Revision of northeast Pacific Paleogene cypraeoidean gastropods, including recognition of three new species: implications for paleobiogeographic distribution and faunal turnover Author Groves, Lindsey Author Squires, Richard L. text PaleoBios 2023 2023-08-10 40 10 1 52 http://dx.doi.org/10.5070/p9401057774 journal article 10.5070/P9401057774 0031-0298 10913295 11600574-2B0E-4C13-BC08-A3A5EF9EE562 NUCLEOLARIA COWLITZIANA GROVES, 1994 A FIGS. 4A–C Nucleolaria cowlitziana Groves, 1994a , p. 247 –248, figs. 6, 7. Nesbitt, 1995 . p. 1066. Groves, 1997 . p. 7. Groves, 2000 . p. 120. Dolin and Lozouet, 2004 . p. 73. Lorenz, 2017 . p. 220. Lorenz, 2018 . p. 664, pl. 328, fig. 5. Jenneria cowlitziana (Groves) . Fehse, 2001 . p. 37. Holotype and Type Locality — UCMP 39837 ( Figs. 4A–C ), length 27.2 mm , width 17.3 mm , height 11.3 mm . UCMP Locality D-8040, Cowlitz Formation , south-central Lewis County , Washington . Occurrence— Middle to upper Eocene (“Tejon Stage”), south-central Lewis County , Washington . Etymology— Originally named for the Eocene Cowlitz Formation, Lewis County, southwest Washington . Description— Shell of size medium size. Shape ovoid. Spire covered. Maximum height slightly posterior of midpoint. Dorsal groove faint; dorsal nodules smooth, circular, connected by fine threads that extend onto basal surface and form prominent denticulation. Marginal cal- lus slight. Aperture curved slightly posteriorly toward columella. Denticulation prominent with smooth inter- stices, labial lip with 19 teeth, and columellar lip with 18 teeth. Fossula with strong denticulation. Anterior and posterior canals prominently lengthened by terminal teeth. ( Groves 1994a : p. 247). Remarks— The holotype of this Cowlitz Formation species is the only known record of this genus in the eastern Pacific. Fehse (2001 : pp. 18, 37) assigned N. cowlitziana erroneously to the ovulid genus Jenneria Jousseaume, 1884 , but later, following an examination of the holotype , Fehse acknowledged his error (personal communication, July 2004 ). Dolin and Lozouet (2004 : p. 73) inexplicably questioned the provenance of this specimen and compared it to the living species N. nucleus ( Linnaeus, 1758 ) . In the same publication they described Nucleolaria bezoyensis and mistakenly claimed that their species (now reassigned to the genus Naria by Lorenz, 2017 ) was the earliest record of Nucleolaria . However, N. cowlitziana remains the earliest record of the genus and its provenance is certain. This earliest known occurrence of Nucleolaria in the NEP region is an enigma. It is likely indicative of an eastward-directed faunistic influx during the Paleogene between the warm waters of the central Pacific Ocean and the NEP region. Nucleolaria today is a tropical Indo-Pacific genus. Like most other extant cypraeoideans, its larvae are planktonic ( Groves 1994a ). Emerson and Chaney (1995) documented 15 species of Indo-Pacific cypraeid species from the eastern Pacific, three of which are based on single specimens. Lorenz (2017) recog- nized only a single Indo-Pacific species established in the eastern Pacific. Lindberg et al. (1980) documented the only other fossil Indo-Pacific species [ Naria cernica (G.B. Sowerby II,1870 ) ] in the eastern Pacific from the Pleistocene of Isla de Guadalupe, Baja California , México .