]" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="13" pageNumber="132">Figs. 3d, g–h</figureCitation>
|
||
). However, in Japanese trechines as a whole, the pore is most strongly postero-dorsally translocated in
|
||
<taxonomicName id="E6F24D16390BFFE7FB96FC58FC94C4C2" pageId="13" pageNumber="132">
|
||
<emphasis id="1386EA87390BFFE7FB96FC58FBB6C4A0" box="[1057,1133,909,933]" italics="true" pageId="13" pageNumber="132">Suzuka</emphasis>
|
||
S. Uéno, 1956 species
|
||
</taxonomicName>
|
||
, which belongs to the
|
||
<taxonomicName id="E6F24D16390BFFE7FB88FC7AFCC1C4EF" authority="Phyletic" authorityName="Phyletic" class="Insecta" family="Carabidae" genus="Trechoblemus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA87390BFFE7FB88FC7AFB0FC4C2" box="[1087,1236,943,967]" italics="true" pageId="13" pageNumber="132">Trechoblemus</emphasis>
|
||
Phyletic
|
||
</taxonomicName>
|
||
Series.
|
||
</paragraph>
|
||
<paragraph id="214D3695390BFFE7FD6AFC20FBA8C357" blockId="13.[707,1237,215,1210]" pageId="13" pageNumber="132">
|
||
<emphasis id="1386EA87390BFFE7FD6AFC20FC60C309" box="[733,955,1013,1037]" italics="true" pageId="13" pageNumber="132">Geographic range.</emphasis>
|
||
Northwestern area of Shikoku, southwestern
|
||
<collectingCountry id="59E57605390BFFE7FC7EFBCDFBD2C32A" box="[969,1033,1048,1071]" name="Japan" pageId="13" pageNumber="132">Japan</collectingCountry>
|
||
; northwest of the range of the nominate subgenus (
|
||
<figureCitation id="B9C92A10390BFFE7FB92FBEFFBBDC357" box="[1061,1126,1082,1106]" captionStart="Fig" captionStartId="16.[163,196,1601,1623]" captionTargetBox="[157,1217,605,1567]" captionTargetId="figure-172@16.[157,1218,605,1568]" captionTargetPageId="16" captionText="Fig. 7. Map showing the distribution of the Rakantrechus Complex (in part) and anophthalmic species of the genus Trechiama in western Japan.—Light green square, Rakantrechus (s. str.); orange square, R. (Izushites); green circle, R. (Iyotrechus); yellow circle, R. (Pilosotrechiama); green triangle, Nipponaphaenops (s. str.); blue open triangle, N. (Tanakaphaenops) subgen. nov.; purple obtriangle, Ishikawatrechus; large purple circle, anophthalmic Trechiama species.—An, the type locality of Ishikawatrechus annulus Sugaya et Hara; Tk, the type locality of Nipponaphaenops (Tanakaphaenops) kojii sugen. et sp. nov. The course of the Hôyo River in the Last Glacial Maximum was drawn according to Kuwashiro (1959)." figureDoi="http://doi.org/10.5281/zenodo.12760060" httpUri="https://zenodo.org/record/12760060/files/figure.png" pageId="13" pageNumber="132">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="214D3695390BFFE7FD6AFB88FB0EC3BF" blockId="13.[707,1237,215,1210]" pageId="13" pageNumber="132">
|
||
<emphasis id="1386EA87390BFFE7FD6AFB88FC94C370" box="[733,847,1117,1141]" italics="true" pageId="13" pageNumber="132">Etymology</emphasis>
|
||
. This new subgenus is named in honour of Mr. Kôji Tanaka, who discovered the
|
||
<typeStatus id="FE498837390BFFE7FD74FB77FD2AC3BF" box="[707,753,1186,1210]" pageId="13" pageNumber="132">type</typeStatus>
|
||
species of this subgenus. Gender: masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |