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<document id="A0193269EFB0E7FABC0779BBA60EFBA2" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="0A00E2F862305E64B88C05A18645C894" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Rishetia basseinensis Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha 2024, comb. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="288FF6D32E86607640CD2EBD9617C534">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:namePart id="17E68DED83A4A9850B08A3AD3F00B989">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="5EC9F44419431578CB5557424E1E9588">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="782499B6DBB2B8EC4C461A2D0701F768">Ablett, Jonathan D.</mods:namePart>
<mods:nameIdentifier id="3ACA270A2C57FB2FE85B036561DB3FAB" type="ORCID">0000-0002-7277-1934</mods:nameIdentifier>
<mods:affiliation id="EDEE470D069A76F754D9C2412FBF1B01">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
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<mods:namePart id="A67C4120C1754A058852ABD25A9F0051">Lwin, Ngwe</mods:namePart>
<mods:nameIdentifier id="7A3270169AC954C049EC14948C10C0D4" type="ORCID">0000-0002-4394-3741</mods:nameIdentifier>
<mods:affiliation id="C8EDE5C22138ABE147FBD20C2F42D469">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="A0975E07FD294961F1CD88382CFB36CF">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="25BBA5A18173BBE96F4AAACDF3B9E5CE" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="270EBC048B5467F45C042ECD2C9F8F93">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:namePart id="135A6A792EA0FA4D663615E73D94E758">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="6D4DFD59E1DCD86E72596ECAEBA8978C" type="ORCID">0000-0002-4431-2458</mods:nameIdentifier>
<mods:affiliation id="248CAD038776641620DBC0D2329EF50C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:typeOfResource id="4AE10AFE4AB34F779C930D05B4BA6625">text</mods:typeOfResource>
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<mods:date id="10E0DCE4418A2BAAEEE40AE796724402">2024</mods:date>
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<mods:number id="8DC9E7EAA9B69455975F160656252081">2024-07-30</mods:number>
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<treatment id="0A00E2F862305E64B88C05A18645C894" ID-DOI="http://doi.org/10.5281/zenodo.13137158" ID-Zenodo-Dep="13137158" ID-arpha="0A00E2F8-6230-5E64-B88C-05A18645C894" LSID="urn:lsid:plazi:treatment:0A00E2F862305E64B88C05A18645C894" httpUri="http://treatment.plazi.org/id/0A00E2F862305E64B88C05A18645C894">
<subSubSection id="6EDD6BA457E8DD53D0F0876D1942130F" type="nomenclature">
<paragraph id="F8B593547456EE31603CC61866BFB2AB">
<heading id="5CBEEB8119B239DA132188E55F37400B" reason="label">29</heading>
<taxonomicName id="6C4B738B88290FA1D9467E1DBF34A29F" authority="(Godwin-Austen, 1920)" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Subulinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="basseinensis" status="comb. nov.">
<emphasis id="65014505F5F759E2D8A98F4B920511DF" italics="true">Rishetia basseinensis</emphasis>
(
<bibRefCitation id="3ACF9F2B4C4F270F19F75B967431A141" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="812EA442BCAFE41D5B003E8B26208670" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3EE49097709A8A19F598AC067BF8171F" type="description">
<paragraph id="12412B31EC60EA43EB0315BBFD70A849">
<figureCitation id="49DA428F07D52FCF6BD1F6858437F62F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">Fig. 13 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7FC305AC6B374C60CACECD9836F2239" type="reference_group">
<paragraph id="B22383E7710F008B1A95031CD7F9C7AF">
<treatmentCitationGroup id="668CCFA07A39ACE38FB4CA5ADE83592F">
<treatmentCitation id="9C4A46C8123269154510713FBB49EB20" author="Blanford WT" page="79" year="1865">
<taxonomicName id="7B21806A2504B1394D1196557308D8A6" authority="Blanford, 1865: 79" authorityName="Blanford" authorityPageNumber="79" authorityYear="1865" class="Gastropoda" family="Achatinidae" genus="Achatina" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pertenuis" variety="major">
<emphasis id="A33F27DA2512454F8C5CB31CCD86FD99" italics="true">Achatina pertenuis var. major</emphasis>
<comment id="C6F900C62F7A01C425CA66A4D8B6790D">
<bibRefCitation id="EC9B564AE03DC78DB4ED4F47654C4004" author="Blanford WT" issue="2" journalOrPublisher="Journal of the Asiatic Society of Bengal" pagination="66-105" refId="B16" refString="Blanford WT (1865) Contributions to Indian Malacology, No. 5. Description of new land shells from Arakan, Pegu, and Ava; with notes on the distributions of described species. Journal of the Asiatic Society of Bengal 34 (2): 66 - 105. https: // www. biodiversitylibrary. org / item / 114408" title="Contributions to Indian Malacology, No. 5. Description of new land shells from Arakan, Pegu, and Ava; with notes on the distributions of described species." url="https://www.biodiversitylibrary.org/item/114408" volume="34" year="1865">Blanford, 1865: 79</bibRefCitation>
[in part].
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="5CC2D0EB2B5F112FE800D7CCEE390E36">
<treatmentCitationGroup id="D9354CFEBDA12FEC122E4DBCBCA5665E">
<taxonomicName id="850C52300AF033BB5CBD0FBAF694EA57" ID-CoL="7RFHW" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="basseinensis">
<emphasis id="7B7B8B803D18E10751370842EB587F3A" italics="true">Glessula basseinensis</emphasis>
</taxonomicName>
<comment id="C43C656907769C12B631845646C62586">
<treatmentCitation id="9AF9886BCEE7E26969D7A2C5DBA38312" author="Godwin-Austen HH" page="54" year="1920">
<bibRefCitation id="18B459B390B8A6C742F766F3D45E1D8F" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="54" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen 1920: 54</bibRefCitation>
</treatmentCitation>
, pl. 161. fig. 3, pl. 164, fig. 12. Type locality: Bassein,
<collectingRegion id="E2DE490FDEFC7802D92AB491D1E52DC2" country="Myanmar" name="Bago">Pegu</collectingRegion>
and Pyema Khyoung, Bassein.
</comment>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E15DK" type="Type specimens">
<paragraph id="4FE075724F080568BA9E9A2CFB58792E">
<heading id="4442C3CB36AB067C11F94B1499EA6626" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="95A42BBEC5410D33929DB2354A125F61">
<materialsCitation id="87691283914D0EFC9377010771C98308" collectionCode="NHMUK" collectorName="Blanford" location="Bassein" specimenCode="1909.3. 15.19" specimenCount="1" stateProvince="Pegu" typeStatus="syntype">
<emphasis id="896CB7FC71F00259AF61A795A7C4DEB5" bold="true" italics="true">
<typeStatus id="BC1DFB74C4A2A913EDD92EA0992C1BE2" type="syntype">Syntypes</typeStatus>
</emphasis>
<named-content id="849DF10234C87475B0566398A5126979" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="8F114980FEE3A9E5B905816DD0A9179B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
<specimenCode id="2C9A30D0EFCCB8A814A26359B2F56020">1909.3. 15.19</specimenCode>
[re-registered in error as 1985139] (3 shells; Fig.
<figureCitation id="B6BD148324FC8F7876AD73A17B848DE6" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 E, F</figureCitation>
) ex.
<collectorName id="07DD53D116AED9C261F4DEC209FB12DF">Blanford</collectorName>
ex. Godwin-Austen collection from
<location id="4550A633D652DD368A47DF402F08BF5A" LSID="urn:lsid:plazi:treatment:0A00E2F862305E64B88C05A18645C894:4550A633D652DD368A47DF402F08BF5A" name="Bassein" stateProvince="Pegu">Bassein</location>
,
<collectingRegion id="6ACF7D3CB876454C7269110B63EF488A" country="Myanmar" name="Bago">Pegu</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EO6DK" type="diagnosis">
<paragraph id="640E95BA67FF1B81568167CC80828C70">
<heading id="5EEB3204A53D87546DBB1F15999608CF" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="5A47F0C0C3B58B6CA1832E3B3EE565C7">Shell slender, elongate turreted and regularly attenuated; apex rounded and blunt; subsequent whorls with fine and equally spaced radial ribs, which more prominent near suture. Suture impressed and whorls flattened convex. Aperture ovate; columella strong, concave, and truncated.</paragraph>
</subSubSection>
<subSubSection id="SECID0ET6DK" type="distribution">
<paragraph id="A89D7657AF1C6BDB45BCBB016DFBCDEF">
<heading id="FD52DAFC93258E9009E814090E505A8A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="9A13F80654B3165C094A848C242B7CB1">
<taxonomicName id="BB117092DD5BEDC15929C63E1B287D72" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="basseinensis">
<emphasis id="C8220DBA4DA6339989CCF75F95436C4C" italics="true">Rishetia basseinensis</emphasis>
</taxonomicName>
appears to be restricted within
<collectingCountry id="66128810124A55EAAEC87969B4974952" name="Myanmar">Myanmar</collectingCountry>
and is known only from the type locality. Based on the current administrative boundary, Bassien (or Pathein), Pegus is in the Pathein Township, Pathein District,
<collectingRegion id="A8934CDF1D171B9019D0932DBCEA6665" country="Myanmar" name="Ayeyawady">Ayeyarwady Region</collectingRegion>
, rather than
<collectingRegion id="4F0CA9D70A29D8ACB8482091BE038205" country="Myanmar" name="Bago">Pegu</collectingRegion>
[
<collectingRegion id="4EFDB0C0D59EF332A5A52E178923CB9B" country="Myanmar" name="Bago">Bago Region</collectingRegion>
].
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEAEK" type="remarks">
<paragraph id="3EDB49FD2E8D14D6953FDDCF12735D1E">
<heading id="B394B2C087ECF51BCDC3D78CEBDF012D" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="DC2FDBF722FB32F9C188FD1426782A0C">
<bibRefCitation id="C6D18F55FEC30306BA697D16EE49E3F6" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="53" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 53)</bibRefCitation>
stated that when examining the type series of
<taxonomicName id="0E7179472BB702E500D0E8380BB3583C" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="variety" species="pertenuis" variety="major">
<emphasis id="F32685EFE7A77D728A33A99E0AA321FC" italics="true">Glessula pertenuis var. major</emphasis>
</taxonomicName>
ex. Blanford collection, he found it very different from the nominotypical taxon, and described this specimen lot under a different name,
<taxonomicName id="BA305B76F4C55F76CFA2E42D0FD304AD" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="basseinensis">
<emphasis id="A1D965BD7106A577E20F77DC3DD62EB6" italics="true">Glessula basseinensis</emphasis>
</taxonomicName>
. In the original description,
<bibRefCitation id="D0BCB27FB0D3FED60FB6FC6049ACF8CB" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" page="54" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920: 54)</bibRefCitation>
provided two collection localities, namely Bassein, Pegu and Pyema Khyoung, Bassein and comparing three and
<specimenCount id="CBBB8B5D196CA21E67C925862A6D2C17" count="6" type="generic">six specimens</specimenCount>
, respectively. The description clearly states a catalogue number Type No. 19.9.3.15
<abbrev content-type="institution" id="ABBRID0EOBEK" xlink_title="used in old labels for the British Museum">
<collectionCode id="593A5BE0CC247577454467994C6D6558">B</collectionCode>
. M
</abbrev>
. and includes an illustration of specimens from Bassein, Pegu. The
<named-content id="B7455E65216B283F20D5D89BC9425081" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="839B2A3D5EFD9F41B9FB20081A620191" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
collection contains this specimen lot
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</named-content>
1909.3. 15.19 consisting of three shells with an original label in Godwin-Austens handwriting stating Type from the H. F. Blanford collection. The specimen with a similar shell measurement given in the original description and with pink wool inside the shell is figured herein (Fig.
<figureCitation id="4EC1A1089DD203EC7107A1B72A185C74" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 E</figureCitation>
). Additionally, one further shell from the same
<typeStatus id="E3D14FC573F32219F4CFC05250AA0A48" type="syntype">syntype</typeStatus>
lot is also illustrated (Fig.
<figureCitation id="0F05BD7C8EF265662E51640CAD89AA29" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A C Rishetia akouktoungensis, syntypes NHMUK 1906.1. 1.2207 from Akouktoung, Pegu D Rishetia baculina, syntypes NHMUK 1909.3. 15.9 from Kursiong, Darjiling E, F Rishetia basseinensis, syntypes NHMUK 1909.3. 15.19 from Bassein G I Rishetia burrailensis maxwelli G holotype NHMUK 1903.7. 1.1717 / 1 from Naga Hills, East of Kohima H specimen NHMUK 1988147 from Somra Tracts, Somra Khulen Post, west of Kyengdwen River, Burma and I paratypes NHMUK 1903.7. 1.1717 / 24 from Naga Hills, East of Kohima." figureDoi="10.3897/zookeys.1208.116083.figure13" httpUri="https://binary.pensoft.net/fig/1104511">13 F</figureCitation>
).
</paragraph>
<paragraph id="1A6C21B346DC1D797699909D03B5914B">
The specimen lot
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</named-content>
1909.1. 1.2208 is excluded from the type series of this species (see also under the
<taxonomicName id="723A5242BFCC449DC74F9AAF355B89F1" authorityName="Man &amp; Ablett &amp; Lwin &amp; Sutcharit &amp; Panha" authorityYear="2024" baseAuthorityName="Blanford" baseAuthorityYear="1865" class="Gastropoda" family="Achatinidae" genus="Rishetia" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subSpecies" species="pertenuis" subSpecies="major">
<emphasis id="468814803E25355D9025FEBB79B8707E" italics="true">Rishetia pertenuis major</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>