351 lines
54 KiB
XML
351 lines
54 KiB
XML
<document id="E9C13F3B9D282EBB2EEBA93EE3B5F8CE" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFCAFFF8BF72FDC00C4F07D0" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus tortuosiflagellatus Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="35" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="33" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="998BE8AB6DB2058C8E3BF2AA581E6C12">
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<mods:title id="374F7264A1998C33FF788E33EF17E979">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="2F000DE7F0493D68EEC7A157956127B3" type="personal">
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<mods:roleTerm id="935C85C6B48F7F39FEBA65DC86F57974">Author</mods:roleTerm>
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<mods:namePart id="EB30DB99DB24790E887A752B1A149FD6">Frisch, Johannes</mods:namePart>
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<mods:name id="6D59DF2D41DC06EE6967D0AA5241F100" type="personal">
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<mods:roleTerm id="509C3550D80D10939587BB44D513CAB8">Author</mods:roleTerm>
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<mods:namePart id="1ED96A45EF911A175D5FDA24B0C7C61A">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="E4C9CCBE2A82EC55B15CB38CFEA2FCC6">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="9A48F4BE5578303F08F3C4FAF1948905">Soil Organisms</mods:title>
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<mods:part id="79376B52D2529F2CE242F3F959359A92">
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<mods:date id="ED6F6272CD6716E40A9FA69DF8C2A894">2022</mods:date>
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<mods:number id="F8CAD3A19E65555351585C14DDE404B5">2022-04-01</mods:number>
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<mods:detail id="22E25CE9CBBB56DDB28DDC1F9F531371" type="volume">
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<mods:number id="536B72A6C049BDFF134BD97034682321">95</mods:number>
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<mods:start id="3DACF838CF7F37E607470D2D0F5F2D8F">23</mods:start>
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<mods:url id="D098252A7ED5D9BA529634BD0A56EB79">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="B6962B4958CB8FA3CC324C163C75A7E4">journal article</mods:classification>
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<mods:identifier id="FA906B6BE18FBE9F4045DB99CED5B8E0" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFCAFFF8BF72FDC00C4F07D0" ID-DOI="http://doi.org/10.5281/zenodo.10724268" ID-Zenodo-Dep="10724268" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFF8BF72FDC00C4F07D0" httpUri="http://treatment.plazi.org/id/03B7C77AFFCAFFF8BF72FDC00C4F07D0" lastPageId="12" lastPageNumber="35" pageId="10" pageNumber="33">
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<subSubSection id="C30425E7FFCAFFFEBF72FDC00841065E" box="[809,1372,568,595]" pageId="10" pageNumber="33" type="nomenclature">
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<paragraph id="8BA1766CFFCAFFFEBF72FDC00841065E" blockId="10.[809,1372,568,628]" box="[809,1372,568,595]" pageId="10" pageNumber="33">
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<heading id="D0E9C100FFCAFFFEBF72FDC00841065E" box="[809,1372,568,595]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="33" reason="5">
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<taxonomicName id="4C1E0DEFFFCAFFFEBF72FDC009C7065F" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[809,1242,568,595]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="tortuosiflagellatus" status="sp. nov.">
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<emphasis id="B96AAA7EFFCAFFFEBF72FDC009C7065F" bold="true" box="[809,1242,568,595]" pageId="10" pageNumber="33">
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<emphasis id="B96AAA7EFFCAFFFEBF72FDC0099C065E" bold="true" box="[809,1153,568,595]" italics="true" pageId="10" pageNumber="33">Scopaeus tortuosiflagellatus</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFCAFFFEB8B3FDC10841065E" box="[1256,1372,569,595]" pageId="10" pageNumber="33" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFCAFFFEBF72FDA109A90679" box="[809,1204,601,628]" pageId="10" pageNumber="33" type="description">
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<paragraph id="8BA1766CFFCAFFFEBF72FDA109A90679" blockId="10.[809,1372,568,628]" box="[809,1204,601,628]" pageId="10" pageNumber="33">
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(
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<figureCitation id="13256AE9FFCAFFFEBF6AFDA10E610679" box="[817,892,601,628]" captionStart="Figures 8–10" captionStartId="5.[189,262,1930,1952]" captionTargetBox="[237,1406,902,1874]" captionTargetId="figure-357@5.[189,1436,889,1911]" captionTargetPageId="5" captionText="Figures 8–10. Habitus of Scopaeus tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (8); S. posoanus, paratype, Sulawesi Tengah: Lake Poso (9); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (10)." figureDoi="http://doi.org/10.5281/zenodo.10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="10" pageNumber="33">Figs 8</figureCitation>
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,
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<figureCitation id="13256AE9FFCAFFFEBFD2FDA20EBA0679" box="[905,935,602,628]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="10" pageNumber="33">33</figureCitation>
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,
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<figureCitation id="13256AE9FFCAFFFEBFE8FDA109180679" box="[947,1029,601,628]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">68 –70</figureCitation>
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,
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<figureCitation id="13256AE9FFCAFFFEB84AFDA109230679" box="[1041,1086,601,628]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="10" pageNumber="33">125</figureCitation>
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,
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<figureCitation id="13256AE9FFCAFFFEB812FDA109B10679" box="[1097,1196,601,628]" captionStart="Figures 146–154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146–154. Sperm pump (146–154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150–152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="10" pageNumber="33">153, 154</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFCAFFFEBF72FD65092F07F3" pageId="10" pageNumber="33" type="materials_examined">
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<paragraph id="8BA1766CFFCAFFFEBF72FD65092F07F3" blockId="10.[786,1399,669,1976]" pageId="10" pageNumber="33">
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<emphasis id="B96AAA7EFFCAFFFEBF72FD650EFA06B8" bold="true" box="[809,999,669,693]" pageId="10" pageNumber="33">Type specimens:</emphasis>
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||
<materialsCitation id="3B767C31FFCAFFFEBFB4FD65086E06FA" collectingDate="2015-09-19" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="870" latitude="-6.755833" location="Cisalimas River" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" pageId="10" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
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<collectingCountry id="F30936FCFFCAFFFEBFB4FD65094606B8" box="[1007,1115,669,693]" name="Indonesia" pageId="10" pageNumber="33">Indonesia</collectingCountry>
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,
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<collectingRegion id="49DAB88EFFCAFFFEB83CFD6509F906B9" box="[1127,1252,669,692]" country="Indonesia" name="Jawa Barat" pageId="10" pageNumber="33">Jawa Barat</collectingRegion>
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:
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<typeStatus id="54A5C8CEFFCAFFFEB8B4FD65084E06B8" box="[1263,1363,669,693]" pageId="10" pageNumber="33" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFCAFFFEB907FD65086F06B9" box="[1372,1394,669,692]" count="1" pageId="10" pageNumber="33" type="male">♂</specimenCount>
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,
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<collectingCounty id="62C00EE0FFCAFFFEBF49FD460E9F06DB" box="[786,898,702,726]" pageId="10" pageNumber="33">Sukabumi</collectingCounty>
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,
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<collectingMunicipality id="6BC5EC16FFCAFFFEBFC8FD460EE006DB" box="[915,1021,702,726]" pageId="10" pageNumber="33">Cisalimas</collectingMunicipality>
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(
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<location id="8EC120B7FFCAFFFEB849FD4609D606DB" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFF8BF72FDC00C4F07D0:8EC120B7FFCAFFFEB849FD4609D606DB" box="[1042,1227,702,726]" country="Indonesia" county="Sukabumi" latitude="-6.755833" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" name="Cisalimas River" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Cisalimas River</location>
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), (
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<geoCoordinate id="EE2A10ABFFCAFFFEB8B7FD46086C06DB" box="[1260,1393,702,726]" degrees="06" direction="south" minutes="45" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="21" value="-6.755833">06°45’21’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFCAFFFEBF49FD270EBA06FA" box="[786,935,735,759]" degrees="106" direction="east" minutes="33" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="38" value="106.56056">106°33’38’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFCAFFFEBF9AFD27091306FB" box="[961,1038,735,759]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="33" unit="m" value="870.0">
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<elevation id="0033915FFFCAFFFEBF9AFD27091306FB" box="[961,1038,735,759]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="33" unit="m" value="870.0">870 m</elevation>
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</quantity>
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,
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<date id="FFA050ACFFCAFFFEB844FD27099006FA" box="[1055,1165,735,759]" pageId="10" pageNumber="33" value="2015-09-19">
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<collectingDate id="EFE4A944FFCAFFFEB844FD27099006FA" box="[1055,1165,735,759]" pageId="10" pageNumber="33" value="2015-09-19">19.9.2015</collectingDate>
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</date>
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, leg.
|
||
<collectorName id="26EB13BAFFCAFFFEB888FD27080B06FA" box="[1235,1302,735,759]" pageId="10" pageNumber="33">Frisch</collectorName>
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(
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||
<collectionCode id="ED0FEEA9FFCAFFFEB970FD27087606FA" box="[1323,1387,735,759]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
|
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)
|
||
</materialsCitation>
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||
.
|
||
<typeStatus id="54A5C8CEFFCAFFFEBF49FCF80E660715" box="[786,891,768,792]" pageId="10" pageNumber="33" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D18BDE5FFCAFFFEBFCBFD07093D0715" box="[912,1056,767,792]" count="6" pageId="10" pageNumber="33" type="generic">6 specimens</specimenCount>
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):
|
||
<materialsCitation id="3B767C31FFCAFFFEB86CFCF80E690734" collectingDate="2015-09-19" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="870" latitude="-6.755833" location="Cisalimas River" longLatPrecision="21" longitude="106.56056" municipality="Cisalimas" pageId="10" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
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<specimenCount id="9D18BDE5FFCAFFFEB86CFCF80978071A" box="[1079,1125,768,792]" count="1" pageId="10" pageNumber="33" type="female">1 ♀</specimenCount>
|
||
, same data as holotype (
|
||
<collectionCode id="ED0FEEA9FFCAFFFEBF41FCD90E710734" box="[794,876,801,825]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="33">MFNB</collectionCode>
|
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)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFCAFFFEBFD3FCD9086F0754" collectingDate="2015-03-10" collectionCode="MZB" collectorName="Frisch" country="Indonesia" elevation="990" latitude="-6.7113886" location="Pamijahan" longLatPrecision="21" longitude="106.684166" municipality="Bogor" pageId="10" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFCAFFFEBFD3FCD90EA60735" box="[904,955,801,825]" count="1" pageId="10" pageNumber="33" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFCAFFFEBF8BFCD909080734" box="[976,1045,801,825]" pageId="10" pageNumber="33">Bogor</collectingMunicipality>
|
||
, SW
|
||
<location id="8EC120B7FFCAFFFEB83AFCD809CE0735" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFF8BF72FDC00C4F07D0:8EC120B7FFCAFFFEB83AFCD809CE0735" box="[1121,1235,800,824]" country="Indonesia" latitude="-6.7113886" longLatPrecision="21" longitude="106.684166" municipality="Bogor" name="Pamijahan" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Pamijahan</location>
|
||
(
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<geoCoordinate id="EE2A10ABFFCAFFFEB8B7FCD8086F0735" box="[1260,1394,800,824]" degrees="06" direction="south" minutes="42" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="41" value="-6.7113886">06°42’41’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFCAFFFEBF49FCB90EBA0754" box="[786,935,833,857]" degrees="106" direction="east" minutes="41" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="03" value="106.684166">106°41’03’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFCAFFFEBF9AFCB909100754" box="[961,1037,833,857]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="10" pageNumber="33" unit="m" value="990.0">
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<elevation id="0033915FFFCAFFFEBF9AFCB909100754" box="[961,1037,833,857]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="10" pageNumber="33" unit="m" value="990.0">990 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFCAFFFEB844FCB909900754" box="[1055,1165,833,857]" pageId="10" pageNumber="33" value="2015-03-10">
|
||
<collectingDate id="EFE4A944FFCAFFFEB844FCB909900754" box="[1055,1165,833,857]" pageId="10" pageNumber="33" value="2015-03-10">3.10.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFCAFFFEB888FCB9080B0754" box="[1235,1302,833,857]" pageId="10" pageNumber="33">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFCAFFFEB970FCB908770754" box="[1323,1386,833,857]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFCAFFFEBF49FC9A095B07B1" collectingDate="2015-09-30" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="1150" latitude="-6.745278" location="Javana Spa" longLatPrecision="21" longitude="106.714165" municipality="Cidahu" pageId="10" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFCAFFFEBF49FC9A0E540777" box="[786,841,866,890]" count="1" pageId="10" pageNumber="33" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D18BDE5FFCAFFFEBF3AFC9A0E880777" box="[865,917,866,890]" count="1" pageId="10" pageNumber="33" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFCAFFFEBFF4FC9A09020777" box="[943,1055,866,890]" pageId="10" pageNumber="33">Sukabumi</collectingCounty>
|
||
, NNW
|
||
<collectingMunicipality id="6BC5EC16FFCAFFFEB8D2FC9A09C60777" box="[1161,1243,866,890]" pageId="10" pageNumber="33">Cidahu</collectingMunicipality>
|
||
:
|
||
<location id="8EC120B7FFCAFFFEB8A9FC9B086B0777" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFF8BF72FDC00C4F07D0:8EC120B7FFCAFFFEB8A9FC9B086B0777" box="[1266,1398,866,890]" country="Indonesia" county="Sukabumi" latitude="-6.745278" longLatPrecision="21" longitude="106.714165" municipality="Cidahu" name="Javana Spa" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Javana Spa</location>
|
||
(
|
||
<geoCoordinate id="EE2A10ABFFCAFFFEBF40FC7B0EBC0796" box="[795,929,899,923]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="43" value="-6.745278">06°44’43’’S</geoCoordinate>
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||
,
|
||
<geoCoordinate id="EE2A10ABFFCAFFFEBF9BFC7B09480796" box="[960,1109,899,923]" degrees="106" direction="east" minutes="42" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="51" value="106.714165">106°42’51’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFCAFFFEB825FC7B09F90796" box="[1150,1252,899,923]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="10" pageNumber="33" unit="m" value="1150.0">
|
||
<elevation id="0033915FFFCAFFFEB825FC7B09F90796" box="[1150,1252,899,923]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="10" pageNumber="33" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFCAFFFEB95FFC7B086F0796" box="[1284,1394,899,923]" pageId="10" pageNumber="33" value="2015-09-30">
|
||
<collectingDate id="EFE4A944FFCAFFFEB95FFC7B086F0796" box="[1284,1394,899,923]" pageId="10" pageNumber="33" value="2015-09-30">30.9.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFCAFFFEBF1CFC5C0E9707B1" box="[839,906,932,956]" pageId="10" pageNumber="33">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFCAFFFEBFC5FC5C0EF307B1" box="[926,1006,932,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="33">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFCAFFFEBFA4FC5C092307B1" box="[1023,1086,932,956]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFCAFFFEB80CFC5C093307F3" collectingDate="2015-05-10" collectionCode="MFNB, MZB" country="Indonesia" county="Sukabumi" elevation="1020" latitude="-6.7461114" location="Cikaniki River" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" pageId="10" pageNumber="33" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFCAFFFEB80CFC5C099807B6" box="[1111,1157,932,956]" count="2" pageId="10" pageNumber="33" type="female">2 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFCAFFFEB8C3FC5C081407B1" box="[1176,1289,932,956]" pageId="10" pageNumber="33">Sukabumi</collectingCounty>
|
||
:
|
||
<collectingMunicipality id="6BC5EC16FFCAFFFEB941FC5C086A07B1" box="[1306,1399,932,956]" pageId="10" pageNumber="33">Cikaniki</collectingMunicipality>
|
||
(
|
||
<location id="8EC120B7FFCAFFFEBF42FC3D0EDC07D0" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFF8BF72FDC00C4F07D0:8EC120B7FFCAFFFEBF42FC3D0EDC07D0" box="[793,961,965,989]" country="Indonesia" county="Sukabumi" latitude="-6.7461114" longLatPrecision="21" longitude="106.540276" municipality="Cikaniki" name="Cikaniki River" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Cikaniki River</location>
|
||
), (
|
||
<geoCoordinate id="EE2A10ABFFCAFFFEBF85FC3D097907D0" box="[990,1124,965,989]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="46" value="-6.7461114">06°44’46’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2A10ABFFCAFFFEB82AFC3D081B07D0" box="[1137,1286,965,989]" degrees="106" direction="east" minutes="32" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="25" value="106.540276">106°32’25’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFCAFFFEB947FC3D086F07D1" box="[1308,1394,965,989]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="10" pageNumber="33" unit="m" value="1020.0">
|
||
<elevation id="0033915FFFCAFFFEB947FC3D086F07D1" box="[1308,1394,965,989]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="10" pageNumber="33" unit="m" value="1020.0">1020 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFCAFFFEBF49FC1E0E6307F3" box="[786,894,998,1022]" pageId="10" pageNumber="33" value="2015-05-10">
|
||
<collectingDate id="EFE4A944FFCAFFFEBF49FC1E0E6307F3" box="[786,894,998,1022]" pageId="10" pageNumber="33" value="2015-05-10">5.10.2015</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFCAFFFEBFD7FC1E0EC107F3" box="[908,988,998,1022]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="33">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFCAFFFEBFB3FC1E093A07F3" box="[1000,1063,998,1022]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFCAFFF8BF72FBFE0FFE06B8" lastPageId="12" lastPageNumber="35" pageId="10" pageNumber="33" type="description">
|
||
<paragraph id="8BA1766CFFCAFFFEBF72FBFE0EBD01A4" blockId="10.[786,1399,669,1976]" pageId="10" pageNumber="33">
|
||
<emphasis id="B96AAA7EFFCAFFFEBF72FBFE0EAB0013" bold="true" box="[809,950,1030,1054]" pageId="10" pageNumber="33">Description:</emphasis>
|
||
Habitus and coloring as in
|
||
<figureCitation id="13256AE9FFCAFFFEB8B6FBFF082D0012" box="[1261,1328,1031,1055]" captionStart="Figures 8–10" captionStartId="5.[189,262,1930,1952]" captionTargetBox="[237,1406,902,1874]" captionTargetId="figure-357@5.[189,1436,889,1911]" captionTargetPageId="5" captionText="Figures 8–10. Habitus of Scopaeus tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (8); S. posoanus, paratype, Sulawesi Tengah: Lake Poso (9); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (10)." figureDoi="http://doi.org/10.5281/zenodo.10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="10" pageNumber="33">Fig. 8</figureCitation>
|
||
. Head subquadrate. Antennae with elongate penultimate segment. Mesotibia slender. Forebody surface with fine, setose punctation, shiny; abdomen matter than forebody. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body black-brown except for narrow, yellow-brown, posterior margin of elytra, light brown tip of abdomen, medium brown maxillary palpi with darker penultimate segment, and yellow-brown legs; antennae with scapus medium brown, from blackish pedicellus gradually lighter towards terminal segments. Total body length
|
||
<quantity id="4CE6DB89FFCAFFFEB84BFA8809860185" box="[1040,1179,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="10" pageNumber="33" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4 – 2.9 mm</quantity>
|
||
; forebody length
|
||
<quantity id="4CE6DB89FFCAFFFEBF49FA690E8101A4" box="[786,924,1425,1449]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="10" pageNumber="33" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5 – 1.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFCAFFF8BF72FA4A0DE505C2" blockId="10.[786,1399,669,1976]" lastBlockId="12.[151,764,241,989]" lastPageId="12" lastPageNumber="35" pageId="10" pageNumber="33">
|
||
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with bisinuate emargination with strongly convex median third in about posterior fifth of sternite length (
|
||
<figureCitation id="13256AE9FFCAFFFEB8C9F9ED09EE0220" box="[1170,1267,1557,1581]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="10" pageNumber="33">Fig. 125</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13256AE9FFCAFFFEBF41F9CE0E700243" box="[794,877,1590,1614]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="10" pageNumber="33">Figs 33</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFCAFFFEBF22F9CD0EDF0243" box="[889,962,1589,1614]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">68 – 70</figureCitation>
|
||
) about
|
||
<quantity id="4CE6DB89FFCAFFFEB84EF9CE09790240" box="[1045,1124,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="33" unit="mm" value="1.1">1.1 mm</quantity>
|
||
long, with asymmetrical distal lobes; dextral apical lobe longer than acute sinistral apical lobe, enlarged apicad and with conspicuous, very slender, proximodorsad curved spine of ventral margin (
|
||
<figureCitation id="13256AE9FFCAFFFEBF28F9410EE502DC" box="[883,1016,1721,1745]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">Figs 68, 69</figureCitation>
|
||
: arrows I,
|
||
<collectionCode id="ED0FEEA9FFCAFFFEB82CF941099102DC" box="[1143,1164,1721,1745]" pageId="10" pageNumber="33">II</collectionCode>
|
||
); dorsal lobe curved ventrad with short, acute end (
|
||
<figureCitation id="13256AE9FFCAFFFEB836F92209A202FF" box="[1133,1215,1754,1778]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">Fig. 68</figureCitation>
|
||
), in dorsal view evenly tapered towards acute apex (
|
||
<figureCitation id="13256AE9FFCAFFFEB8F3F90309E6031E" box="[1192,1275,1787,1811]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">Fig. 70</figureCitation>
|
||
); flagellum monstrous, strongly projecting ventrally, with strong, convex enlargement (
|
||
<figureCitation id="13256AE9FFCAFFFEB859F8C5099A0358" box="[1026,1159,1853,1877]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="10" pageNumber="33">Figs 68, 69</figureCitation>
|
||
: arrows
|
||
<collectionCode id="ED0FEEA9FFCAFFFEB8B0F8C508150358" box="[1259,1288,1853,1877]" pageId="10" pageNumber="33">III</collectionCode>
|
||
) at large, thick proximal winding, thinner and meandering towards end; lateral lobes small, narrow, projecting, each bearing apical group of long setae and proximal group of shorter setae (
|
||
<figureCitation id="13256AE9FFCCFFF8BC84FF090C7F0504" box="[223,354,241,265]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="12" pageNumber="35">Figs 68, 69</figureCitation>
|
||
); ventral lobe asymmetrical, divided in distodextral portion not much projecting ventrally and proximosinistral, transverse portion strongly triangularly projecting ventrally and laterally projecting from base of sinistral apical lobe (
|
||
<figureCitation id="13256AE9FFCCFFF8BD2CFE8D0CD40580" box="[375,457,373,397]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="12" pageNumber="35">Figs 33</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFCCFFF8BD8FFE8D0F020580" box="[468,543,373,397]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="12" pageNumber="35">68 – 70</figureCitation>
|
||
); transverse ridge of median foramen strongly, triangularly projecting ventrad (
|
||
<figureCitation id="13256AE9FFCCFFF8BCC4FE4F0DF105C2" box="[159,236,439,463]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="12" pageNumber="35">Fig. 33</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6126E4FFCBFFFFBCE6F88C085003BA" ID-DOI="http://doi.org/10.5281/zenodo.10724250" ID-Zenodo-Dep="10724250" httpUri="https://zenodo.org/record/10724250/files/figure.png" pageId="11" pageNumber="34" startId="11.[189,262,1908,1930]" targetBox="[279,1382,235,1877]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BA1766CFFCBFFFFBCE6F88C085003BA" blockId="11.[189,1436,1908,1976]" pageId="11" pageNumber="34">
|
||
<emphasis id="B96AAA7EFFCBFFFFBCE6F88C0C550384" bold="true" box="[189,328,1908,1930]" pageId="11" pageNumber="34">Figures 17–22.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBDF4F88D0F5F0384" authorityName="Frisch" authorityYear="2022" box="[431,578,1908,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="velifer">
|
||
<emphasis id="B96AAA7EFFCBFFFFBDF4F88D0F5F0384" box="[431,578,1908,1930]" italics="true" pageId="11" pageNumber="34">Scopaeus velifer</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Banten, Lebak: Citorek Kidul (
|
||
<emphasis id="B96AAA7EFFCBFFFFBFE1F88C0ECC0384" bold="true" box="[954,977,1908,1929]" pageId="11" pageNumber="34">17</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBFB8F88D097C0384" authorityName="Cameron" authorityYear="1930" box="[995,1121,1908,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="elegantulus">
|
||
<emphasis id="B96AAA7EFFCBFFFFBFB8F88D097C0384" box="[995,1121,1908,1930]" italics="true" pageId="11" pageNumber="34">S. elegantulus</emphasis>
|
||
</taxonomicName>
|
||
, Jawa Barat, Sukabumi: Djampang Kulon (
|
||
<emphasis id="B96AAA7EFFCBFFFFBD58F8730C0703AD" bold="true" box="[259,282,1931,1952]" pageId="11" pageNumber="34">18</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBD75F8730C8303AC" authorityName="Frisch" authorityYear="2022" box="[302,414,1931,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="uncinatus">
|
||
<emphasis id="B96AAA7EFFCBFFFFBD75F8730C2403AD" box="[302,313,1931,1952]" italics="true" pageId="11" pageNumber="34">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFCBFFFFBD1DF8740C8303AC" box="[326,414,1932,1953]" italics="true" pageId="11" pageNumber="34">uncinatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Jawa Barat, Sukabumi: Cidahu (
|
||
<emphasis id="B96AAA7EFFCBFFFFBF76F8730E5903AD" bold="true" box="[813,836,1931,1952]" pageId="11" pageNumber="34">19</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBF03F8730EF703AD" authorityName="Frisch" authorityYear="2022" box="[856,1002,1931,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="bipenicillatus">
|
||
<emphasis id="B96AAA7EFFCBFFFFBF03F8730E7E03AD" box="[856,867,1931,1952]" italics="true" pageId="11" pageNumber="34">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFCBFFFFBF34F8730EF703AD" box="[879,1002,1931,1952]" italics="true" pageId="11" pageNumber="34">bipenicillatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Sulawesi Tengah, Sigi: Pagana (
|
||
<emphasis id="B96AAA7EFFCBFFFFB923F873089203AD" bold="true" box="[1400,1423,1931,1952]" pageId="11" pageNumber="34">20</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBCE6F85A0C5A03BA" authorityName="Frisch" authorityYear="2022" box="[189,327,1954,1975]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="bipectenatus">
|
||
<emphasis id="B96AAA7EFFCBFFFFBCE6F85A0DD503BA" box="[189,200,1954,1975]" italics="true" pageId="11" pageNumber="34">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFCBFFFFBC8FF85A0C5A03BA" box="[212,327,1954,1975]" italics="true" pageId="11" pageNumber="34">bipectenatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Sulawesi Tengah: Lake Poso (
|
||
<emphasis id="B96AAA7EFFCBFFFFBEE6F85A0FC903BA" bold="true" box="[701,724,1954,1975]" pageId="11" pageNumber="34">21</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFCBFFFFBEBCF85A0E9603BA" authorityName="Frisch" authorityYear="2022" box="[743,907,1954,1975]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="crassipunctatus">
|
||
<emphasis id="B96AAA7EFFCBFFFFBEBCF85A0E9603BA" box="[743,907,1954,1975]" italics="true" pageId="11" pageNumber="34">S. crassipunctatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
|
||
<emphasis id="B96AAA7EFFCBFFFFB972F85A085D03BA" bold="true" box="[1321,1344,1954,1975]" pageId="11" pageNumber="34">22</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA1766CFFCCFFF8BCF5FE200C8B067E" blockId="12.[151,764,241,989]" pageId="12" pageNumber="35">
|
||
Female: Chamber of sperm pump subrectangular, truncate towards process segment, with long, distad widened apophysis (
|
||
<figureCitation id="13256AE9FFCCFFF8BD20FDE10CC7063C" box="[379,474,537,561]" captionStart="Figures 146–154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146–154. Sperm pump (146–154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150–152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="12" pageNumber="35">Fig. 154</figureCitation>
|
||
); bursal duct short, about twice as long as sperm pump, weakly sclerotized; bursa membranous (
|
||
<figureCitation id="13256AE9FFCCFFF8BD6AFDA30C97067E" box="[305,394,603,627]" captionStart="Figures 146–154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146–154. Sperm pump (146–154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150–152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="12" pageNumber="35">Fig. 153</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFCCFFF8BCF5FD840FFE06B8" blockId="12.[151,764,241,989]" pageId="12" pageNumber="35">
|
||
<emphasis id="B96AAA7EFFCCFFF8BCF5FD840C300699" bold="true" box="[174,301,636,660]" pageId="12" pageNumber="35">Phylogeny:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFCCFFF8BD6EFD840F770699" authorityName="Frisch" authorityYear="2022" box="[309,618,636,660]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="tortuosiflagellatus">
|
||
<emphasis id="B96AAA7EFFCCFFF8BD6EFD840F770699" box="[309,618,636,660]" italics="true" pageId="12" pageNumber="35">Scopaeus tortuosiflagellatus</emphasis>
|
||
</taxonomicName>
|
||
is a member of the
|
||
<taxonomicName id="4C1E0DEFFFCCFFF8BC80FD650C5706B8" authorityName="Oke" authorityYear="1933" box="[219,330,669,693]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96AAA7EFFCCFFF8BC80FD650C5706B8" box="[219,330,669,693]" italics="true" pageId="12" pageNumber="35">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="EF8F0B9DFFCCFFF8BDAAFD650FCA06B8" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[497,727,669,693]" pageId="12" pageNumber="35" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFCCFFF8BCF5FD460FE10715" pageId="12" pageNumber="35" type="distribution">
|
||
<paragraph id="8BA1766CFFCCFFF8BCF5FD460FE10715" blockId="12.[151,764,241,989]" pageId="12" pageNumber="35">
|
||
<emphasis id="B96AAA7EFFCCFFF8BCF5FD460C5906DB" bold="true" box="[174,324,702,726]" pageId="12" pageNumber="35">Distribution:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFCCFFF8BD37FD460FDC06DB" authorityName="Frisch" authorityYear="2022" box="[364,705,702,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="tortuosiflagellatus">
|
||
<emphasis id="B96AAA7EFFCCFFF8BD37FD460FDC06DB" box="[364,705,702,726]" italics="true" pageId="12" pageNumber="35">Scopaeus tortuosiflagellatus</emphasis>
|
||
</taxonomicName>
|
||
is distributed in
|
||
<collectingRegion id="49DAB88EFFCCFFF8BD6FFD270CB306FB" box="[308,430,735,758]" country="Indonesia" name="Jawa Barat" pageId="12" pageNumber="35">Jawa Barat</collectingRegion>
|
||
and was collected within and at the edge of the Mount Halimun – Salak National Park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFCCFFF8BCF5FCD90C4F07D0" pageId="12" pageNumber="35" type="etymology">
|
||
<paragraph id="8BA1766CFFCCFFF8BCF5FCD90C4F07D0" blockId="12.[151,764,241,989]" pageId="12" pageNumber="35">
|
||
<emphasis id="B96AAA7EFFCCFFF8BCF5FCD90C290734" bold="true" box="[174,308,801,825]" pageId="12" pageNumber="35">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFCCFFF8BDE4FCD80F990735" authorityName="Frisch" authorityYear="2022" box="[447,644,800,824]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="tortuosiflagellatus">
|
||
<emphasis id="B96AAA7EFFCCFFF8BDE4FCD80F990735" box="[447,644,800,824]" italics="true" pageId="12" pageNumber="35">tortuosiflagellatus</emphasis>
|
||
</taxonomicName>
|
||
[adjective, Latin, composed of the adjectives
|
||
<emphasis id="B96AAA7EFFCCFFF8BE79FCBA0F950754" box="[546,648,834,857]" italics="true" pageId="12" pageNumber="35">tortuosus</emphasis>
|
||
(tortuous, winding) and
|
||
<emphasis id="B96AAA7EFFCCFFF8BD1BFC9A0CAD0777" box="[320,432,866,890]" italics="true" pageId="12" pageNumber="35">flagellatus</emphasis>
|
||
(indicating a characteristic/ feature of the flagellum)] refers to the huge, remarkably tortuous flagellum of the aedeagus of this new species (
|
||
<figureCitation id="13256AE9FFCCFFF8BCC4FC3D0DED07D0" box="[159,240,965,989]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="12" pageNumber="35">Figs 33</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFCCFFF8BCA0FC3D0C5B07D0" box="[251,326,965,989]" captionStart="Figures 68–73" captionStartId="23.[189,262,1865,1887]" captionTargetBox="[261,1375,305,1707]" captionTargetId="figure-154@23.[189,1436,256,1845]" captionTargetPageId="23" captionText="Figures 68–73. Apex of aedeagus in lateral (68, 71), ventral (69, 72), dorsal view (70, 72) of Scopaeus tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (68–70); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (71–73). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, s – groups of setae, tr– transverse ridge, vl – ventral lobe, vld – dextral portion of ventral lobe, vls – sinistral portion of ventral lobe.Arrows: ventral spine of apical lobe (I, II), proximal enlargement of flagellum (III), ventral tooth of apical lobe (IV), tip of dorsal lobe (V, VI)." figureDoi="http://doi.org/10.5281/zenodo.10724272" httpUri="https://zenodo.org/record/10724272/files/figure.png" pageId="12" pageNumber="35">68 – 70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |