407 lines
54 KiB
XML
407 lines
54 KiB
XML
<document id="7A41D9DB84FDB987FFE738E051F3B898" ID-DOI="10.50826/bnmnszool.48.3_119" ID-ISSN="2434-091X" ID-Zenodo-Dep="12760044" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721224709816" checkinUser="felipe" docAuthor="Karaoğlan, Beliz Bahar, Yekedüz, Emre, Yazgan, Satı Coşkun, Mocan, Eda Eylemer, Köksoy, Elif Berna, Yaşar, Hatime Arzu, Şenler, Filiz Çay, Utkan, Güngör, Demirkazık, Ahmet, Akbulut, Hakan & Ürün, Yüksel" docDate="2022" docId="A95B87833908FFFBFF1CFF37FE06C67F" docLanguage="en" docName="BullNatlMusNatSciSerA.48.3.119-138.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology (Zoology) 48 (3)" docSource="http://dx.doi.org/10.1080/1750743X.2024.2370231" docStyle="DocumentStyle:EEA6C589EB70DFF4CCECB2264A05AF4C.2:BullNatlMusNatSciSerA.2021-.journal_article.english.noindex" docStyleId="EEA6C589EB70DFF4CCECB2264A05AF4C" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.noindex" docStyleVersion="2" docTitle="Nipponaphaenops (Tanakaphaenops) kojii Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün 2022" docType="treatment" docVersion="2" lastPageNumber="136" masterDocId="5562FFFB3906FFEAFFB7FFD5FFDBC705" masterDocTitle="Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan" masterLastPageNumber="138" masterPageNumber="119" pageNumber="133" updateTime="1721225289731" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FB92F8A5665CA66F094887AF5403AA83">Contribution to the Knowledge of the Rakantrechus Complex (Coleoptera, Carabidae, Trechini), with Description of a New Subgenus and a New Species of the Genus Nipponaphaenops S. Uéno, 1971 from Northwestern Shikoku, Western Japan</mods:title>
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<mods:namePart id="845921F57C50F7D00AA7C78BB879FE33">Karaoğlan, Beliz Bahar</mods:namePart>
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<mods:namePart id="EBF86B1CE16D80CE933E0E786CCB9946">Yekedüz, Emre</mods:namePart>
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<mods:namePart id="0F92EB433B23ECDB209910003734981B">Yazgan, Satı Coşkun</mods:namePart>
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<mods:namePart id="1A22C772FB419C64D9961BC192207080">Yaşar, Hatime Arzu</mods:namePart>
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<mods:namePart id="24025D427D1ADE671670FD2ABED961D4">Şenler, Filiz Çay</mods:namePart>
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<mods:namePart id="BEAB8872A5C3E79994FA9A23735E4B86">Utkan, Güngör</mods:namePart>
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<mods:namePart id="722DA8DAAAF8742DF968B869A25BD88F">Akbulut, Hakan</mods:namePart>
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<mods:namePart id="4B65176BC5F8DC1D1C29D59AB6FFE84D">Ürün, Yüksel</mods:namePart>
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<mods:date id="B8EDFE1ADDBF82D3805EB8E227497637">2022</mods:date>
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<heading id="7A0581F93908FFE4FF1CFF37FDA5C7FF" bold="true" box="[171,638,226,250]" centered="true" fontSize="36" level="1" pageId="14" pageNumber="133" reason="1">
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<taxonomicName id="E6F24D163908FFE4FF1CFF37FDA5C7FF" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[171,638,226,250]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="kojii" status="sp. nov." subGenus="Tanakaphaenops">
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<emphasis id="1386EA873908FFE4FF1CFF37FEA8C7FF" bold="true" box="[171,371,226,250]" italics="true" pageId="14" pageNumber="133">Nipponaphaenops</emphasis>
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(
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<emphasis id="1386EA873908FFE4FE35FF37FDE6C7FF" bold="true" box="[386,573,226,250]" italics="true" pageId="14" pageNumber="133">Tanakaphaenops</emphasis>
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)
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<emphasis id="1386EA873908FFE4FDFAFF37FDA5C7FF" bold="true" box="[589,638,226,250]" italics="true" pageId="14" pageNumber="133">kojii</emphasis>
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<paragraph id="214D36953908FFE4FEDCFED3FE65C618" blockId="14.[171,638,226,285]" box="[363,446,262,285]" pageId="14" pageNumber="133">sp. nov.</paragraph>
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<subSubSection id="69E8651E3908FFFBFEADFEE0FE06C67F" lastPageId="17" lastPageNumber="136" pageId="14" pageNumber="133" type="description">
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<paragraph id="214D36953908FFE4FEADFEE0FDD4C64F" blockId="14.[282,527,308,331]" box="[282,527,308,331]" pageId="14" pageNumber="133">
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(
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<figureCitation id="B9C92A103908FFE4FE96FEE0FE58C64F" box="[289,387,309,331]" captionStart="Fig" captionStartId="4.[163,196,919,941]" captionTargetBox="[177,1199,214,885]" captionTargetId="figure-331@4.[176,1200,213,886]" captionTargetPageId="4" captionText="Fig. 1. Right mandible in dorsal view (a–i) and labial (right) and maxillary (left) palpi in ventral view (j–p).—a, Chaetotrechiama procerus S. Uéno; b, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); c & l, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d & m, R. (Uozumitrechus) sp.; e, R. (Pilosotrechiama) mirabilis S. Uéno; f & k, R. (Izushites) sp.; g & j, R. (Rakantrechus) kawasawai S. Uéno; h & o, Nipponaphaenops (N.) erraticus S. Uéno; i & n, N. (Tanakaphaenops) kojii subgen. et sp. nov.; p, Trechiama pluto S. Uéno (this species is unique in having trisetose penultimate labial palpomere in the Trechiama Series)." figureDoi="http://doi.org/10.5281/zenodo.12760048" httpUri="https://zenodo.org/record/12760048/files/figure.png" pageId="14" pageNumber="133">Figs. 1h, n</figureCitation>
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,
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<figureCitation id="B9C92A103908FFE4FE3AFEE0FE60C64E" box="[397,443,309,331]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="14" pageNumber="133">2e, k</figureCitation>
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,
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<figureCitation id="B9C92A103908FFE4FE73FEE0FE29C64E" box="[452,498,309,331]" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="14" pageNumber="133">3c–d</figureCitation>
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,
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<figureCitation id="B9C92A103908FFE4FE4BFEE1FDD3C64F" box="[508,520,308,330]" captionStart="Fig" captionStartId="14.[163,196,1777,1799]" captionTargetBox="[178,1198,1196,1744]" captionTargetId="figure-373@14.[176,1200,1184,1744]" captionTargetPageId="14" captionText="Fig. 6. Habitus.—Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female). Scale: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.12760058" httpUri="https://zenodo.org/record/12760058/files/figure.png" pageId="14" pageNumber="133">6</figureCitation>
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)
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</paragraph>
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<uri id="55633A973908FFE4FF3BFEB1FEE5C69D" pageId="14" pageNumber="133">
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urn:lsid:zoobank.org:act:
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<uuid id="55540C403908FFE4FEC4FEB1FEE5C69D" pageId="14" pageNumber="133">A6A097BE-955F-486E-9832- 17531A260D2A</uuid>
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<paragraph id="214D36953908FFE4FF3BFE67FD57C6E0" blockId="14.[140,652,434,485]" pageId="14" pageNumber="133">
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Semiaphaenopsoid ˂ [sic]:
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<bibRefCitation id="45634B643908FFE4FE3AFE67FDACC6C2" author="Ueno, S. - I. & T. Naito" box="[397,631,434,456]" pageId="14" pageNumber="133" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno & Naitô, 2009: 247</bibRefCitation>
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||
. Semiaphaenopsoid specimen:
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<bibRefCitation id="45634B643908FFE4FE29FE1AFD53C6E0" author="Ueno, S. - I. & T. Naito" box="[414,648,463,485]" pageId="14" pageNumber="133" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno & Naitô, 2009: 252</bibRefCitation>
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||
.
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||
</paragraph>
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<paragraph id="214D36953908FFE4FF10FDC5FF02C41C" blockId="14.[140,670,525,1141]" pageId="14" pageNumber="133">
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<emphasis id="1386EA873908FFE4FF10FDC5FE88C522" box="[167,339,527,551]" italics="true" pageId="14" pageNumber="133">Type material</emphasis>
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||
.
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||
<materialsCitation id="919A3CC83908FFE4FEC3FDDAFE5BC5F2" collectingDate="2009-04-10" collectingDateMax="2009-05-09" collectingDateMin="2009-04-10" collectionCode="NSMT" collectorName="K. Tanaka" country="Japan" county="northwestern Shikoku" elevation="480" location="Furuhashi" municipality="Uchiko-cho" pageId="14" pageNumber="133" specimenCount="1" stateProvince="Ehime" typeStatus="holotype">
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<typeStatus id="FE4988373908FFE4FEC3FDDAFE00C522" box="[372,475,527,551]" pageId="14" pageNumber="133" type="holotype">Holotype</typeStatus>
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||
: ˂ (
|
||
<collectionCode id="47E3AE503908FFE4FDF5FDDAFD4AC522" box="[578,657,527,551]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="14" pageNumber="133">NSMT</collectionCode>
|
||
), lKuroyama,
|
||
<quantity id="E60A9B703908FFE4FEAAFDE7FE86C54C" box="[285,349,562,586]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="133" unit="m" value="480.0">
|
||
<elevation id="AADFD1A63908FFE4FEAAFDE7FE86C54C" box="[285,349,562,586]" metricMagnitude="2" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="133" unit="m" value="480.0">480 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<location id="242D604E3908FFE4FE12FDE7FDC9C54F" LSID="urn:lsid:plazi:treatment:A95B87833908FFFBFF1CFF37FE06C67F:242D604E3908FFE4FE12FDE7FDC9C54F" box="[421,530,562,586]" country="Japan" county="northwestern Shikoku" municipality="Uchiko-cho" name="Furuhashi" pageId="14" pageNumber="133" stateProvince="Ehime">Furuhashi</location>
|
||
,
|
||
<location id="242D604E3908FFE4FD91FDE7FD42C54F" LSID="urn:lsid:plazi:treatment:A95B87833908FFFBFF1CFF37FE06C67F:242D604E3908FFE4FD91FDE7FD42C54F" box="[550,665,562,586]" country="Japan" county="northwestern Shikoku" municipality="Uchiko-cho" name="Ishidatami" pageId="14" pageNumber="133" stateProvince="Ehime">Ishidatami</location>
|
||
,
|
||
<collectingMunicipality id="C129ACEF3908FFE4FF3BFD80FED1C568" box="[140,266,597,621]" pageId="14" pageNumber="133">Uchiko-chô</collectingMunicipality>
|
||
[typed on white label] /
|
||
<collectingRegion id="E336F8773908FFE4FDABFD80FD45C568" box="[540,670,597,621]" country="Japan" name="Ehime" pageId="14" pageNumber="133">Ehime Pref.</collectingRegion>
|
||
<collectingCounty id="C82C4E193908FFE4FF3BFDA2FEA6C58A" box="[140,381,631,655]" pageId="14" pageNumber="133">northwestern Shikoku</collectingCounty>
|
||
, western
|
||
<collectingCountry id="59E576053908FFE4FE5EFDADFDF3C58A" box="[489,552,632,655]" name="Japan" pageId="14" pageNumber="133">Japan</collectingCountry>
|
||
,
|
||
<date id="554C10553908FFE4FD81FDA2FD42C58A" box="[566,665,631,655]" pageId="14" pageNumber="133" value="2009-05-09">
|
||
<collectingDate id="4508E9BD3908FFE4FD81FDA2FD42C58A" box="[566,665,631,655]" pageId="14" pageNumber="133" value="2009-05-09">9.V.2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="8C0753433908FFE4FF3BFD4FFF22C5B7" box="[140,249,666,690]" pageId="14" pageNumber="133">K. Tanaka</collectorName>
|
||
leg. [typed on white label] / found in a
|
||
<collectingMethod id="F8B34E823908FFE4FF3BFD68FEDCC5D1" box="[140,263,701,725]" pageId="14" pageNumber="133">baited trap</collectingMethod>
|
||
set by
|
||
<collectorName id="8C0753433908FFE4FEC4FD68FE30C5D0" box="[371,491,701,725]" pageId="14" pageNumber="133">K. Tanaka</collectorName>
|
||
on
|
||
<date id="554C10553908FFE4FD90FD68FD46C5D0" box="[551,669,701,725]" pageId="14" pageNumber="133" value="2009-04-10">
|
||
<collectingDate id="4508E9BD3908FFE4FD90FD68FD46C5D0" box="[551,669,701,725]" pageId="14" pageNumber="133" value="2009-04-10">10.IV.2009</collectingDate>
|
||
</date>
|
||
[typed on white label]
|
||
</materialsCitation>
|
||
/
|
||
<typeStatus id="FE4988373908FFE4FE2CFD0AFE25C5F2" box="[411,510,735,759]" pageId="14" pageNumber="133" type="holotype">Holotype</typeStatus>
|
||
[typed on red label]z.
|
||
</paragraph>
|
||
<paragraph id="214D36953908FFE4FF10FCF0FE76C45A" blockId="14.[140,670,525,1141]" pageId="14" pageNumber="133">
|
||
<emphasis id="1386EA873908FFE4FF10FCF0FE70C438" box="[167,427,805,829]" italics="true" pageId="14" pageNumber="133">Diagnostic characters.</emphasis>
|
||
See the diagnostic characters of the subgenus.
|
||
</paragraph>
|
||
<paragraph id="214D36953908FFE4FF10FCBEFDE2C370" blockId="14.[140,670,525,1141]" pageId="14" pageNumber="133">
|
||
<emphasis id="1386EA873908FFE4FF10FCBEFEF7C487" box="[167,300,875,898]" italics="true" pageId="14" pageNumber="133">Description.</emphasis>
|
||
Female. Body size small, BL:
|
||
<quantity id="E60A9B703908FFE4FF3BFC59FF30C4A1" box="[140,235,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="5.39" pageId="14" pageNumber="133" unit="mm" value="5.39">5.39 mm</quantity>
|
||
. Colour of dorsum, venter, most of the appendages entirely dark reddish brown, maxilla and labial palpi pale yellowish brown. Surface polished, faintly iridescent. Microsculpture composed of fine transverse lines and strongly transverse meshes on head; very fine anastomosing transverse lines on pronotum and elytra.
|
||
</paragraph>
|
||
<paragraph id="214D36953908FFE4FD6AFF02FB56C45A" blockId="14.[707,1236,215,1141]" pageId="14" pageNumber="133">Head elongate subquadrate, longer than wide, only feebly dilated anteriad, with the widest point located at the level of antennal insertion; apical margin of labrum moderately emarginate; labial suture only partially visible as a superficial transverse lineation; frontovertexal areas sharply convex, subcarinate along mid-line; genae flat, glabrous; neck constriction shallow; eye vestiges represented by small indistinct parallel-sided patches, with transverse wrinkles behind them; frontal furrows complete, rapidly divergent anteriad in anterior 2/11 of their length, widely gradually divergent posteriad in posterior 4/9 of their length, subparallel in between; anterior and posterior supraorbital pores widely distant from each other, on lines convergent posteriad; HW/HL 0.98, AL/EL 1.32, relative length of each antennal segment from I to XI: 1: 0.94: 1.55: 1.76: 1.76: 1.77: 1.55: 1.44: 1.37: 1.20: 1.55.</paragraph>
|
||
<paragraph id="214D36953908FFE5FD6AFCBFFE79C4C2" blockId="14.[707,1236,215,1141]" lastBlockId="15.[140,670,215,1799]" lastPageId="15" lastPageNumber="134" pageId="14" pageNumber="133">Pronotum elongate subcordate, widest at about apical 3/14; apex wider than base, slightly arcuate at middle; base nearly straight; sides somewhat irregularly curved, arcuately convergent apicad from the widest point, partially rectilinearly convergent posteriad from that point to basal 1/10, then slightly divergent to base; lateral borders entire, narrow in basal 7/9, wider in apical 2/9; anterior lateral seta inserted slightly behind the widest point, posterior one distinctly anteriorly shifted, located at 1/10 from base; dorsum convex in apical and median parts, baso-lateral areas before basal transverse impression narrowly longitudinally depressed along lateral borders, the bottom of the depressed areas transversely wrinkled; disc with a foveole on each side behind the widest parts; median-line distinct, deepening before vague apical transverse impression and also in and before basal transverse impression, but becoming shallower towards both ends; apical area rather strongly depressed at middle, rather deeply longitudinally strigose along apical margin; basal transverse impression deep and continuous, parallel to basal margin; basal area with a longitudinal paramedian impression and some deep longitudinal impressions; basal foveae rather small and deep; postangular carinae absent; PW/HW 1.30, PL/HL 1.50, PW/PL 0.86, PW/PA 1.33, PW/PB 1.56, PB/PA 0.85, PSa 22.7, PwL 21.5.</paragraph>
|
||
<caption id="758D661D3908FFE4FF14F924FB6CC003" ID-DOI="http://doi.org/10.5281/zenodo.12760058" ID-Zenodo-Dep="12760058" box="[163,1207,1776,1799]" httpUri="https://zenodo.org/record/12760058/files/figure.png" pageId="14" pageNumber="133" startId="14.[163,196,1777,1799]" targetBox="[178,1198,1196,1744]" targetPageId="14" targetType="figure">
|
||
<paragraph id="214D36953908FFE4FF14F924FB6CC003" blockId="14.[163,1207,1776,1799]" box="[163,1207,1776,1799]" pageId="14" pageNumber="133">
|
||
Fig. 6. Habitus.—
|
||
<taxonomicName id="E6F24D163908FFE4FEE0F924FD0CC003" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[343,727,1777,1798]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="133" phylum="Arthropoda" rank="species" species="kojii" subGenus="Tanakaphaenops">
|
||
<emphasis id="1386EA873908FFE4FEE0F924FE27C003" box="[343,508,1777,1798]" italics="true" pageId="14" pageNumber="133">Nipponaphaenops</emphasis>
|
||
(
|
||
<emphasis id="1386EA873908FFE4FDBEF924FD79C003" box="[521,674,1777,1798]" italics="true" pageId="14" pageNumber="133">Tanakaphaenops</emphasis>
|
||
)
|
||
<emphasis id="1386EA873908FFE4FD18F924FD0CC003" box="[687,727,1777,1798]" italics="true" pageId="14" pageNumber="133">kojii</emphasis>
|
||
</taxonomicName>
|
||
subgen et sp. nov. (holotype female). Scale: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="214D36953909FFE5FF10FC07FB5DC617" blockId="15.[140,670,215,1799]" lastBlockId="15.[707,1237,215,1799]" pageId="15" pageNumber="134">Elytra oval, large, more than twice as wide as pronotum, widest slightly behind middle; prehumeral basal parts strongly constricted basad in nearly straight lines; sides narrowly bordered and arcuate from behind shoulders to apices through slight preapical emargination; apices separately rounded; striae narrowly crenulate and weakly punctulate, 1–5 entirely moderately impressed, 6–7 superficial, partially represented by row of punctures, 8 partially impressed around marginal umbilicate pores; scutellar striole vanished; apical recurrent striole short, well incurved and directed to apical anastomosis of striae 3–5; intervals somewhat convex near suture and in basal parts, but flat in other parts; two discal setiferous pores on interval 4 next to stria 3 before middle; single discal setiferous pore on stria 5 behind middle; parascutellar pores on the base of stria 2; preapical pore on apical anastomosis of striae 2 and 3 before the level of anterior end of recurrent striole; apical triangle elongate, with inner edge away from mid-line anteriad; EW/PW 2.19, EL/PL 2.69, EL/EW 1.43. The positions of the elytral setae are as follows: D1 16.4, D2 34.8, D3 64.6, DP 87.8, U1 15.8, U2 17.5, U3 21.7, U4 29.2, U5 63.2, U6 68.7, U7 81.7, U8 89.1.</paragraph>
|
||
<paragraph id="214D36953909FFE5FD6AFEC8FB4BC54F" blockId="15.[707,1237,215,1799]" pageId="15" pageNumber="134">Legs fairly long and stout; protibia gradually dilated apicad, nearly straight, only slightly bowed near apex; mesotibia about 4/9 as long as elytra; metatibia feebly outcurved, about 5/9 as long as elytra; mesotarsus about 9/14 as long as mesotibia; mesotarsomere 1 longer than the mesotarsomeres 2–4 combined; metatarsus about 4/5 as long as metatibia; metatarsomere 1 much longer than metatarsomeres 2–4 combined.</paragraph>
|
||
<paragraph id="214D36953909FFE5FD6AFD80FCB8C5F2" blockId="15.[707,1237,215,1799]" pageId="15" pageNumber="134">
|
||
Apical segment of the female gonopods unguiform, with a sensory fovea bearing a pair of sensory setae present on ventral face near apex; laterotergite with about ten or more apical fringing setae (
|
||
<figureCitation id="B9C92A103909FFE5FCB0FD0AFC8CC5F2" box="[775,855,735,759]" captionStart="Fig" captionStartId="5.[163,196,1055,1077]" captionTargetBox="[177,1198,214,1020]" captionTargetId="figure-238@5.[176,1199,213,1021]" captionTargetPageId="5" captionText="Fig. 2. Labium (a–e), prosternum (f), and female abdominal sterna (setae on metasternum and metacoxae are omitted).—a & j, Rakantrechus (Uozumitrechus) sp.; b & h, R. (Rakantrechus) kawasawai S. Uéno; c & m, R. (Iyotrechus) hisamatsui (S. Uéno et Naitô) comb. nov.; d, R. (Paratrechiama) sp. (nomurai group from Shikoku); e & k, Nipponaphaenops (Tanakaphaenops) kojii subgen. et sp. nov; f–g, R. (Pilosotrechiama) peninsularis S. Uéno et Naitô; i, R. (Izushites) sp.; l, N. (N.) erraticus S. Uéno; n, Kusumia takahasii S. Uéno." figureDoi="http://doi.org/10.5281/zenodo.12760050" httpUri="https://zenodo.org/record/12760050/files/figure.png" pageId="15" pageNumber="134">Fig. 2k</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="214D36953909FFE5FD6AFCD7FB95C41F" blockId="15.[707,1237,215,1799]" box="[733,1102,770,794]" pageId="15" pageNumber="134">Reproductive tract. Not examined.</paragraph>
|
||
<paragraph id="214D36953909FFE5FD6AFCF0FC49C438" blockId="15.[707,1237,215,1799]" box="[733,914,805,829]" pageId="15" pageNumber="134">Male. Unknown.</paragraph>
|
||
<paragraph id="214D36953909FFFAFD6AFC92FDC7C652" blockId="15.[707,1237,215,1799]" lastBlockId="16.[140,669,214,551]" lastPageId="16" lastPageNumber="135" pageId="15" pageNumber="134">
|
||
<emphasis id="1386EA873909FFE5FD6AFC92FC99C45A" box="[733,834,839,863]" italics="true" pageId="15" pageNumber="134">Remarks.</emphasis>
|
||
As mentioned in the introduction of this paper, the
|
||
<typeStatus id="FE4988373909FFE5FC32FCBFFC39C487" box="[901,994,874,898]" pageId="15" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
specimen of this new species has already been introduced in
|
||
<bibRefCitation id="45634B643909FFE5FBDBFC58FC8EC4C2" author="Ueno, S. - I. & T. Naito" pageId="15" pageNumber="134" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno and Naitô (2009)</bibRefCitation>
|
||
, but it was treated as one of the problematical specimens supposedly related to (or a variant of)
|
||
<taxonomicName id="E6F24D163909FFE5FCC4FC20FC98C32A" authority="S. Ueno et Naito, 2009" authorityName="S. Ueno et Naito" authorityYear="2009" class="Insecta" family="Carabidae" genus="Iyotrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="hisamatsui">
|
||
<emphasis id="1386EA873909FFE5FCC4FC20FB86C308" box="[883,1117,1013,1037]" italics="true" pageId="15" pageNumber="134">Iyotrechus hisamatsui</emphasis>
|
||
S.
|
||
<bibRefCitation id="45634B643909FFE5FB37FC20FC98C32A" author="Ueno, S. - I. & T. Naito" pageId="15" pageNumber="134" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno et Naitô, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="E6F24D163909FFE5FCCDFBC2FB52C32A" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" baseAuthorityName="S. Ueno et Naito" baseAuthorityYear="2009" box="[890,1161,1047,1071]" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="hisamatsui" status="comb. nov.">
|
||
<emphasis id="1386EA873909FFE5FCCDFBC2FB52C32A" box="[890,1161,1047,1071]" italics="true" pageId="15" pageNumber="134">Rakantrechus hisamatsui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3909FFE5FB25FBC2FD2FC357" pageId="15" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
|
||
). A brief description of it (merely as a supposed variant of
|
||
<taxonomicName id="E6F24D163909FFE5FC3BFB88FB5AC370" authorityName="S. Ueno et Naito" authorityYear="2009" box="[908,1153,1117,1141]" class="Insecta" family="Carabidae" genus="Iyotrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="hisamatsui">
|
||
<emphasis id="1386EA873909FFE5FC3BFB88FB5AC370" box="[908,1153,1117,1141]" italics="true" pageId="15" pageNumber="134">Iyotrechus hisamatsui</emphasis>
|
||
</taxonomicName>
|
||
) given therein was, however, not to the point, because important characteristics that differ between this new species and
|
||
<taxonomicName id="E6F24D163909FFE5FCCCFB10FCD1C3FA" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" baseAuthorityName="S. Ueno et Naito" baseAuthorityYear="2009" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="hisamatsui" status="comb. nov." subGenus="Iyotrechus">
|
||
<emphasis id="1386EA873909FFE5FCCCFB10FBD6C3D8" box="[891,1037,1221,1245]" italics="true" pageId="15" pageNumber="134">Rakantrechus</emphasis>
|
||
(
|
||
<emphasis id="1386EA873909FFE5FBA9FB10FB55C3D8" box="[1054,1166,1221,1245]" italics="true" pageId="15" pageNumber="134">Iyotrechus</emphasis>
|
||
)
|
||
<emphasis id="1386EA873909FFE5FB28FB10FCD1C3FA" italics="true" pageId="15" pageNumber="134">hisamatsui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3909FFE5FCA5FB32FC57C3FA" box="[786,908,1255,1279]" pageId="15" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, such as the shape of mentum tooth, presence/absence of pronotal postangular setae, and condition of labial suture were not mentioned. Further, we were completely erroneous in stating that the humeral set of marginal umbilicate pores of elytra is lalmost aggregatedz (
|
||
<bibRefCitation id="45634B643909FFE5FD7DFA62FC6FC2CA" author="Ueno, S. - I. & T. Naito" box="[714,948,1463,1487]" pageId="15" pageNumber="134" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno & Naitô, 2009</bibRefCitation>
|
||
, p. 252). In fact, humeral set of
|
||
<taxonomicName id="E6F24D163909FFE5FCBDFA0FFBE8C2F7" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[778,1075,1498,1522]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="kojii" status="sp. nov." subGenus="Tanakaphaenops">
|
||
<emphasis id="1386EA873909FFE5FCBDFA0FFC17C2F7" box="[778,972,1498,1522]" italics="true" pageId="15" pageNumber="134">Nipponaphaenops</emphasis>
|
||
(
|
||
<emphasis id="1386EA873909FFE5FC69FA0EFC29C2F7" box="[990,1010,1499,1522]" italics="true" pageId="15" pageNumber="134">T.</emphasis>
|
||
)
|
||
<emphasis id="1386EA873909FFE5FBB3FA0FFBE8C2F7" box="[1028,1075,1498,1522]" italics="true" pageId="15" pageNumber="134">kojii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3909FFE5FB8AFA0FFB49C2F4" box="[1085,1170,1498,1521]" pageId="15" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is far from being aggregated as described above (
|
||
<figureCitation id="B9C92A103909FFE5FB23FA28FD08C132" captionStart="Fig" captionStartId="6.[163,196,1164,1186]" captionTargetBox="[178,1198,215,1129]" captionTargetId="figure-144@6.[176,1200,213,1131]" captionTargetPageId="6" captionText="Fig. 3. Basal parts of pronotum and elytra in the Trechiama Series.—a–d, f & h–i, basal parts of pronotum and elytra; e & g, basal parts of elytra (a, c & e–i, dorsal view; b & d; oblique dorsal view).—a,Ishikawatrechus septemtrionalis S. Uéno; b, Trechiama ohruii S. Uéno; c–d, Nipponaphaenops (Tanakaphaenops) kojii subgen et sp. nov. (holotype female); e, Rakantrechus (Paratrechiama) sp. (nomurai group from Shikoku); f, Nipponaphaenops (s. str.) erraticus S. Uéno; g, Yamautidius (Miyamaidius) anaulax S. Uéno; h, Rakantrechus (s. str.) kawasawai S. Uéno; i, Himiseus sp.—D1, first elytral discal seta on or near stria 3; D2, second elytral discal seta on or near stria 3; D5, first elytral discal seta on or near stria 5; ds, discal setae on pronotum; fur, basal transverse furrow; hb, prehumeral border of elytra; icl, inclined face of elytral basal peduncle; tvc, transverse carina bordering elytral disc and inclined face; ©–®, pores 1–4 of marginal umbilicate series of elytra, respectively; III, elytral stria 3; V, elytral stria 5. Scale: 0.25mm for a, d & h; 0.5mm for b; 0.21mm for c & g; 0.32mm for e, f & i." figureDoi="http://doi.org/10.5281/zenodo.12760052" httpUri="https://zenodo.org/record/12760052/files/figure.png" pageId="15" pageNumber="134">Fig. 3 d</figureCitation>
|
||
). Having compared the
|
||
<typeStatus id="FE4988373909FFE5FC6EF9CAFBEDC132" box="[985,1078,1567,1591]" pageId="15" pageNumber="134" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E6F24D163909FFE5FBEAF9F5FB76C132" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[1117,1197,1567,1591]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="kojii" status="sp. nov.">
|
||
<emphasis id="1386EA873909FFE5FBEAF9F5FBB4C132" box="[1117,1135,1568,1591]" italics="true" pageId="15" pageNumber="134">N</emphasis>
|
||
.
|
||
<emphasis id="1386EA873909FFE5FBC9F9CAFB76C132" box="[1150,1197,1567,1591]" italics="true" pageId="15" pageNumber="134">kojii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3909FFE5FB02F9F5FD2BC15C" pageId="15" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
|
||
with the type series of
|
||
<taxonomicName id="E6F24D163909FFE5FBB5F997FCD1C179" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" baseAuthorityName="S. Ueno et Naito" baseAuthorityYear="2009" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="134" phylum="Arthropoda" rank="species" species="hisamatsui" status="comb. nov.">
|
||
<emphasis id="1386EA873909FFE5FBB5F997FCD1C179" italics="true" pageId="15" pageNumber="134">Rakantrechus hisamatsui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3909FFE5FCA4F9B0FC50C179" box="[787,907,1637,1661]" pageId="15" pageNumber="134" rank="species">comb. nov.</taxonomicNameLabel>
|
||
in that paper may seem to be widely misled or irrelevant to the subject. However, the late Dr. Uéno had intended to give a taxonomic consideration over a broader scope; he wrote (pers. comm.) in a letter to me dated 31 March, 2009 lI have an expectation that a certain affinity, like that between
|
||
<taxonomicName id="E6F24D163916FFFAFE1CFF2FFDB6C617" authorityName="S. Ueno" authorityYear="1971" box="[427,621,250,274]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFE1CFF2FFDB6C617" box="[427,621,250,274]" italics="true" pageId="16" pageNumber="135">Nipponaphaenops</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="1386EA873916FFFAFF3BFEC8FEF4C630" box="[140,303,285,309]" italics="true" pageId="16" pageNumber="135">Ishikawatrechu</emphasis>
|
||
s, will be shown also between the former and
|
||
<taxonomicName id="E6F24D163916FFFAFEBFFEEAFEA3C652" authorityName="S. Ueno et Naito" authorityYear="2009" box="[264,376,319,343]" class="Insecta" family="Carabidae" genus="Iyotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFEBFFEEAFEA3C652" box="[264,376,319,343]" italics="true" pageId="16" pageNumber="135">Iyotrechus</emphasis>
|
||
</taxonomicName>
|
||
z (in Japanese).
|
||
</paragraph>
|
||
<paragraph id="214D36953916FFFAFF10FEB6FB9CC617" blockId="16.[140,669,214,551]" lastBlockId="16.[707,1236,215,551]" pageId="16" pageNumber="135">
|
||
<emphasis id="1386EA873916FFFAFF10FEB6FE5CC67F" box="[167,391,355,378]" italics="true" pageId="16" pageNumber="135">Notes on synonymy.</emphasis>
|
||
Two spellings cited in the synonymy of this new species are not available. These spellings were used to refer to the
|
||
<typeStatus id="FE4988373916FFFAFF3BFE1FFF32C6E7" box="[140,233,458,482]" pageId="16" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
specimen (designated in this paper) of this new species in
|
||
<bibRefCitation id="45634B643916FFFAFEDEFE38FDB6C500" author="Ueno, S. - I. & T. Naito" box="[361,621,493,517]" pageId="16" pageNumber="135" pagination="245 - 253" refId="ref13685" refString="Ueno, S. - I. and T. Naito 2009. A remarkable blind trechine beetle (Coleoptera, Trechinae) from northwestern Shikoku, southwest Japan. Japanese Journal of Systematic Entomology 15: 245 - 253." type="journal article" year="2009">Uéno and Naitô (2009)</bibRefCitation>
|
||
, but were not proposed as scientific names for a new species, as the taxonomic position of the specimen was not determined at the time.
|
||
</paragraph>
|
||
<paragraph id="214D36953916FFFBFD6AFEC8FEEEC630" blockId="16.[707,1236,215,551]" lastBlockId="17.[140,669,215,378]" lastPageId="17" lastPageNumber="136" pageId="16" pageNumber="135">
|
||
<emphasis id="1386EA873916FFFAFD6AFEC8FC02C630" box="[733,985,285,309]" italics="true" pageId="16" pageNumber="135">Habitat and bionomics</emphasis>
|
||
. The type locality of this remarkable new species is on the right (eastern) side of the Hiji-kawa River, about
|
||
<quantity id="E60A9B703916FFFAFB3AFEB7FB0FC67F" box="[1165,1236,354,378]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="16" pageNumber="135" unit="km" value="0.7">0.7 km</quantity>
|
||
north of the type locality of
|
||
<taxonomicName id="E6F24D163916FFFAFBB4FE50FCD1C6BA" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" baseAuthorityName="S. Ueno et Naito" baseAuthorityYear="2009" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="species" species="hisamatsui" status="comb. nov.">
|
||
<emphasis id="1386EA873916FFFAFBB4FE50FCD1C6BA" italics="true" pageId="16" pageNumber="135">Rakantrechus hisamatsui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="08B557FC3916FFFAFCA6FE72FC5CC6BA" box="[785,903,423,447]" pageId="16" pageNumber="135" rank="species">comb. nov.</taxonomicNameLabel>
|
||
From nearby places, the members of
|
||
<taxonomicName id="E6F24D163916FFFAFCA3FE1FFC7DC6E7" authorityName="S. Ueno" authorityYear="1951" box="[788,934,458,482]" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFCA3FE1FFC7DC6E7" box="[788,934,458,482]" italics="true" pageId="16" pageNumber="135">Rakantrechus</emphasis>
|
||
</taxonomicName>
|
||
of the subgenera
|
||
<taxonomicName id="E6F24D163916FFFAFBD3FE1FFB0FC6E7" authorityName="S. Ueno et Naito" authorityYear="2009" box="[1124,1236,458,482]" class="Insecta" family="Carabidae" genus="Iyotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFBD3FE1FFB0FC6E7" box="[1124,1236,458,482]" italics="true" pageId="16" pageNumber="135">Iyotrechus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E6F24D163916FFFAFD43FE38FC6CC500" authority="S. Ueno" authorityName="S. Ueno" box="[756,951,493,517]" class="Insecta" family="Carabidae" genus="Izushites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFD43FE38FC8AC500" box="[756,849,493,517]" italics="true" pageId="16" pageNumber="135">Izushites</emphasis>
|
||
S. Uéno
|
||
</taxonomicName>
|
||
are also discovered (
|
||
<figureCitation id="B9C92A103916FFFAFB1EFE38FD08C522" captionStart="Fig" captionStartId="16.[163,196,1601,1623]" captionTargetBox="[157,1217,605,1567]" captionTargetId="figure-172@16.[157,1218,605,1568]" captionTargetPageId="16" captionText="Fig. 7. Map showing the distribution of the Rakantrechus Complex (in part) and anophthalmic species of the genus Trechiama in western Japan.—Light green square, Rakantrechus (s. str.); orange square, R. (Izushites); green circle, R. (Iyotrechus); yellow circle, R. (Pilosotrechiama); green triangle, Nipponaphaenops (s. str.); blue open triangle, N. (Tanakaphaenops) subgen. nov.; purple obtriangle, Ishikawatrechus; large purple circle, anophthalmic Trechiama species.—An, the type locality of Ishikawatrechus annulus Sugaya et Hara; Tk, the type locality of Nipponaphaenops (Tanakaphaenops) kojii sugen. et sp. nov. The course of the Hôyo River in the Last Glacial Maximum was drawn according to Kuwashiro (1959)." figureDoi="http://doi.org/10.5281/zenodo.12760060" httpUri="https://zenodo.org/record/12760060/files/figure.png" pageId="16" pageNumber="135">Fig. 7</figureCitation>
|
||
). The
|
||
<typeStatus id="FE4988373916FFFAFC94FDDAFC5BC522" box="[803,896,527,551]" pageId="16" pageNumber="135" type="holotype">holotype</typeStatus>
|
||
specimen was obtained by a baited pit-fall trap set in upper-hypogean environment (cf.
|
||
<bibRefCitation id="45634B643917FFFBFE87FF2FFD4CC617" author="Juberthie, C. & M. Bouillon" box="[304,663,250,274]" pageId="17" pageNumber="136" pagination="91 - 98" refId="ref12914" refString="Juberthie, C. and M. Bouillon 1983. Presence des Aphaenops (Coleopteres, Trechinae) dans le Milieu Souterrain Superficiel des Pyrenees francaises. Memoires de biospeologie 10: 91 - 98." type="journal article" year="1983">Juberthie and Bouillon, 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="45634B643917FFFBFF3BFEC8FEF2C630" author="Yoshida, M." box="[140,297,285,309]" pageId="17" pageNumber="136" pagination="1 - 6" refId="ref13726" refString="Yoshida, M. 2012. How to collect insects living in upper hypogean by using underground traps. The Nature and Insects 47: 6 - 12, pl. 1, figs. 1 - 6 (in Japanese, with English title)." type="journal article" year="2012">Yoshida, 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="758D661D3916FFFAFF14F994FC88C002" ID-DOI="http://doi.org/10.5281/zenodo.12760060" ID-Zenodo-Dep="12760060" httpUri="https://zenodo.org/record/12760060/files/figure.png" pageId="16" pageNumber="135" startId="16.[163,196,1601,1623]" targetBox="[157,1217,605,1567]" targetPageId="16" targetType="figure">
|
||
<paragraph id="214D36953916FFFAFF14F994FC88C002" blockId="16.[163,1214,1601,1799]" pageId="16" pageNumber="135">
|
||
Fig. 7. Map showing the distribution of the
|
||
<taxonomicName id="E6F24D163916FFFAFDE1F994FD09C153" authorityName="S. Ueno" authorityYear="1951" box="[598,722,1601,1622]" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFDE1F994FD09C153" box="[598,722,1601,1622]" italics="true" pageId="16" pageNumber="135">Rakantrechus</emphasis>
|
||
</taxonomicName>
|
||
Complex (in part) and anophthalmic species of the genus
|
||
<taxonomicName id="E6F24D163916FFFAFEBBF98BFEB7C176" box="[268,364,1630,1651]" class="Insecta" family="Carabidae" genus="Trechiama" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFEBBF98BFEB7C176" box="[268,364,1630,1651]" italics="true" pageId="16" pageNumber="135">Trechiama</emphasis>
|
||
</taxonomicName>
|
||
in western Japan.—Light green square,
|
||
<taxonomicName id="E6F24D163916FFFAFD55F98BFC85C176" authorityName="S. Ueno" authorityYear="1951" box="[738,862,1630,1651]" class="Insecta" family="Carabidae" genus="Rakantrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFD55F98BFC85C176" box="[738,862,1630,1651]" italics="true" pageId="16" pageNumber="135">Rakantrechus</emphasis>
|
||
</taxonomicName>
|
||
(s. str.); orange square,
|
||
<emphasis id="1386EA873916FFFAFB89F98AFB97C171" box="[1086,1100,1631,1652]" italics="true" pageId="16" pageNumber="135">R</emphasis>
|
||
. (
|
||
<taxonomicName id="E6F24D163916FFFAFBD7F98BFB74C176" box="[1120,1199,1630,1651]" class="Insecta" family="Carabidae" genus="Izushites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFBD7F98BFB74C176" box="[1120,1199,1630,1651]" italics="true" pageId="16" pageNumber="135">Izushites</emphasis>
|
||
</taxonomicName>
|
||
); green circle,
|
||
<emphasis id="1386EA873916FFFAFEFCF9A9FE82C194" box="[331,345,1660,1681]" italics="true" pageId="16" pageNumber="135">R</emphasis>
|
||
. (
|
||
<taxonomicName id="E6F24D163916FFFAFED8F9A9FE15C194" authorityName="S. Ueno et Naito" authorityYear="2009" box="[367,462,1660,1681]" class="Insecta" family="Carabidae" genus="Iyotrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFED8F9A9FE15C194" box="[367,462,1660,1681]" italics="true" pageId="16" pageNumber="135">Iyotrechus</emphasis>
|
||
</taxonomicName>
|
||
); yellow circle,
|
||
<emphasis id="1386EA873916FFFAFDDCF9A9FDA2C194" box="[619,633,1660,1681]" italics="true" pageId="16" pageNumber="135">R</emphasis>
|
||
. (
|
||
<taxonomicName id="E6F24D163916FFFAFD39F9A9FCF8C194" authorityName="S. Ueno" authorityYear="1958" box="[654,803,1660,1681]" class="Insecta" family="Carabidae" genus="Pilosotrechiama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFD39F9A9FCF8C194" box="[654,803,1660,1681]" italics="true" pageId="16" pageNumber="135">Pilosotrechiama</emphasis>
|
||
</taxonomicName>
|
||
); green triangle,
|
||
<taxonomicName id="E6F24D163916FFFAFC7FF9A9FBB6C194" authorityName="S. Ueno" authorityYear="1971" box="[968,1133,1660,1681]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFC7FF9A9FBB6C194" box="[968,1133,1660,1681]" italics="true" pageId="16" pageNumber="135">Nipponaphaenops</emphasis>
|
||
</taxonomicName>
|
||
(s. str.); blue open triangle,
|
||
<emphasis id="1386EA873916FFFAFE36F94FFE4CC1AA" box="[385,407,1690,1711]" italics="true" pageId="16" pageNumber="135">N.</emphasis>
|
||
(
|
||
<taxonomicName id="E6F24D163916FFFAFE13F94CFDE6C1AB" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[420,573,1689,1710]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="131" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Tanakaphaenops">
|
||
<emphasis id="1386EA873916FFFAFE13F94CFDE6C1AB" box="[420,573,1689,1710]" italics="true" pageId="16" pageNumber="135">Tanakaphaenops</emphasis>
|
||
</taxonomicName>
|
||
) subgen. nov.; purple obtriangle,
|
||
<taxonomicName id="E6F24D163916FFFAFCC6F94CFBDEC1AB" authorityName="Habu" authorityYear="1950" box="[881,1029,1689,1710]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1386EA873916FFFAFCC6F94CFBDEC1AB" box="[881,1029,1689,1710]" italics="true" pageId="16" pageNumber="135">Ishikawatrechus</emphasis>
|
||
</taxonomicName>
|
||
; large purple circle, anophthalmic
|
||
<taxonomicName id="E6F24D163916FFFAFEE5F963FDD9C1C9" box="[338,514,1718,1740]" class="Insecta" family="Carabidae" genus="Trechiama" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="1386EA873916FFFAFEE5F963FE69C1CE" box="[338,434,1718,1739]" italics="true" pageId="16" pageNumber="135">Trechiama</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
—An, the type locality of
|
||
<taxonomicName id="E6F24D163916FFFAFD40F963FBAAC1C9" authority="Sugaya et Hara" authorityName="Sugaya et Hara" box="[759,1137,1718,1740]" class="Insecta" family="Carabidae" genus="Ishikawatrechus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="species" species="annulus">
|
||
<emphasis id="1386EA873916FFFAFD40F963FC01C1CE" box="[759,986,1718,1739]" italics="true" pageId="16" pageNumber="135">Ishikawatrechus annulus</emphasis>
|
||
Sugaya et Hara
|
||
</taxonomicName>
|
||
; Tk, the type locality of
|
||
<taxonomicName id="E6F24D163916FFFAFED5F901FD3FC1EC" authorityName="Karaoğlan & Yekedüz & Yazgan & Mocan & Köksoy & Yaşar & Şenler & Utkan & Demirkazık & Akbulut & Ürün" authorityYear="2022" box="[354,740,1748,1769]" class="Insecta" family="Carabidae" genus="Nipponaphaenops" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="135" phylum="Arthropoda" rank="species" species="kojii" subGenus="Tanakaphaenops">
|
||
<emphasis id="1386EA873916FFFAFED5F901FDDCC1EC" box="[354,519,1748,1769]" italics="true" pageId="16" pageNumber="135">Nipponaphaenops</emphasis>
|
||
(
|
||
<emphasis id="1386EA873916FFFAFDA2F901FD75C1EC" box="[533,686,1748,1769]" italics="true" pageId="16" pageNumber="135">Tanakaphaenops</emphasis>
|
||
)
|
||
<emphasis id="1386EA873916FFFAFD0BF901FD3FC1EC" box="[700,740,1748,1769]" italics="true" pageId="16" pageNumber="135">kojii</emphasis>
|
||
</taxonomicName>
|
||
sugen. et sp. nov. The course of the Hôyo River in the Last Glacial Maximum was drawn according to
|
||
<bibRefCitation id="45634B643916FFFAFD10F924FC94C002" author="Kuwashiro, I." box="[679,847,1777,1799]" pageId="16" pageNumber="135" pagination="24 - 35" refId="ref12950" refString="Kuwashiro, I. 1959. Submarine topography of Japanese Inland Sea Setonaikai. Geographical Review of Japan 32: 24 - 35 (in Japanese, with English title and summary)." type="journal article" year="1959">Kuwashiro (1959)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="214D36953917FFFBFF10FEEAFE06C67F" blockId="17.[140,669,215,378]" pageId="17" pageNumber="136">
|
||
<emphasis id="1386EA873917FFFBFF10FEEAFEC2C652" box="[167,281,319,343]" italics="true" pageId="17" pageNumber="136">Etymology</emphasis>
|
||
. This new species is named after its discoverer, Mr. Kôji Tanaka.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |