treatments-xml/data/C8/52/BA/C852BA0F4594C0A61091B97ECAC9DB42.xml
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<mods:title id="E384B561549DCA5D1EF0269E8027C4C1">A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae).</mods:title>
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<mods:namePart id="164BAFA5D1998AE5B45ADF4D0D0EC691">Eguchi, K.</mods:namePart>
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<treatment id="C852BA0F4594C0A61091B97ECAC9DB42" ID-DOI="http://doi.org/10.5281/zenodo.6230490" ID-GBIF-Taxon="100133931" ID-Zenodo-Dep="6230490" LSID="urn:lsid:plazi:treatment:C852BA0F4594C0A61091B97ECAC9DB42" httpUri="http://treatment.plazi.org/id/C852BA0F4594C0A61091B97ECAC9DB42" lastPageNumber="77" pageNumber="73">
<subSubSection id="77A694A6E8F3F4C17A2AA839D0F782D0" type="nomenclature">
<paragraph id="FCC0FFA1DF3E86E40F345F0BFD1A79A8" pageNumber="73">
<taxonomicName id="D6A09C1AF0CBD612B500FA791BEAA3E8" ID-CoL="4FYJ4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria F. Smith</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0B2F3DB0623CFA6ED879373519F15E8A" type="description">
<paragraph id="5A085A54A3EBA3F0121CA7F7E449376B" pageNumber="73">Figs. 20a-g, 32e, 32f</paragraph>
</subSubSection>
<subSubSection id="43A97929D115AE6D66C4F2ACC0232899" type="reference_group">
<paragraph id="3ECAFB92A61EA3D8927FE1B56CE5F088" pageNumber="73">
<taxonomicName id="01E58DDAB576EF2056FE580930B89561" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria F. Smith</taxonomicName>
, 1860: 112. Eguchi, Bui et al. 2005: 90 (checklist). Syntypes: 3 majors &amp; 3 minors, &quot;Bac&quot; [Batjan, Indonesia], OXUM TYPE HYM: 983 1-2/6, 3-4/6 &amp; 5-6/6, examined.
</paragraph>
<paragraph id="4C8B897CD9BCDF8A9197042010BDA00D" pageNumber="73">
<taxonomicName id="6DFE105C327610AFF782F4B7CA0A284E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142504" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole divergens Mayr" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="divergens">Pheidole divergens Mayr</taxonomicName>
, 1867: 97. Mayr, 1879: 675 (junior synonym of P
<taxonomicName id="6E3D143C56D925C8A66951B5A2FA95BD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="plagiaria">plagiaria</taxonomicName>
). Syntypes: major, minor, queen &amp; male, Indonesia, not examined.
</paragraph>
<paragraph id="5CDF5F0FD2375620F2C1718BA4BEBFDE" pageNumber="73">
<taxonomicName id="54298C96E3EB2C0BBCDB365992DD7F4A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151468" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole peguensis yomensis Forel" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="peguensis" subSpecies="yomensis">Pheidole peguensis r. yomensis Forel</taxonomicName>
, 1903: 253.
<taxonomicNameLabel id="A551F25E47B1437F619E4FF205A71CEA">Syn.n.</taxonomicNameLabel>
Syntype: 1 major, &quot;Lower Burma Pegu Yoma 1 1900 Bingham&quot; [Myanmer], MHNG, examined, designated as lectotype [Fig. 32e, 32f].
</paragraph>
<paragraph id="E2FBB3040B77153C9C023FB44CA44C4E" pageNumber="73">
Current subspecies (Bolton, 1995): nominal plus
<taxonomicName id="BEE679D9B79E1B86EC303B29FDEB9E0F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151477" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria moica Forel" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="moica">moica Forel</taxonomicName>
, 1911c: 222, syntype(s): major, Central part of Southern Vietnam, not examined;
<taxonomicName id="EFCC9BD04EB6DD7459684A8D57799748" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151479" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria palawanica Stitz" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="palawanica">palawanica Stitz</taxonomicName>
, 1925: 118, syntypes: 2 majors, &quot;N. Palawan Binaluan Boettcher&quot; [Philippines], ZMHB, examined;
<taxonomicName id="947E9AC9E9A4ADD7A3884018D770DB28" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151480" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria rectilineata Viehmeyer" order="Hymenoptera" pageNumber="73" phylum="Arthropoda" rank="species" species="rectilineata">rectilineata Viehmeyer</taxonomicName>
, 1916b: 288-289, syntype(s): major, Sulawesi (in copal), not examined.
</paragraph>
</subSubSection>
<subSubSection id="5C4EDD9D181ED7482982B3FFA68DA09A" type="materials_examined">
<paragraph id="B39791B19830397CFD65CA04CE979AA6" pageNumber="74">
<pageStartToken id="1A0ECDC9976B6464B127E8299CF15E87" pageNumber="74">Other</pageStartToken>
material examined: S. China: Hainan: Jianling N.R., Wanling [J. Fellowes]. Vietnam: Thai Nguyen: My Yen Commune Forest, 21°35'N, 135°36'E, Na Hau Village [Eg01-VN-147]; Vinh Phuc: Tam Dao N.P., 21°27'N, 105°38'E, 800-900 m alt. [Eg99-VN-037, -038], Tam Dao N.P., 900 m alt. [Eg99-VN- 001, -005, -008, -009; Eg01-VN-104, -106], Tam Dao N.P., 900-1100 m alt. [Eg99-VN-052], Tam Dao N.P., 1100 m alt. [VN98-SKY-14]; Ha Tay (mislabeled as &quot;Ha Tai&quot;): Ba Vi N.P., 400-800 m alt. [Eg99-VN-119]. Thailand: Chiang Mai: Doi Ang Khang HQ, 1300 m alt. [TH98-SKY-28], Doi Chiang Dao [Eg01-TH-122, - 134, -148], Doi Chiang Dao, 500-600 m alt. [TH98-SKY-18], Doi Suthep-Pui N.P., 600 m alt. [TH98-SKY- 04], Doi Suthep-Pui N.P., 800 m alt. [TH98-SKY-08], Doi Suthep-Pui N.P., 800-900 m alt. [Eg01-TH-069, - 096, -098, -100], Doi Suthep-Pui N.P., 900 m alt. [Eg01-TH-082, -083, -086, -087, -089, -090], Doi SuthepPui N.P., 1200 m alt. (Doi Pui) [Eg01-TH-110, -118]; Nakhonratchasima: Khao Yai N.P. [TH00-SKY-24], nr. Sakaerat Environmental Research Centre [Eg99-TH-020; Eg01-TH-532, -549]; Chanthabri: Khao Soi Dao [Eg01-TH-021]; Chacheongsao: Lumchangwat Station, Khao Ang Reu Nai W.S. [TH03-SKY-69]; Pattani: Soi Khao [TH98-SKY-40]; Trang: Khao Chong Waterfall [Eg01-TH-708, -714, -723, -736, -745, -746, -758]; Songkhla: Ton Nga Chang, nr. Hat Yai [Eg01-TH-613, -615, -626, -628], Songkhla Univ. (Khao Kor Hong) [Eg01-TH-597, -598]; Narathiwat: Bala area of Bala-Hala W.S. [Eg01-TH-636]. Myanmar: Chin: between Kampetlet and Natmataung N.P., 1670 m alt. [MM02-SKY-46], between Kampetlet and Natmataung N.P., 1700 m alt. [MM02-SKY-02], between Kampetlet and Natmataung N.P., 1760 m alt. [MM02-SKY-34], Natmataung N.P., 2000 m alt. [MM02-SKY-10], Natmataung N.P., 21°13'N, 93°56'E, 2000 m alt. [MM02-SKY- 24, -26, -29]. W. Malaysia: Selangor: Ulu Gombak [FI92MG-174, -383]. E. Malaysia: Sabah: Deramakot Forest Reserve [C. Briihl], Gunong Rara [Eg96-BOR-351, -363; Eg97-BOR-526, -534, -556, -566], Poring, 450- 500 m alt. [Eg96-BOR-271], Poring, 500-550 m alt. [Eg96-BOR-294], Poring, 600 m alt., [T. Kikuta's colony: 8-A, 43A, 731, 6XI2306S1-4], Sepilok Forest [Eg97-BOR-412, -418, -460, -464; Eg98-BOR-871], Tawau Hills Park [Sk. Yamane]; Sarawak: Lambir Hills N.P. [Eg98-BOR-803, -808, -817, -819, -825, -829], Lambir Hills N.P. (Inoue Trail) [SR04-SKY-67]. Brunei: Belalong Forest Section [Eg99-BOR-211], Merimbun Heritage Park (=Tasek Merimbun) [Eg99-BOR-007, -008, -028, -030, -035, -070, -074, -123, -150, -552, -567, -568; Eg00-BOR-019, -022, -064]. Indonesia: E. Kalimantan: Sangkimah, Kutai N.P. [Sk. Yamane]; W. Sumatra: Sukarami, nr. Padang [FI92-66, -83; FI96-162; M. Kawamura's colony: 9/29c (1999)]; Krakatau Is.: P. Rakata Besar [Kagoshima Univ. Krakatau Exp.]; W. Java: Cibodas [M. Kawamura's colony: D28 (1997)], Kebun Raya, Bogor [FI95-534, -784; Sk. Yamane; M. Kawamura's colony: 154 (1998), 9/26a (1999), 10/5b (1999), 10/5c (1999)], G. Halimun [FI98-373; M. Kawamura's colony: 5/6a (1998)], Pangandaran [FI95-705], Ciater, 1350 m alt. [Syaukani]; C. Java: Borobudur, nr. Yogyakarta [Sk. Yamane], Kaliadem, 800-1000 m alt., G. Merapi [JV02/03-SKY-38]; E. Java: Sumberbrantas, 1600 m alt., Tulungrejo, Batu [JV02/03-SKY-78, -79]; Bali: Kebun Raya [FI94-137]. Philippines: Leyte: Baybay, Mt. Pangasugan, Calbiga-a River [Zettel].
</paragraph>
</subSubSection>
<subSubSection id="297C72DB3E7392A12066E4FE2C6629E2" type="description">
<paragraph id="DB68176419C4F1899C3B7ED0D160AC97" pageNumber="74">Worker measurement &amp; indices: Major (n=5). - HL 1.65-1.89 mm, HW 1.59-1.77 mm, CI 91-98, SL 1.03-1.22 mm, SI 59-71, FL 1.42-1.70 mm, FI 85-99.</paragraph>
<paragraph id="7BBFD63911778F5FC03B3C351651C949" pageNumber="74">Minor (n=5). - HL 0.79-0.93 mm, HW 0.67-0.75 mm, CI 81-85, SL 1.01-1.20 mm, SI 146-163, FL 1.12-1.36 mm, FI 162-181.</paragraph>
<paragraph id="878BE0AE10997FE4BB18A3D60C568289" pageNumber="74">Worker description</paragraph>
<paragraph id="020694666F409B123BD3037395C32FF1" lastPageNumber="76" pageNumber="74">Major. - Head in lateral view not or only very weakly impressed on vertex; posterior margin of head in full-face view relatively widely and deeply concave; longitudinal rugulae running on the frons and then spreading posterolaterad as they run almost transversely on the posteriormost part of the dorsum of vertexal lobe; posterolateral face of head with oblique rugulae which run anteroventrad from dorsal face of vertexal lobe; frontal carina conspicuous; antennal scrobe inconspicuous or only weakly impressed; median longitudinal carina of clypeus present but weak, or rarely absent; median and submedian processes of hypostoma inconspicuous or absent; lateral processes conspicuous; outer surface of mandible covered with relatively long decumbent hairs; antenna with a 3-segmented club; maximal diameter of eye a little longer, as long as, or a little shorter than antennal segment X. Promesonotal dome in dorsal view transversely rugose with interspaces smooth to shagreened, or smooth to shagreened with sparse several transverse rugulae; the dome in lateral view with a low prominence or mound on its posterior slope; humerus of the dome not or hardly produced laterad; the dome at the humeri narrower than at the bottom; mesopleuron, metapleuron and lateral face of propodeum rugoso-reticulate, with enclosures punctured. Outer surface of foretibia bearing relatively long decumbent-suberect hairs. Petiole longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite largely smooth and shining, but sometimes weakly punctured around its articulation with postpetiole.</paragraph>
<caption id="BD34A7FE917F6C2986700C199AEE3307">
<paragraph id="82CE8C94CFD1564968089BEA17B3B58E" pageNumber="75">
<pageStartToken id="8A12C1478878A9CDB981434CA8B206E5" pageNumber="75">FIGURE</pageStartToken>
20a-d,
<taxonomicName id="52605F4AD8FA5C1F07DECA67CC0FB486" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="75" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria</taxonomicName>
, major [Eg99-VN-037] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
</paragraph>
</caption>
<caption id="F936500559A98A08CAFDB73357E5A7BA">
<paragraph id="BD08B85707888E1AB78E338FC79FDACC" pageNumber="76">
<pageStartToken id="926FA56763B556E9DC4B09C9A2145327" pageNumber="76">FIGURE</pageStartToken>
20e-g,
<taxonomicName id="628931989E2C56A7C025CAA595DCB1DC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="76" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria</taxonomicName>
, minor [Eg99-VN-037] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
</paragraph>
</caption>
<paragraph id="2C0082355C71BEA67963BC2B98FC9EB5" pageNumber="77">Minor. - Frons and dorsal face of vertex smooth and shining, or very weakly punctured or rugoso-punctate; area between antennal insertion and eye rugose or rugoso-punctate; preoccipital carina well-developed; median part of clypeus with a zigzag or ramified (but rarely straight) median carina which is usually accompanied with weak rugulae; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than antennal segment X. Promesonotal dome smooth and shining, or in dorsal view concentrically rugose, with a median area smooth and shining or very weakly punctured; the posterior slope of the dome with a low mound which bears several (&gt;2) standing hairs; humerus in dorsooblique view not or hardly produced; mesopleuron, metapleuron and lateral face of propodeum punctured, usually overlain by weak rugoso-reticulation; propodeal spine horn-like or elongate-triangular, narrowly based, directing relatively upward. Petiole longer than postpetiole (excluding helcium); postpetiole not massive.</paragraph>
<paragraph id="8755CF442FE1AA5F1C31CA21A131325E" pageNumber="77">
Recognition: The syntype major of
<taxonomicName id="490720951BBE9129DB83208E0D7F0F34" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151468" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole peguensis yomensis Forel" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="peguensis" subSpecies="yomensis">Pheidole peguensis r. yomensis Forel</taxonomicName>
agrees with the majors of Thai populations of
<taxonomicName id="A58333217C3E37A7EC70F3BD4F4DEF51" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria</taxonomicName>
(e.g., Eg01-TH-083). Thus, I here synonymize
<taxonomicName id="C9A190FE5E1E5CD164593B9DEAEFD62E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151468" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole peguensis yomensis Forel" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="subSpecies" species="peguensis" subSpecies="yomensis">Pheidole peguensis r. yomensis</taxonomicName>
with
<taxonomicName id="B517A1642BEF9552F15D934BA88AEA2F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="species" species="plagiaria">P. plagiaria</taxonomicName>
. Three
<taxonomicName id="D61EFEFC7BCAFE7AC6745576FFE70440" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
minors which are pinned together and labeled as &quot;Ph
<taxonomicName id="1B4F51920286D5C81045D293643DA286" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="species" species="plagiaria">plagiaria Sm</taxonomicName>
<taxonomicName id="70AE2812F155420687AFBDE454ED2B26" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151477" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria moica Forel" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="variety" species="plagiaria" variety="moica">v. moica For</taxonomicName>
.... Mois Cochinchine (Dugas)&quot; were examined (MHNG). The top of the pin is undoubtedly a minor of
<taxonomicName id="DCEF8FD742E3DAF7F7042E555839AC1E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="species" species="plagiaria">Pheidole plagiaria</taxonomicName>
, while the middle is the other
<taxonomicName id="B3E96657BCD9BC9BC6F9D03144E08534" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
species, and the bottom is
<taxonomicName id="9346A21DAAE13308E84D24A07FBFA8F6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
sp. Forel (1911) described &quot;
<taxonomicName id="1E5A101498194BD7AF1DEC06125D61D0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151477" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria moica Forel" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="variety" species="plagiaria" variety="moica">P. plagiaria var. moica</taxonomicName>
&quot; based on the major subcaste, and so I refrain from resolving the synonymy.
</paragraph>
</subSubSection>
<subSubSection id="2DF8DA46D03FDEFBAEE2624793125E7A" type="discussion">
<paragraph id="6B09AC568FB6BA10168EE3407A2C56EB" pageNumber="77">
This species is characterized among Indo-Chinese species by the combination of the following features: in the major posterior margin of head in full-face view relatively widely and deeply concave; in the major the posteriormost part of the dorsum of vertexal lobe almost transversely rugose; in the major and minor the posterior slope of promosonotal dome with a prominence or mound; in the major outer surface of foretibia bearing relatively long decumbent-suberect hairs; in the minor median part of clypeus with a zigzag or ramified (but rarely straight) median carina which is usually accompanied with weak rugulae. The minor of this species is similar to that of
<taxonomicName id="835E2CC380FD6113A240E8B2E6044A3C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33623" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole binghamii Forel" order="Hymenoptera" pageNumber="77" phylum="Arthropoda" rank="species" species="binghami">P. binghami</taxonomicName>
, but in the latter median part of clypeus has no rugula and the mound on the posterior slope of promesonotal dome has only a pair of hairs.
</paragraph>
</subSubSection>
<subSubSection id="BCEF5B82D45D6C2765EDEA8015A5993E" type="distribution">
<paragraph id="A9E045A940A8F923FD26C04D6A988977" pageNumber="77">Distribution &amp; bionomics: Widely distributed in the Indo-Chinese, Indo-Malayan subregions and Austro-Malayan subregion. Indo-Malayan populations inhabit well-developed forests, and usually nest in rotting logs and wood fragments. On the other hand, Indo-Chinese populations often occur in forest edges and even in open lands adjacent to forests, and usually nest in the soil (Eguchi 2001).</paragraph>
</subSubSection>
</treatment>
</document>