Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species
Author
Frisch, Johannes
Author
Narakusumo, Raden Pramesa
MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia
text
Soil Organisms
2022
2022-04-01
95
1
23
73
http://dx.doi.org/10.25674/so95iss1id311
journal article
10.25674/so95iss1id311
2509-9523
10724236
Scopaeus tortuosiflagellatus
Frisch
,
spec. nov.
(
Figs 8
,
33
,
68 –70
,
125
,
153, 154
)
Type specimens:
Indonesia
,
Jawa Barat
:
Holotype
♂
,
Sukabumi
,
Cisalimas
(
Cisalimas River
), (
06°45’21’’S
,
106°33’38’’E
),
870 m
,
19.9.2015
, leg.
Frisch
(
MZB
)
.
Paratypes
(
6 specimens
):
1 ♀
, same data as holotype (
MFNB
)
;
1 ♂
,
Bogor
, SW
Pamijahan
(
06°42’41’’S
,
106°41’03’’E
),
990 m
,
3.10.2015
, leg.
Frisch
(
MZB
)
;
1 ♂
,
1 ♀
,
Sukabumi
, NNW
Cidahu
:
Javana Spa
(
06°44’43’’S
,
106°42’51’’E
),
1150 m
,
30.9.2015
, leg.
Frisch
(
MFNB
,
MZB
)
;
2 ♀
,
Sukabumi
:
Cikaniki
(
Cikaniki River
), (
06°44’46’’S
,
106°32’25’’E
),
1020 m
,
5.10.2015
(
MFNB
,
MZB
)
.
Description:
Habitus and coloring as in
Fig. 8
. Head subquadrate. Antennae with elongate penultimate segment. Mesotibia slender. Forebody surface with fine, setose punctation, shiny; abdomen matter than forebody. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body black-brown except for narrow, yellow-brown, posterior margin of elytra, light brown tip of abdomen, medium brown maxillary palpi with darker penultimate segment, and yellow-brown legs; antennae with scapus medium brown, from blackish pedicellus gradually lighter towards terminal segments. Total body length
2.4 – 2.9 mm
; forebody length
1.5 – 1.6 mm
.
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with bisinuate emargination with strongly convex median third in about posterior fifth of sternite length (
Fig. 125
). Aedeagus (
Figs 33
,
68 – 70
) about
1.1 mm
long, with asymmetrical distal lobes; dextral apical lobe longer than acute sinistral apical lobe, enlarged apicad and with conspicuous, very slender, proximodorsad curved spine of ventral margin (
Figs 68, 69
: arrows I,
II
); dorsal lobe curved ventrad with short, acute end (
Fig. 68
), in dorsal view evenly tapered towards acute apex (
Fig. 70
); flagellum monstrous, strongly projecting ventrally, with strong, convex enlargement (
Figs 68, 69
: arrows
III
) at large, thick proximal winding, thinner and meandering towards end; lateral lobes small, narrow, projecting, each bearing apical group of long setae and proximal group of shorter setae (
Figs 68, 69
); ventral lobe asymmetrical, divided in distodextral portion not much projecting ventrally and proximosinistral, transverse portion strongly triangularly projecting ventrally and laterally projecting from base of sinistral apical lobe (
Figs 33
,
68 – 70
); transverse ridge of median foramen strongly, triangularly projecting ventrad (
Fig. 33
).
Figures 17–22.
Habitus of
Scopaeus velifer
, paratype, Banten, Lebak: Citorek Kidul (
17
);
S. elegantulus
, Jawa Barat, Sukabumi: Djampang Kulon (
18
);
S
.
uncinatus
, paratype, Jawa Barat, Sukabumi: Cidahu (
19
);
S
.
bipenicillatus
, paratype, Sulawesi Tengah, Sigi: Pagana (
20
);
S
.
bipectenatus
, paratype, Sulawesi Tengah: Lake Poso (
21
);
S. crassipunctatus
, paratype, Jawa Barat, Sukabumi: Sirnarasa (
22
).
Female: Chamber of sperm pump subrectangular, truncate towards process segment, with long, distad widened apophysis (
Fig. 154
); bursal duct short, about twice as long as sperm pump, weakly sclerotized; bursa membranous (
Fig. 153
).
Phylogeny:
Scopaeus tortuosiflagellatus
is a member of the
S. gracilis
species group (
Frisch et al. 2002: 39
).
Distribution:
Scopaeus tortuosiflagellatus
is distributed in
Jawa Barat
and was collected within and at the edge of the Mount Halimun – Salak National Park.
Etymology:
The epithet
tortuosiflagellatus
[adjective, Latin, composed of the adjectives
tortuosus
(tortuous, winding) and
flagellatus
(indicating a characteristic/ feature of the flagellum)] refers to the huge, remarkably tortuous flagellum of the aedeagus of this new species (
Figs 33
,
68 – 70
).