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<paragraph id="1A09685AFFDAFFA1FF67F936FD01F942" blockId="9.[197,715,1677,1738]" box="[197,715,1677,1703]" pageId="9" pageNumber="8">
<heading id="4141DF36FFDAFFA1FF67F936FD01F942" allCaps="true" box="[197,715,1677,1703]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="8" reason="2">
<taxonomicName id="DDB613D9FFDAFFA1FF67F936FD7FF942" ID-CoL="C4NHK" authority="GROVES, 1994" authorityName="Groves" authorityYear="1994" box="[197,693,1677,1703]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="cowlitziana">
<emphasis id="28C2B448FFDAFFA1FF67F936FDC0F942" box="[197,522,1677,1703]" italics="true" pageId="9" pageNumber="8">NUCLEOLARIA COWLITZIANA</emphasis>
GROVES, 1994
</taxonomicName>
A
</heading>
</paragraph>
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<figureCitation id="828D74DFFFDAFFA1FE25F90BFDC3F92F" box="[391,521,1712,1738]" captionStart="Figure 4" captionStartId="9.[112,199,195,221]" captionTargetId="figure-13@8.[123,1510,192,1968]" captionTargetPageId="8" captionText="Figure 4 (previous page). AC. Nucleolaria cowlitzensis Groves, 1994, Cowlitz Formation (middle Eocene, upper “Tejon Stage”), Vader area, south-central Lewis County, Washington. Basal (A), dorsal (B), and left (columellar) side (C) views, 1.9x, length 27.2 mm, width 17.3 mm, height 11.3 mm, of holotype UCMP 39837, UCMP Locality D-8040. DH. Propustularia kemperae (Nelson, 1925), Santa Susana Formation, (Paleocene, Selandian Stage), north side Simi Hills, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F), and views, 2.2x, length 21 mm, width 17.3, height 12 mm, of holotype CASG 391.01 [ex CASG 987], Locality CASG 391. Dorsal (G) view, 2.6x, length 18.2 mm, width 14.9, height 11.2 mm, of hypotype LACMIP 22307.6, LACMIP Type 14918 (LACMIP Locality 22307). Basal (H) view, 2.2x, length 21 mm, width 17.3, height 12 mm, of paratype UCMP 30541, UCMP Locality 3764. IN. Subepona goedertorum (Groves and Squires, 1995), Crescent Forma- tion (Eocene, “Capay Stage”), Black Hills, Thurston County, Washington. Basal (I), dorsal (J), left (columellar) lateral (K), and right (labral) (L) views, 2.8x, length 19.4 mm, width 12.3, height 10.8 mm, of holotype LACMIP 16655.23, LACMIP Type 12375, LACMIP Locality 16655 [=LACMIP Locality 41563; ex CSUN Locality 1563]. MN, Juncal Formation, Canton Canyon, Whitaker Peak area, Ventura County, southern California. Basal (M) and dorsal (N) views, both 2.4x, length 19.4 mm, width 13.9, height 9.4 mm, of hypotype LACMIP 40807.1, LACMIP Type 14935 (LACMIP Locality 40807 [= LACMIP Locality 16191, ex CSUN Locality 807]). OQ, Subepona leahae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (O), dorsal (P), and left (labral) side (Q) views, 1.6x, length 29.5 mm, width 13.9, height 15.1 mm, of holotype LACMIP 40374.79, LACMIP Type 14936, (LACMIP Locality 40374)." figureDoi="http://doi.org/10.5281/zenodo.13869592" httpUri="https://zenodo.org/record/13869592/files/figure.png" pageId="9" pageNumber="8">FIGS. 4AC</figureCitation>
</paragraph>
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<subSubSection id="52AC3BD1FFDAFFA1FF3BF949FD25F87F" pageId="9" pageNumber="8" type="reference_group">
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<treatmentCitation id="9B174E4BFFDAFFA1FF3BF949FD10F8E8" author="Groves, L. T." box="[153,730,1778,1806]" page="247" pageId="9" pageNumber="8" year="1994">
<taxonomicName id="DDB613D9FFDAFFA1FF3BF949FD10F8E8" ID-CoL="C4NHK" authority="Groves, 1994 a, p. 247" authorityName="Groves" authorityPageNumber="247" authorityYear="1994" box="[153,730,1778,1806]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="cowlitziana">
<emphasis id="28C2B448FFDAFFA1FF3BF949FE75F8EB" box="[153,447,1778,1806]" italics="true" pageId="9" pageNumber="8">Nucleolaria cowlitziana</emphasis>
<bibRefCitation id="7E2715ABFFDAFFA1FE65F948FDB1F8E8" author="Groves, L. T." box="[455,635,1779,1805]" pageId="9" pageNumber="8" pagination="79 - 87" refId="ref37896" refString="Groves, L. T. 1994 a. New species of Cypraeidae (Mollusca: Gastropoda) from the Miocene of California and the Eocene of Washington. The Veliger 37 (3): 244 - 252, figs. 1 - 13 [reprinted in Bulletin of the Southern California Paleontological Society 26 (9 - 10): 79 - 87]." type="journal article" year="1994">Groves, 1994a</bibRefCitation>
, p. 247
</taxonomicName>
</treatmentCitation>
248, figs. 6, 7.
<bibRefCitation id="7E2715ABFFDAFFA1FF7BF8ADFEB6F8D4" author="Nesbitt, E. A." box="[217,380,1814,1841]" pageId="9" pageNumber="8" pagination="1060 - 1073" refId="ref40036" refString="Nesbitt, E. A. 1995. Paleoecological analysis of molluscan assemblages from the middle Eocene Cowlitz Forma- tion, southwestern Washington. Journal of Paleontology 69 (6): 1060 - 1073, figs. 1 - 6." type="journal article" year="1995">Nesbitt, 1995</bibRefCitation>
. p. 1066.
<bibRefCitation id="7E2715ABFFDAFFA1FE48F8ACFD42F8D4" author="Groves, L. T." box="[490,648,1815,1841]" pageId="9" pageNumber="8" pagination="7 - 10" refId="ref38049" refString="Groves, L. T. 1997. Fossil and recent species of eastern Pacific Cypraeacea (Cypraeidae and Eocypraeinae [Ovulidae]): An update. The Western Society of Malacology Annual Report 29: 7 - 10." type="book chapter" year="1997">Groves, 1997</bibRefCitation>
. p. 7.
<bibRefCitation id="7E2715ABFFDAFFA1FD64F8ACFF7BF8B1" author="Groves, L. T." pageId="9" pageNumber="8" pagination="116 - 124" refId="ref38090" refString="Groves, L. T. 2000. Catalog of Recent and fossil Cypraeidae and Eocypraeinae (Ovulidae): 1994 through 1999. The Festivus 32 (8): 116 - 124." type="journal article" year="2000">Groves, 2000</bibRefCitation>
. p. 120.
<bibRefCitation id="7E2715ABFFDAFFA1FE81F881FDABF8B1" author="Dolin, L. &amp; P. Lozouet" box="[291,609,1850,1876]" pageId="9" pageNumber="8" pagination="1 - 164" refId="ref36614" refString="Dolin, L. and P. Lozouet. 2004. Nouvelles especes de gasteropodes (Mollusca: Gastropoda) de l'Oligocene et du Miocene inferieur de l'Aquitaine (Sud-Ouest de la France). Partie 3. Cypraeidae et Ovulidae. Cossmanniana, hors serie, 4: 1 - 164, figs. 1 - 27, pls. 1 - 36." type="journal article" year="2004">Dolin and Lozouet, 2004</bibRefCitation>
. p. 73.
<bibRefCitation id="7E2715ABFFDAFFA1FD60F881FF7BF892" author="Lorenz, F." pageId="9" pageNumber="8" refId="ref39506" refString="Lorenz, F. 2017. Cowries. A guide to the gastropod family Cypraeidae. Volume 1. Biology and Systematics. Conch- Books, Harxheim, Germany. 644 pp., numerous figures." type="book" year="2017">Lorenz, 2017</bibRefCitation>
. p. 220.
<bibRefCitation id="7E2715ABFFDAFFA1FEB5F8E6FE76F892" author="Lorenz, F." box="[279,444,1885,1911]" pageId="9" pageNumber="8" refId="ref39543" refString="Lorenz, F. 2018. Cowries. A guide to the gastropod fam- ily Cypraeidae. Volume 2. Shells and Animals. Conch- Books, Harxheim, Germany. 715 pp., 345 pls." type="book" year="2018">Lorenz, 2018</bibRefCitation>
. p. 664, pl. 328, fig. 5.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1A09685AFFDAFFA1FF32F8C4FD25F87F" blockId="9.[112,800,1778,1947]" box="[144,751,1919,1947]" pageId="9" pageNumber="8">
<taxonomicName id="DDB613D9FFDAFFA1FF32F8C4FDCAF87F" authority="(Groves)" baseAuthorityName="Groves" box="[144,512,1919,1947]" class="Gastropoda" family="Pediculariidae" genus="Jenneria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="cowlitziana">
<emphasis id="28C2B448FFDAFFA1FF32F8C4FE41F87E" box="[144,395,1919,1947]" italics="true" pageId="9" pageNumber="8">Jenneria cowlitziana</emphasis>
(Groves)
</taxonomicName>
.
<bibRefCitation id="7E2715ABFFDAFFA1FDA8F83BFD6AF87F" author="Fehse, D." box="[522,672,1920,1946]" pageId="9" pageNumber="8" pagination="1 - 51" refId="ref37011" refString="Fehse, D. 2001. Beitrage zur Kenntnis der Ovuliade (Mollusca: Cypraeoidea). 8. Einleitung zur familie sowie Katalog, Taxonomy und Bibliographie und Bemerkungen zu verwandten Gruppen. Acta Conchyliorum 5: 1 - 51, figs. 1 - 3." type="journal article" year="2001">Fehse, 2001</bibRefCitation>
. p. 37.
</paragraph>
</subSubSection>
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<paragraph id="1A09685AFFDAFFA1FCC2FCC9FB45FC13" blockId="9.[832,1521,882,1965]" pageId="9" pageNumber="8">
<materialsCitation id="AADE6207FFDAFFA1FCC2FCC9FB45FC13" collectionCode="UCMP" county="Lewis County" location="Cowlitz Formation" municipality="Locality" pageId="9" pageNumber="8" specimenCode="UCMP 39837" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="28C2B448FFDAFFA1FCC2FCC9FB16FC69" bold="true" box="[864,1244,882,908]" pageId="9" pageNumber="8">
<typeStatus id="C50DD6F8FFDAFFA1FCC2FCC9FC16FC69" box="[864,988,882,908]" pageId="9" pageNumber="8" type="holotype">Holotype</typeStatus>
and Type Locality
</emphasis>
<specimenCode id="4A10C021FFDAFFA1FB5AFCC9FA68FC69" box="[1272,1442,882,908]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP 39837</specimenCode>
(
<figureCitation id="828D74DFFFDAFFA1FA16FCC9FC4CFC55" captionStart="Figure 4" captionStartId="9.[112,199,195,221]" captionTargetId="figure-13@8.[123,1510,192,1968]" captionTargetPageId="8" captionText="Figure 4 (previous page). AC. Nucleolaria cowlitzensis Groves, 1994, Cowlitz Formation (middle Eocene, upper “Tejon Stage”), Vader area, south-central Lewis County, Washington. Basal (A), dorsal (B), and left (columellar) side (C) views, 1.9x, length 27.2 mm, width 17.3 mm, height 11.3 mm, of holotype UCMP 39837, UCMP Locality D-8040. DH. Propustularia kemperae (Nelson, 1925), Santa Susana Formation, (Paleocene, Selandian Stage), north side Simi Hills, Ventura County, southern California. Basal (D), dorsal (E), and right (labral) side (F), and views, 2.2x, length 21 mm, width 17.3, height 12 mm, of holotype CASG 391.01 [ex CASG 987], Locality CASG 391. Dorsal (G) view, 2.6x, length 18.2 mm, width 14.9, height 11.2 mm, of hypotype LACMIP 22307.6, LACMIP Type 14918 (LACMIP Locality 22307). Basal (H) view, 2.2x, length 21 mm, width 17.3, height 12 mm, of paratype UCMP 30541, UCMP Locality 3764. IN. Subepona goedertorum (Groves and Squires, 1995), Crescent Forma- tion (Eocene, “Capay Stage”), Black Hills, Thurston County, Washington. Basal (I), dorsal (J), left (columellar) lateral (K), and right (labral) (L) views, 2.8x, length 19.4 mm, width 12.3, height 10.8 mm, of holotype LACMIP 16655.23, LACMIP Type 12375, LACMIP Locality 16655 [=LACMIP Locality 41563; ex CSUN Locality 1563]. MN, Juncal Formation, Canton Canyon, Whitaker Peak area, Ventura County, southern California. Basal (M) and dorsal (N) views, both 2.4x, length 19.4 mm, width 13.9, height 9.4 mm, of hypotype LACMIP 40807.1, LACMIP Type 14935 (LACMIP Locality 40807 [= LACMIP Locality 16191, ex CSUN Locality 807]). OQ, Subepona leahae n. sp., Llajas Formation (Eocene, “Domengine Stage”), north side Simi Valley, Ventura County, southern California. Basal (O), dorsal (P), and left (labral) side (Q) views, 1.6x, length 29.5 mm, width 13.9, height 15.1 mm, of holotype LACMIP 40374.79, LACMIP Type 14936, (LACMIP Locality 40374)." figureDoi="http://doi.org/10.5281/zenodo.13869592" httpUri="https://zenodo.org/record/13869592/files/figure.png" pageId="9" pageNumber="8">Figs. 4AC</figureCitation>
), length
<quantity id="DD4EC5BFFFDAFFA1FC4EFC2DFB90FC55" box="[1004,1114,918,944]" metricMagnitude="-2" metricUnit="m" metricValue="2.7199999999999998" pageId="9" pageNumber="8" unit="mm" value="27.2">27.2 mm</quantity>
, width
<quantity id="DD4EC5BFFFDAFFA1FB10FC2DFAEAFC55" box="[1202,1312,918,944]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="9" pageNumber="8" unit="mm" value="17.3">17.3 mm</quantity>
, height
<quantity id="DD4EC5BFFFDAFFA1FADCFC2DFA26FC55" box="[1406,1516,918,944]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="9" pageNumber="8" unit="mm" value="11.3">11.3 mm</quantity>
.
<collectionCode id="7CA7F09FFFDAFFA1FCE2FC02FC41FC36" box="[832,907,953,979]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="9" pageNumber="8" type="University or college">UCMP</collectionCode>
<collectingMunicipality id="FA6DF220FFDAFFA1FC32FC02FC38FC36" box="[912,1010,953,979]" pageId="9" pageNumber="8">Locality</collectingMunicipality>
D-8040,
<location id="1F693E81FFDAFFA1FBFCFC02FA88FC36" LSID="urn:lsid:plazi:treatment:921FD94CFFDAFFA2FF67F936FCEBFCD7:1F693E81FFDAFFA1FBFCFC02FA88FC36" box="[1118,1346,953,979]" county="Lewis County" municipality="Locality" name="Cowlitz Formation" pageId="9" pageNumber="8" stateProvince="Washington">Cowlitz Formation</location>
, south-central
<collectingCounty id="F36810D6FFDAFFA1FCE2FC67FC23FC13" box="[832,1001,988,1014]" pageId="9" pageNumber="8">Lewis County</collectingCounty>
,
<collectingRegion id="D872A6B8FFDAFFA1FC51FC67FB41FC13" box="[1011,1163,988,1014]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
.
</materialsCitation>
</paragraph>
<paragraph id="1A09685AFFDAFFA1FCC2FC44FAF7FBD9" blockId="9.[832,1521,882,1965]" pageId="9" pageNumber="8">
<emphasis id="28C2B448FFDAFFA1FCC2FC44FBDFFBFC" bold="true" box="[864,1045,1023,1049]" pageId="9" pageNumber="8">Occurrence—</emphasis>
<materialsCitation id="AADE6207FFDAFFA1FBB6FC44FAF3FBD9" county="Lewis County" location="upper Eocene" pageId="9" pageNumber="8" specimenCount="1" stateProvince="Washington">
Middle to
<location id="1F693E81FFDAFFA1FB28FBBBFAFBFBFF" LSID="urn:lsid:plazi:treatment:921FD94CFFDAFFA2FF67F936FCEBFCD7:1F693E81FFDAFFA1FB28FBBBFAFBFBFF" box="[1162,1329,1024,1050]" county="Lewis County" name="upper Eocene" pageId="9" pageNumber="8" stateProvince="Washington">upper Eocene</location>
(“Tejon Stage”), south-central
<collectingCounty id="F36810D6FFDAFFA1FC4CFB99FB5DFBD9" box="[1006,1175,1058,1084]" pageId="9" pageNumber="8">Lewis County</collectingCounty>
,
<collectingRegion id="D872A6B8FFDAFFA1FB03FB99FAF3FBD9" box="[1185,1337,1058,1084]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="52AC3BD1FFDAFFA1FCC2FBFDFA6CFB66" pageId="9" pageNumber="8" type="etymology">
<paragraph id="1A09685AFFDAFFA1FCC2FBFDFA6CFB66" blockId="9.[832,1521,882,1965]" pageId="9" pageNumber="8">
<emphasis id="28C2B448FFDAFFA1FCC2FBFDFBC0FB85" bold="true" box="[864,1034,1094,1120]" pageId="9" pageNumber="8">Etymology—</emphasis>
Originally named for the Eocene Cowlitz Formation, Lewis County, southwest
<collectingRegion id="D872A6B8FFDAFFA1FAA8FBD2FA68FB66" box="[1290,1442,1129,1155]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="52AC3BD1FFDAFFA1FCC2FB30FB7BF9E3" pageId="9" pageNumber="8" type="description">
<paragraph id="1A09685AFFDAFFA1FCC2FB30FB7BF9E3" blockId="9.[832,1521,882,1965]" pageId="9" pageNumber="8">
<emphasis id="28C2B448FFDAFFA1FCC2FB30FBD0FB40" bold="true" box="[864,1050,1163,1189]" pageId="9" pageNumber="8">Description—</emphasis>
Shell of size medium size. Shape ovoid. Spire covered. Maximum height slightly posterior of midpoint. Dorsal groove faint; dorsal nodules smooth, circular, connected by fine threads that extend onto basal surface and form prominent denticulation. Marginal cal- lus slight. Aperture curved slightly posteriorly toward columella. Denticulation prominent with smooth inter- stices, labial lip with 19 teeth, and columellar lip with 18 teeth. Fossula with strong denticulation. Anterior and posterior canals prominently lengthened by terminal teeth. (
<bibRefCitation id="7E2715ABFFDAFFA1FC3AFA57FB8EF9E3" author="Groves, L. T." box="[920,1092,1516,1542]" pageId="9" pageNumber="8" pagination="79 - 87" refId="ref37896" refString="Groves, L. T. 1994 a. New species of Cypraeidae (Mollusca: Gastropoda) from the Miocene of California and the Eocene of Washington. The Veliger 37 (3): 244 - 252, figs. 1 - 13 [reprinted in Bulletin of the Southern California Paleontological Society 26 (9 - 10): 79 - 87]." type="journal article" year="1994">Groves 1994a</bibRefCitation>
: p. 247).
</paragraph>
</subSubSection>
<subSubSection id="52AC3BD1FFDAFFA2FCC2F9B4FCEBFCD7" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="discussion">
<paragraph id="1A09685AFFDAFFA2FCC2F9B4FE17FEC7" blockId="9.[832,1521,882,1965]" lastBlockId="10.[112,801,192,818]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
<emphasis id="28C2B448FFDAFFA1FCC2F9B4FC3CF9CC" bold="true" box="[864,1014,1551,1577]" pageId="9" pageNumber="8">Remarks—</emphasis>
The
<typeStatus id="C50DD6F8FFDAFFA1FB92F9B4FB55F9CC" box="[1072,1183,1551,1577]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
of this Cowlitz Formation species is the only known record of this genus in the eastern Pacific.
<bibRefCitation id="7E2715ABFFDAFFA1FBADF9EEFB7AF995" author="Fehse, D." box="[1039,1200,1621,1648]" pageId="9" pageNumber="8" pagination="1 - 51" refId="ref37011" refString="Fehse, D. 2001. Beitrage zur Kenntnis der Ovuliade (Mollusca: Cypraeoidea). 8. Einleitung zur familie sowie Katalog, Taxonomy und Bibliographie und Bemerkungen zu verwandten Gruppen. Acta Conchyliorum 5: 1 - 51, figs. 1 - 3." type="journal article" year="2001">Fehse (2001</bibRefCitation>
: pp. 18, 37) assigned
<taxonomicName id="DDB613D9FFDAFFA1FA75F9EFFC1BF976" class="Gastropoda" family="Pediculariidae" genus="Jenneria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="cowlitziana">
<emphasis id="28C2B448FFDAFFA1FA75F9EFFC1BF976" italics="true" pageId="9" pageNumber="8">N. cowlitziana</emphasis>
</taxonomicName>
erroneously to the ovulid genus
<taxonomicName id="DDB613D9FFDAFFA1FA2AF9CCFBECF953" authority="Jousseaume, 1884" authorityName="Jousseaume" authorityYear="1884" class="Gastropoda" family="Pediculariidae" genus="Jenneria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="28C2B448FFDAFFA1FA2AF9CCFA3BF976" box="[1416,1521,1655,1683]" italics="true" pageId="9" pageNumber="8">Jenneria</emphasis>
<bibRefCitation id="7E2715ABFFDAFFA1FCE2F927FBECF953" author="Jousseaume, F. P." box="[832,1062,1692,1718]" pageId="9" pageNumber="8" pagination="81 - 100" refId="ref39191" refString="Jousseaume, F. P. 1884. Etude sur la famille des Cypaeidae. Bulletin de la Societe Zoologique 9: 81 - 100." type="journal article" year="1884">Jousseaume, 1884</bibRefCitation>
</taxonomicName>
, but later, following an examination of the
<typeStatus id="C50DD6F8FFDAFFA1FC28F904FC30F93C" box="[906,1018,1727,1753]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
, Fehse acknowledged his error (personal communication,
<date id="6E084E9AFFDAFFA1FBB1F959FB47F919" box="[1043,1165,1762,1788]" pageId="9" pageNumber="8" value="2004-07">July 2004</date>
).
<bibRefCitation id="7E2715ABFFDAFFA1FB0AF959FA22F919" author="Dolin, L. &amp; P. Lozouet" box="[1192,1512,1762,1789]" pageId="9" pageNumber="8" pagination="1 - 164" refId="ref36614" refString="Dolin, L. and P. Lozouet. 2004. Nouvelles especes de gasteropodes (Mollusca: Gastropoda) de l'Oligocene et du Miocene inferieur de l'Aquitaine (Sud-Ouest de la France). Partie 3. Cypraeidae et Ovulidae. Cossmanniana, hors serie, 4: 1 - 164, figs. 1 - 27, pls. 1 - 36." type="journal article" year="2004">Dolin and Lozouet (2004</bibRefCitation>
: p. 73) inexplicably questioned the provenance of this specimen and compared it to the living species
<taxonomicName id="DDB613D9FFDAFFA1FAD5F89CFBD8F883" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Cypraeidae" genus="Cypraea" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="nucleus">
<emphasis id="28C2B448FFDAFFA1FAD5F89CFA3AF8A6" box="[1399,1520,1831,1859]" italics="true" pageId="9" pageNumber="8">N. nucleus</emphasis>
(
<bibRefCitation id="7E2715ABFFDAFFA1FCE9F8F7FBCCF882" author="Linnaeus, C." box="[843,1030,1868,1895]" pageId="9" pageNumber="8" refId="ref39466" refString="Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae … Editio decimal, reformata. Vol. (Regnum animale), part. Holmiae. Impensis Direct. Laurentii Salvii. Stockholm. 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
. In the same publication they described
<taxonomicName id="DDB613D9FFDAFFA1FCE2F8D5FB98F86E" authorityName="Dolin &amp; Lozouet" authorityYear="2004" box="[832,1106,1902,1931]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="species" species="bezoyensis">
<emphasis id="28C2B448FFDAFFA1FCE2F8D5FB98F86E" box="[832,1106,1902,1931]" italics="true" pageId="9" pageNumber="8">Nucleolaria bezoyensis</emphasis>
</taxonomicName>
and mistakenly claimed that their species (now reassigned to the genus
<taxonomicName id="DDB613D9FFDAFFA1FABDF82AFAAFF848" authorityName="Gray" authorityYear="1837" box="[1311,1381,1937,1965]" class="Gastropoda" family="Cypraeidae" genus="Naria" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="28C2B448FFDAFFA1FABDF82AFAAFF848" box="[1311,1381,1937,1965]" italics="true" pageId="9" pageNumber="8">Naria</emphasis>
</taxonomicName>
by
<bibRefCitation id="7E2715ABFFDAFFA2FA31F828FF7BFF39" author="Lorenz, F." lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" refId="ref39506" refString="Lorenz, F. 2017. Cowries. A guide to the gastropod family Cypraeidae. Volume 1. Biology and Systematics. Conch- Books, Harxheim, Germany. 644 pp., numerous figures." type="book" year="2017">Lorenz, 2017</bibRefCitation>
) was the earliest record of
<emphasis id="28C2B448FFD9FFA2FDB2FF7BFD6EFF39" box="[528,676,192,220]" italics="true" pageId="10" pageNumber="9">
<taxonomicName id="DDB613D9FFD9FFA2FDB2FF7BFD6BFF39" authorityName="OYAMA" authorityYear="1959" box="[528,673,192,220]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="9" phylum="Mollusca" rank="genus">Nucleolaria</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="DDB613D9FFD9FFA2FFD2FF58FED5FF1A" box="[112,287,227,255]" class="Gastropoda" family="Pediculariidae" genus="Jenneria" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="9" phylum="Mollusca" rank="species" species="cowlitziana">
<emphasis id="28C2B448FFD9FFA2FFD2FF58FED5FF1A" box="[112,287,227,255]" italics="true" pageId="10" pageNumber="9">N. cowlitziana</emphasis>
</taxonomicName>
remains the earliest record of the genus and its provenance is certain.
</paragraph>
<paragraph id="1A09685AFFD9FFA2FF32FE90FCEBFCD7" blockId="10.[112,801,192,818]" pageId="10" pageNumber="9">
This earliest known occurrence of
<taxonomicName id="DDB613D9FFD9FFA2FDC5FE91FD37FEA3" authorityName="OYAMA" authorityYear="1959" box="[615,765,298,326]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="28C2B448FFD9FFA2FDC5FE91FD37FEA3" box="[615,765,298,326]" italics="true" pageId="10" pageNumber="9">Nucleolaria</emphasis>
</taxonomicName>
in the NEP region is an enigma. It is likely indicative of an eastward-directed faunistic influx during the Paleogene between the warm waters of the central Pacific Ocean and the NEP region.
<taxonomicName id="DDB613D9FFD9FFA2FECBFE0CFE32FE36" authorityName="OYAMA" authorityYear="1959" box="[361,504,439,467]" class="Gastropoda" family="Cypraeidae" genus="Nucleolaria" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="28C2B448FFD9FFA2FECBFE0CFE32FE36" box="[361,504,439,467]" italics="true" pageId="10" pageNumber="9">Nucleolaria</emphasis>
</taxonomicName>
today is a tropical Indo-Pacific genus. Like most other extant cypraeoideans, its larvae are planktonic (
<bibRefCitation id="7E2715ABFFD9FFA2FE11FE45FDAAFDFD" author="Groves, L. T." box="[435,608,510,536]" pageId="10" pageNumber="9" pagination="79 - 87" refId="ref37896" refString="Groves, L. T. 1994 a. New species of Cypraeidae (Mollusca: Gastropoda) from the Miocene of California and the Eocene of Washington. The Veliger 37 (3): 244 - 252, figs. 1 - 13 [reprinted in Bulletin of the Southern California Paleontological Society 26 (9 - 10): 79 - 87]." type="journal article" year="1994">Groves 1994a</bibRefCitation>
).
<bibRefCitation id="7E2715ABFFD9FFA2FDDBFE44FEE7FDD9" author="Emerson, W. K. &amp; H. W. Chaney" pageId="10" pageNumber="9" pagination="8 - 21" refId="ref36961" refString="Emerson, W. K. and H. W. Chaney. 1995. A zoogeographic review of the Cypraeidae (Mollsuca: Gastropoda) occurring in the eastern Pacific Ocean. The Veliger 38 (1): 8 - 21, figs. 1 - 15." type="journal article" year="1995">Emerson and Chaney (1995)</bibRefCitation>
documented 15 species of Indo-Pacific cypraeid species from the eastern Pacific, three of which are based on single specimens.
<bibRefCitation id="7E2715ABFFD9FFA2FDAEFDD3FD0CFD67" author="Lorenz, F." box="[524,710,616,642]" pageId="10" pageNumber="9" refId="ref39506" refString="Lorenz, F. 2017. Cowries. A guide to the gastropod family Cypraeidae. Volume 1. Biology and Systematics. Conch- Books, Harxheim, Germany. 644 pp., numerous figures." type="book" year="2017">Lorenz (2017)</bibRefCitation>
recog- nized only a single Indo-Pacific species established in the eastern Pacific.
<bibRefCitation id="7E2715ABFFD9FFA2FEC4FD15FDB1FD2C" author="Lindberg, D. R. &amp; B. Roth &amp; M. G. Kellogg &amp; C. L. Hubbs" box="[358,635,686,713]" pageId="10" pageNumber="9" refId="ref39381" refString="Lindberg, D. R., B. Roth, M. G. Kellogg, and C. L. Hubbs. 1980. Invertebrate megafossils of Pleistocene (Sangamon In- terglagial) age from Isla de Guadalupe, Baja California, Mexico. Pp. 41 - 62, figs. 1 - 4 in D. M. Power (ed.). The California islands: Proceedings of a multidisciplinary symposium. Santa Barbara Museum of Natural History." type="proceedings paper" year="1980">Lindberg et al. (1980)</bibRefCitation>
documented the only other fossil Indo-Pacific species [
<taxonomicName id="DDB613D9FFD9FFA2FDDBFD6BFE5BFCEA" authority="(G. B. Sowerby II, 1870)" baseAuthorityName="G. B. Sowerby II" baseAuthorityYear="1870" class="Gastropoda" family="Cypraeidae" genus="Naria" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="9" phylum="Mollusca" rank="species" species="cernica">
<emphasis id="28C2B448FFD9FFA2FDDBFD6BFCEAFD09" box="[633,800,720,748]" italics="true" pageId="10" pageNumber="9">Naria cernica</emphasis>
(G.B. Sowerby
<bibRefCitation id="7E2715ABFFD9FFA2FE8BFD4FFE4DFCEA" author="Sowerby, G. B. II." box="[297,391,756,783]" pageId="10" pageNumber="9" pagination="1 - 58" refId="ref42574" refString="Sowerby, G. B. II. 1870. A monograph of the genus Cy- praea. Thesaurus Conchyliorum, or figures and descrip- tions of recent shells 4: 1 - 58, pls. 1 - 37." type="journal article" year="1870">II,1870</bibRefCitation>
)
</taxonomicName>
] in the eastern Pacific from the Pleistocene of Isla de Guadalupe,
<collectingRegion id="D872A6B8FFD9FFA2FDA1FCA3FD73FCD7" box="[515,697,792,818]" country="Mexico" name="Baja California" pageId="10" pageNumber="9">Baja California</collectingRegion>
,
<collectingCountry id="62A128CAFFD9FFA2FD60FCA3FCD7FCD7" box="[706,797,792,818]" name="Mexico" pageId="10" pageNumber="9">México</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>