179 lines
18 KiB
XML
179 lines
18 KiB
XML
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<mods:title id="8D6B143936A5F1EE1DF4A95D5164B7C3">Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan</mods:title>
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<mods:namePart id="D09B34AF2022F5F59DD88955F32A7A42">Karaoğlan, Beliz Bahar</mods:namePart>
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<mods:namePart id="9E688A05C3B4A5D4ECAAB802D34FA9D2">Yekedüz, Emre</mods:namePart>
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<mods:namePart id="FA0554ACA078F47718EA806A2F6ACE34">Yazgan, Satı Coşkun</mods:namePart>
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<mods:namePart id="5E6FA55FC83A7DE20CC42AB8D5FE6FA7">Yaşar, Hatime Arzu</mods:namePart>
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<mods:namePart id="7C2285274AC60132CFE9E124EF79DB35">Şenler, Filiz Çay</mods:namePart>
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<mods:namePart id="6EDCAB4D353B5FDD2CD38E07EF28BB4C">Utkan, Güngör</mods:namePart>
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<mods:namePart id="5AEE3034E8DA10EBCDD8CA0EA5578A29">Akbulut, Hakan</mods:namePart>
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<mods:namePart id="4BD36A1F6EEFACE3C261A56D0777A14D">Ürün, Yüksel</mods:namePart>
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<mods:title id="A7601EF64AE282AFBC4EBD30B96D2A49">Bulletin of the National Museum of Nature and Science. Series A, Zoology</mods:title>
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<mods:date id="B9C35DFFB5E4459EC98EB6E56AC7EAFC">2022</mods:date>
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<mods:title id="4390156F1FA55C14BB4CEB4695AF7C2C">Zoology</mods:title>
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<mods:number id="9E75C6320CF46A483F4C4277ADD805F8">2022-08-22</mods:number>
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<mods:number id="0FCF6C99F2C41AECE33248C68636B4CC">48</mods:number>
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<mods:start id="C7BE04721201A45258E8461FF12FEC8C">119</mods:start>
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<treatment id="A95B8783390AFFE6FF1DFC80FD3BC568" ID-DOI="http://doi.org/10.5281/zenodo.12760131" ID-Zenodo-Dep="12760131" LSID="urn:lsid:plazi:treatment:A95B8783390AFFE6FF1DFC80FD3BC568" httpUri="http://treatment.plazi.org/id/A95B8783390AFFE6FF1DFC80FD3BC568" lastPageNumber="131" pageId="12" pageNumber="131">
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<subSubSection id="69E8651E390AFFE6FF1DFC80FEFDC002" pageId="12" pageNumber="131" type="nomenclature">
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<heading id="7A0581F9390AFFE6FF1DFC80FDA4C468" box="[170,639,852,877]" centered="true" fontSize="36" level="2" pageId="12" pageNumber="131" reason="3">
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Subgenus
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<taxonomicName id="E6F24D16390AFFE6FEAFFC81FDA4C468" ID-CoL="8KXYH" authority="S. Ueno, 1971" authorityName="S. Ueno" authorityYear="1971" box="[280,639,852,877]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis id="1386EA87390AFFE6FEAFFC81FE3BC469" bold="true" box="[280,480,852,876]" italics="true" pageId="12" pageNumber="131">Nipponaphaenops</emphasis>
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S.
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<bibRefCitation id="45634B64390AFFE6FDB4FC80FDA4C468" author="Ueno, S. - I." box="[515,639,853,877]" pageId="12" pageNumber="131" pagination="451 - 462" refId="ref13267" refString="Ueno, S. - I. 1971. Occurrence of an aphaenopsoid trechine beetle in Japan. Annales de speleologie 26: 451 - 462." type="journal article" year="1971">Uéno, 1971</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="214D3695390AFFE6FF10FC58FEFDC002" blockId="12.[140,671,909,1799]" pageId="12" pageNumber="131">
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<emphasis id="1386EA87390AFFE6FF10FC58FE47C4A0" box="[167,412,909,933]" italics="true" pageId="12" pageNumber="131">Diagnostic characters.</emphasis>
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<materialsCitation id="919A3CC8390AFFE6FE03FC58FEF9C002" collectionCode="BL, PW, PL, HW, EL" pageId="12" pageNumber="131" specimenCode="HW 1.14, 1.27, HL 1.08, 1.20, EW 1.48, 1.57" specimenCount="1">
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As far as known from the
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<typeStatus id="FE498837390AFFE6FF01FC65FF3FC4CD" box="[182,228,944,968]" pageId="12" pageNumber="131">type</typeStatus>
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species, this subgenus is different from another subgenus in the following character states; larger (
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<collectionCode id="47E3AE50390AFFE6FE92FC20FE97C308" box="[293,332,1013,1037]" pageId="12" pageNumber="131">BL</collectionCode>
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:
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); head more macrocephalic (
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<collectionCode id="47E3AE50390AFFE6FEA3FBC2FEE5C32A" box="[276,318,1047,1071]" country="Thailand" name="Paleontological Collections" pageId="12" pageNumber="131">PW</collectionCode>
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/
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<specimenCode id="71549EEE390AFFE6FEF1FBC2FE68C32A" box="[326,435,1047,1071]" collectionCode="HW" pageId="12" pageNumber="131">HW 1.14</specimenCode>
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–
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<specimenCode id="71549EEE390AFFE6FE74FBC2FE2FC32A" box="[451,500,1047,1071]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="12" pageNumber="131">1.27</specimenCode>
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,
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<collectionCode id="47E3AE50390AFFE6FDB1FBC2FDFDC32A" box="[518,550,1047,1071]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="12" pageNumber="131" type="Museum">PL</collectionCode>
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/
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<specimenCode id="71549EEE390AFFE6FD9AFBC2FD55C32A" box="[557,654,1047,1071]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="12" pageNumber="131" type="Museum">HL 1.08</specimenCode>
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–
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<specimenCode id="71549EEE390AFFE6FF3BFBEFFF65C357" box="[140,190,1082,1106]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="12" pageNumber="131" type="Museum">1.20</specimenCode>
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), more elongate (
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<collectionCode id="47E3AE50390AFFE6FEC1FBEFFE78C357" box="[374,419,1082,1106]" pageId="12" pageNumber="131">HW</collectionCode>
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/HL 0.76–0.84); frontal furrows abbreviated posteriorly; tempora to genae only with a genal seta; penultimate labial palpomere about 1.15 times longer than ultimate one, with two subapical setae on the external face of the former reduced in size (
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<figureCitation id="B9C92A10390AFFE6FE6FFB32FDFCC3FA" box="[472,551,1255,1279]" captionStart="Fig" captionStartId="4.[163,196,919,941]" captionTargetBox="[177,1199,214,885]" captionTargetId="figure-331@4.[176,1200,213,886]" captionTargetPageId="4" captionText="Fig. 1. Right mandible in dorsal view (a–i) and labial (right) and maxillary (left) palpi in ventral view (j–p).—a, Chaetotrechiama procerus S. Uéno; b, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); c & l, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d & m, R. (Uozumitrechus) sp.; e, R. (Pilosotrechiama) mirabilis S. Uéno; f & k, R. (Izushites) sp.; g & j, R. (Rakantrechus) kawasawai S. Uéno; h & o, Nipponaphaenops (N.) erraticus S. Uéno; i & n, N. (Tanakaphaenops) kojii subgen. et sp. nov.; p, Trechiama pluto S. Uéno (this species is unique in having trisetose penultimate labial palpomere in the Trechiama Series)." figureDoi="http://doi.org/10.5281/zenodo.12760048" httpUri="https://zenodo.org/record/12760048/files/figure.png" pageId="12" pageNumber="131">Fig. 1o</figureCitation>
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); antennae longer, slenderer, more than a half longer than elytra; pronotum with hind angles with acute denticle produced posteriad; posterior latero-marginal setae of pronotum absent; elytra more elongate (
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<collectionCode id="47E3AE50390AFFE6FF41FA40FECCC2A8" box="[246,279,1429,1453]" pageId="12" pageNumber="131">EL</collectionCode>
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/
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||
<specimenCode id="71549EEE390AFFE6FEA9FA40FE5AC2A9" box="[286,385,1428,1453]" collectionCode="EW" pageId="12" pageNumber="131">EW 1.48</specimenCode>
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–
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<specimenCode id="71549EEE390AFFE6FE38FA41FE64C2A9" box="[399,447,1428,1452]" collectionCode="EL" pageId="12" pageNumber="131">1.57</specimenCode>
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), with longer prehumeral borders more arcuately sinuate; pore 1 of the marginal umbilicate series of elytra closer to marginal gutter and more distant from elytral base (U1 17.2–18.8), pore 4 of the series more distant from elytral base (U4 33.6–36.1); elytral discal seta of the external series more inwardly (in interval 4), more anteriorly (basal 6/11–10/17) positioned; legs longer and slenderer; ventrites 4–6 each with two or three pairs of paramedian setae (
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<figureCitation id="B9C92A10390AFFE6FF67F93AFEC2C002" box="[208,281,1775,1799]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="12" pageNumber="131">Fig. 2l</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="69E8651E390AFFE6FD6AFF0DFD3BC568" pageId="12" pageNumber="131" type="description">
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<paragraph id="214D3695390AFFE6FD6AFF0DFBA9C500" blockId="12.[707,1236,215,621]" pageId="12" pageNumber="131">
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<emphasis id="1386EA87390AFFE6FD6AFF0DFC41C7EA" box="[733,922,216,239]" italics="true" pageId="12" pageNumber="131">Taxonomic notes.</emphasis>
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This subgenus is presently monotypic. One of the diagnostic characters of this subgenus, posteriorly abbreviated frontal furrows, makes the
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<typeStatus id="FE498837390AFFE6FC3BFE95FC61C65D" box="[908,954,320,344]" pageId="12" pageNumber="131">type</typeStatus>
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species, in combination with the so-called aphaenopsian body form, a unique truly aphaenopsian representative within the members of the
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<taxonomicName id="E6F24D16390AFFE6FC2FFE72FBBCC6BA" authority="Phyletic" authorityName="Phyletic" box="[920,1127,423,447]" class="Insecta" family="Carabidae" genus="Trechiama" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="genus">
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<emphasis id="1386EA87390AFFE6FC2FFE72FBD2C6BA" box="[920,1033,423,447]" italics="true" pageId="12" pageNumber="131">Trechiama</emphasis>
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Phyletic
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</taxonomicName>
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Series, as far as the currently included members of the series are considered (
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<bibRefCitation id="45634B64390AFFE6FC18FE38FBEBC500" author="Ueno, S. - I." box="[943,1072,493,517]" pageId="12" pageNumber="131" pagination="1 - 13" refId="ref13334" refString="Ueno, S. - I. 1978. The cave trechines of the genus Allotrechiama (Coleoptera, Trechinae). Journal of the Speleological Society of Japan 3: 1 - 13." type="journal article" year="1978">Uéno, 1978</bibRefCitation>
|
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, p. 9).
|
||
</paragraph>
|
||
<paragraph id="214D3695390AFFE6FD6AFDDAFD3BC568" blockId="12.[707,1236,215,621]" pageId="12" pageNumber="131">
|
||
<emphasis id="1386EA87390AFFE6FD6AFDDAFC70C522" box="[733,939,527,551]" italics="true" pageId="12" pageNumber="131">Geographic range.</emphasis>
|
||
Ohnogahara Karst, west-central area of Shikoku, southwestern
|
||
<collectingCountry id="59E57605390AFFE6FBEAFDE7FB41C54C" box="[1117,1178,562,585]" name="Japan" pageId="12" pageNumber="131">Japan</collectingCountry>
|
||
(
|
||
<figureCitation id="B9C92A10390AFFE6FB1EFDE7FD08C568" captionStart="Fig" captionStartId="16.[163,196,1601,1623]" captionTargetBox="[157,1217,605,1567]" captionTargetId="figure-172@16.[157,1218,605,1568]" captionTargetPageId="16" captionText="Fig. 7. Map showing the distribution of the Rakantrechus Complex (in part) and anophthalmic species of the genus Trechiama in western Japan.—Light green square, Rakantrechus (s. str.); orange square, R. (Izushites); green circle, R. (Iyotrechus); yellow circle, R. (Pilosotrechiama); green triangle, Nipponaphaenops (s. str.); blue open triangle, N. (Tanakaphaenops) subgen. nov.; purple obtriangle, Ishikawatrechus; large purple circle, anophthalmic Trechiama species.—An, the type locality of Ishikawatrechus annulus Sugaya et Hara; Tk, the type locality of Nipponaphaenops (Tanakaphaenops) kojii sugen. et sp. nov. The course of the Hôyo River in the Last Glacial Maximum was drawn according to Kuwashiro (1959)." figureDoi="http://doi.org/10.5281/zenodo.12760060" httpUri="https://zenodo.org/record/12760060/files/figure.png" pageId="12" pageNumber="131">Fig. 7</figureCitation>
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).
|
||
</paragraph>
|
||
</subSubSection>
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</treatment>
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</document> |