1155 lines
151 KiB
XML
1155 lines
151 KiB
XML
<document id="6CECC9743127AB21AEC33AC2F01E5B7F" ID-CLB-Dataset="3875" ID-DOI="10.11646/zootaxa.5057.4.3" ID-GBIF-Dataset="168685a6-5bad-4fe1-9a63-ce464dfec698" ID-ISSN="1175-5326" ID-Zenodo-Dep="5598859" ID-ZooBank="A3C378BB-58ED-4A18-8F52-1D00E58E6F1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635231831341" checkinUser="plazi" docAuthor="Munir, Misbahul, Nishikawa, Kanto, Hamidy, Amir & Smith, Eric N." docDate="2021" docId="C83EE36BC448FD68FF192A53600AFD76" docLanguage="en" docName="zootaxa.5057.4.3.pdf" docOrigin="Zootaxa 5057 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megophrys acehensis Munir & Nishikawa & Hamidy & Smith 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="520" masterDocId="34079B13C443FD79FF8E2C58636FFFDF" masterDocTitle="Two new species of Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from Sumatra, Indonesia" masterLastPageNumber="529" masterPageNumber="503" pageNumber="514" updateTime="1699464381098" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5E152FEB49915238664B196549D322ED">
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<mods:title id="289E59481BE2059F2F287A761002CF05">Two new species of Megophrys Kuhl and Van Hasselt (Amphibia: Megophryidae) from Sumatra, Indonesia</mods:title>
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<mods:namePart id="7360F5F7D9E127958362A096195C8861">Munir, Misbahul</mods:namePart>
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<mods:affiliation id="1DB30EBE155E63171DD33624E4381200">Graduate School of Global Environmental Studies, Kyoto University, Yoshida-Honmachi, Sakyo-ku, Kyoto, 606 - 8501, JAPAN & misbahul. munir. 87 a @ st. kyoto-u. ac. jp; munir _ 1912 @ outlook. com; https: // orcid. org / 0000 - 0001 - 7115 - 6625</mods:affiliation>
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<mods:nameIdentifier id="EC012CC5482F7B00E6C9FAADDE74EE7B" type="email">misbahul.munir.87a@st.kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="3FCB489A42CC64EF2B228AA6520AE683">Nishikawa, Kanto</mods:namePart>
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<mods:nameIdentifier id="E4129BA0576E7E46AA35C30A4D2C16C8" type="ORCID">0000-0002-6274-4959</mods:nameIdentifier>
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<mods:affiliation id="078E03130AFEAD03240220DCC70FB650">Graduate School of Global Environmental Studies, Kyoto University, Yoshida-Honmachi, Sakyo-ku, Kyoto, 606 - 8501, JAPAN & Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-Nihonmatsu-cho, Sakyo-ku, Kyoto 606 - 8501, JAPAN nishikawa. kanto. 8 v @ kyoto-u. ac. jp; http: // orcid. org / 0000 - 0002 - 6274 - 4959</mods:affiliation>
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<mods:nameIdentifier id="B21C4C399C5A31070B042E867771A60C" type="email">nishikawa.kanto.8v@kyoto-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="E1DCFE243E34961E825CC8687DB16068">Hamidy, Amir</mods:namePart>
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<mods:nameIdentifier id="A5608023D180AD3F9A4F08273C510763" type="ORCID">0000-0001-6426-7458</mods:nameIdentifier>
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<mods:affiliation id="876928B27313428EA64C13F2FE4B229F">Museum Zoologicum Bogoriense, Research Centre for Biology, National Research and Innovation Agency of Indonesia, Gd. Widyasatwaloka, Jl. Raya Jakarta-Bogor Km 46, Cibinong, West Java, INDONESIA & hamidyamir @ gmail. com; http: // orcid. org / 0000 - 0001 - 6426 - 7458</mods:affiliation>
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<mods:nameIdentifier id="17E0294251BED57D58F7ABF818C194B8" type="email">hamidyamir@gmail.com</mods:nameIdentifier>
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<mods:namePart id="4A3A666212EF047EB937795F7E0A30DD">Smith, Eric N.</mods:namePart>
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<mods:nameIdentifier id="85528DEF6842CAC749E1999F6414AF95" type="ORCID">0000-0002-4257-741X</mods:nameIdentifier>
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<mods:affiliation id="1ED418D0DDB4318D568FD72F57FA7BA2">The Amphibian and Reptile Diversity Research Center and Department of Biology, University of Texas at Arlington, Arlington, TX 76010, USA & e. smith @ uta. edu; https: // orcid. org / 0000 - 0002 - 4257 - 741 X</mods:affiliation>
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<mods:nameIdentifier id="95D88F027A6BAADAB567A8852914C0D7" type="email">e.smith@uta.edu</mods:nameIdentifier>
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<mods:title id="8E690E5B3B7894F7AC1464B66C3E2541">Zootaxa</mods:title>
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<mods:date id="9D3041BB9A63386D1317CCFAB9ADAEEE">2021</mods:date>
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<treatment id="C83EE36BC448FD68FF192A53600AFD76" ID-DOI="http://doi.org/10.5281/zenodo.5633955" ID-GBIF-Taxon="189461797" ID-Zenodo-Dep="5633955" LSID="urn:lsid:plazi:treatment:C83EE36BC448FD68FF192A53600AFD76" httpUri="http://treatment.plazi.org/id/C83EE36BC448FD68FF192A53600AFD76" lastPageId="17" lastPageNumber="520" pageId="11" pageNumber="514">
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<taxonomicName id="879729FEC448FD72FF192A5362F4F9F9" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[151,411,1547,1574]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="11" pageNumber="514" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
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<emphasis id="72E38E6FC448FD72FF192A5362F4F9F9" bold="true" box="[151,411,1547,1574]" italics="true" pageId="11" pageNumber="514">Megophrys acehensis</emphasis>
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<taxonomicNameLabel id="69D03314C448FD72FE2C2A54616FF9F9" box="[418,512,1548,1574]" pageId="11" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="088D01F6C448FD72FF192A6862EFF994" box="[151,384,1584,1611]" pageId="11" pageNumber="514" type="description">
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<paragraph id="4028527DC448FD72FF192A6862EFF994" blockId="11.[151,512,1547,1611]" box="[151,384,1584,1611]" pageId="11" pageNumber="514">
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(
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<figureCitation id="D8AC4EF8C448FD72FF112A686382F995" box="[159,237,1584,1610]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="11" pageNumber="514">Figs. 4</figureCitation>
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A−D, 5A−E)
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</paragraph>
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<subSubSection id="088D01F6C448FD72FF192A206642F8B4" pageId="11" pageNumber="514" type="materials_examined">
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<materialsCitation id="F0FF5820C448FD72FF192A2061FFF920" ID-GBIF-Occurrence="3393350303" accessionNumber="ENS 18664, MT710708" collectingDate="2015-08-05" collectionCode="MZB, MT" county="Amph" elevation="1638" latitude="4.506444" location="Linge District" longLatPrecision="1" longitude="96.86075" municipality="Tengah Regency" pageId="11" pageNumber="514" specimenCount="1" stateProvince="Aceh Province, Aceh" typeStatus="holotype">
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<typeStatus id="9F2CECDFC448FD72FF192A20626BF94D" box="[151,260,1656,1682]" pageId="11" pageNumber="514" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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26098 (field number ENS 18664; GenBank accession no.
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710708
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;
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<figureCitation id="D8AC4EF8C448FD72FB312A206663F94D" box="[1215,1292,1656,1682]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="11" pageNumber="514">Figs. 4</figureCitation>
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A−B, 5A−E;
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<taxonomicName id="879729FEC448FD72FF192AC46252F968" box="[151,317,1692,1719]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="11" pageNumber="514" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="72E38E6FC448FD72FF192AC4627AF969" box="[151,277,1692,1718]" italics="true" pageId="11" pageNumber="514">Megophrys</emphasis>
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sp.
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</taxonomicName>
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north in
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<figureCitation id="D8AC4EF8C448FD72FE2A2AC46289F969" box="[420,486,1692,1718]" captionStart="FIGURE 1" captionStartId="4.[151,250,1009,1033]" captionTargetBox="[201,1352,206,977]" captionTargetId="figure-22@4.[163,1424,181,985]" captionTargetPageId="4" captionText="FIGURE 1. Phylogenetic tree based on Maximum Likelihood tree from 471 bp of 16S rRNA gene, values above or below branches represent bootstrap support for Ultrafast Bootstrap Probability, Bayesian Inference posterior probability (UFB/ BPP). The number at the tip of branches corresponds to sample numbers in Appendix 1." figureDoi="http://doi.org/10.5281/zenodo.5598861" httpUri="https://zenodo.org/record/5598861/files/figure.png" pageId="11" pageNumber="514">Fig. 1</figureCitation>
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and
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<tableCitation id="0D1567C6C448FD72FD932AC46100F969" box="[541,623,1692,1718]" captionStart="TABLE 1" captionStartId="3.[151,239,637,663]" captionTargetPageId="3" captionText="TABLE 1. Uncorrected p-distance (%) among Sumatran and Javan Megophrys based on 471 bp fragment of 16S rRNA fragment (lower diagonal) and standard error estimate(s) (above diagonal). GenBank accession numbers for the samples are referenced in Appendix 1 along with voucher number and general locality details." httpUri="http://table.plazi.org/id/14E802F5C440FD7AFF192E2565A4FD64" pageId="11" pageNumber="514" tableUuid="14E802F5C440FD7AFF192E2565A4FD64">Table 1</tableCitation>
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), adult male collected at 2330 h on
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</date>
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from
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<collectingRegion id="82539C9FC448FD72FB612AC463BDF905" isEnumeration="true" pageId="11" pageNumber="514">Aceh Province, Aceh</collectingRegion>
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<collectingMunicipality id="A04CC807C448FD72FF552A9862F5F904" box="[219,410,1728,1755]" pageId="11" pageNumber="514">Tengah Regency</collectingMunicipality>
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,
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,
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(
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,
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,
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a.s.l), by
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<location id="454804A6C448FD72FA872A9866F6F905" LSID="urn:lsid:plazi:treatment:C83EE36BC448FD68FF192A53600AFD76:454804A6C448FD72FA872A9866F6F905" box="[1289,1433,1728,1754]" county="Amph" latitude="4.506444" longLatPrecision="1" longitude="96.86075" municipality="Tengah Regency" name="Elijah Wostl" pageId="11" pageNumber="514" stateProvince="Aceh Province, Aceh">Elijah Wostl</location>
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,
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<location id="454804A6C448FD72FF192ABC6246F920" LSID="urn:lsid:plazi:treatment:C83EE36BC448FD68FF192A53600AFD76:454804A6C448FD72FF192ABC6246F920" box="[151,297,1764,1791]" county="Amph" latitude="4.506444" longLatPrecision="1" longitude="96.86075" municipality="Tengah Regency" name="Ilham Fonna" pageId="11" pageNumber="514" stateProvince="Aceh Province, Aceh">Ilham Fonna</location>
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, and
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<location id="454804A6C448FD72FEED2ABC6141F921" LSID="urn:lsid:plazi:treatment:C83EE36BC448FD68FF192A53600AFD76:454804A6C448FD72FEED2ABC6141F921" box="[355,558,1764,1790]" county="Amph" latitude="4.506444" longLatPrecision="1" longitude="96.86075" municipality="Tengah Regency" name="Muhammad Iksan" pageId="11" pageNumber="514" stateProvince="Aceh Province, Aceh">Muhammad Iksan</location>
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(
|
||
<figureCitation id="D8AC4EF8C448FD72FDB32ABC61EDF920" box="[573,642,1764,1791]" captionStart="FIGURE 6" captionStartId="18.[151,250,1833,1857]" captionTargetBox="[306,1280,924,1777]" captionTargetId="figure-341@18.[274,1314,885,1810]" captionTargetPageId="18" captionText="FIGURE 6. Map of Sumatra showing the known distribution of Sumatran Megophrys. Numbers on map correspond to sample identifiers with molecular data (see Appendix 1), filled markers showed the holotype locality, and markers without numbers are based on the referred specimens or references (see Munir et al., 2018). Black dashed lines indicate major biogeographic breaks within Sumatran Megophrys, modified from (O’Connell et al., 2018a; Shaney et al., 2020). The biogeographic boundary between central and northern Sumatra in this study is different from the one proposed by O’Connell et al. (2018a) and Shaney et al. (2020) because there is still uncertainty as to where that exact boundary occurs." figureDoi="http://doi.org/10.5281/zenodo.5598873" httpUri="https://zenodo.org/record/5598873/files/figure.png" pageId="11" pageNumber="514">Fig. 6</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4028527DC448FD72FF492B506642F8B4" blockId="11.[151,1437,1656,1899]" pageId="11" pageNumber="514">
|
||
<materialsCitation id="F0FF5820C448FD72FF492B506191F899" ID-GBIF-Occurrence="3393350304" accessionNumber="ENS 21030, MT710709" collectingDate="2016-06-01" collectionCode="MT" location="UTA" pageId="11" pageNumber="514" specimenCount="1" stateProvince="Utah" typeStatus="paratype">
|
||
<emphasis id="72E38E6FC448FD72FF492B50625BF8FD" bold="true" box="[199,308,1800,1826]" pageId="11" pageNumber="514">
|
||
<typeStatus id="9F2CECDFC448FD72FF492B50625BF8FD" box="[199,308,1800,1826]" pageId="11" pageNumber="514" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
<collectingRegion id="82539C9FC448FD72FEB32B506218F8FD" box="[317,375,1800,1826]" country="United States of America" name="Utah" pageId="11" pageNumber="514">UTA</collectingRegion>
|
||
A–66178 (field number ENS 21030; GenBank accession no.
|
||
<accessionNumber id="5FC4CF9EC448FD72FBB12B5067AEF8FD" box="[1087,1217,1800,1826]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT710709" pageId="11" pageNumber="514">
|
||
<collectionCode id="2686CAB8C448FD72FBB12B506705F8FD" box="[1087,1130,1800,1826]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="11" pageNumber="514">MT</collectionCode>
|
||
710709
|
||
</accessionNumber>
|
||
;
|
||
<figureCitation id="D8AC4EF8C448FD72FB402B506625F8FD" box="[1230,1354,1800,1826]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="11" pageNumber="514">Fig. 4C–D</figureCitation>
|
||
), a sub adult male collected on at 1856 h on
|
||
<date id="342974BDC448FD72FDB62B7461AEF899" box="[568,705,1836,1863]" pageId="11" pageNumber="514" value="2016-06-01">
|
||
<collectingDate id="246D8D55C448FD72FDB62B7461AEF899" box="[568,705,1836,1863]" pageId="11" pageNumber="514" value="2016-06-01">1 June 2016</collectingDate>
|
||
</date>
|
||
from
|
||
</materialsCitation>
|
||
<materialsCitation id="F0FF5820C448FD72FC8A2B746646F8B4" ID-GBIF-Occurrence="3393350307" collectionCode="V" collectorName="Michael B. Harvey & Eric N. Smith" county="Pidie Regency" elevation="1086" latitude="4.85454" location="Geumpang" longLatPrecision="1" longitude="96.21654" municipality="Geumpang District" pageId="11" pageNumber="514" specimenCount="1" stateProvince="Aceh" typeStatus="paratype">
|
||
<collectingRegion id="82539C9FC448FD72FC8A2B7460C2F899" box="[772,941,1836,1862]" country="Indonesia" name="Aceh" pageId="11" pageNumber="514">Aceh Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="A9492AF1C448FD72FC372B746732F898" box="[953,1117,1836,1863]" pageId="11" pageNumber="514">Pidie Regency</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="A04CC807C448FD72FBE62B74662DF899" box="[1128,1346,1836,1862]" pageId="11" pageNumber="514">Geumpang District</collectingMunicipality>
|
||
, UPT
|
||
<collectionCode id="2686CAB8C448FD72FA072B7566F2F898" box="[1417,1437,1837,1863]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="514" type="Museum">V</collectionCode>
|
||
<location id="454804A6C448FD72FF192B08627DF8B5" LSID="urn:lsid:plazi:treatment:C83EE36BC448FD68FF192A53600AFD76:454804A6C448FD72FF192B08627DF8B5" box="[151,274,1872,1898]" county="Pidie Regency" latitude="4.85454" longLatPrecision="1" longitude="96.21654" municipality="Geumpang District" name="Geumpang" pageId="11" pageNumber="514" stateProvince="Aceh">Geumpang</location>
|
||
(
|
||
<geoCoordinate id="25A334BAC448FD72FEAE2B0862DBF8B5" box="[288,436,1872,1899]" degrees="4.854540" direction="north" orientation="latitude" pageId="11" pageNumber="514" precision="1" value="4.85454">4.854540 °N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="25A334BAC448FD72FE4E2B086133F8B5" box="[448,604,1872,1899]" degrees="96.216540" direction="east" orientation="longitude" pageId="11" pageNumber="514" precision="1" value="96.21654">96.216540 °E</geoCoordinate>
|
||
,
|
||
<quantity id="876FFF98C448FD72FDE62B0861D2F8B4" box="[616,701,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.0859999999999999" pageId="11" pageNumber="514" unit="m" value="1086.0">
|
||
<elevation id="CBBAB54EC448FD72FDE62B0861D2F8B4" box="[616,701,1872,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.0859999999999999" pageId="11" pageNumber="514" unit="m" value="1086.0">1086 m</elevation>
|
||
</quantity>
|
||
a.s.l) by
|
||
<collectorName id="ED6237ABC448FD72FCAA2B086095F8B4" box="[804,1018,1872,1899]" pageId="11" pageNumber="514">Michael B. Harvey</collectorName>
|
||
and
|
||
<collectorName id="ED6237ABC448FD72FBBF2B0867A4F8B5" box="[1073,1227,1872,1899]" pageId="11" pageNumber="514">Eric N. Smith</collectorName>
|
||
(
|
||
<figureCitation id="D8AC4EF8C448FD72FB552B086670F8B4" box="[1243,1311,1872,1899]" captionStart="FIGURE 6" captionStartId="18.[151,250,1833,1857]" captionTargetBox="[306,1280,924,1777]" captionTargetId="figure-341@18.[274,1314,885,1810]" captionTargetPageId="18" captionText="FIGURE 6. Map of Sumatra showing the known distribution of Sumatran Megophrys. Numbers on map correspond to sample identifiers with molecular data (see Appendix 1), filled markers showed the holotype locality, and markers without numbers are based on the referred specimens or references (see Munir et al., 2018). Black dashed lines indicate major biogeographic breaks within Sumatran Megophrys, modified from (O’Connell et al., 2018a; Shaney et al., 2020). The biogeographic boundary between central and northern Sumatra in this study is different from the one proposed by O’Connell et al. (2018a) and Shaney et al. (2020) because there is still uncertainty as to where that exact boundary occurs." figureDoi="http://doi.org/10.5281/zenodo.5598873" httpUri="https://zenodo.org/record/5598873/files/figure.png" pageId="11" pageNumber="514">Fig. 6</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="14E802F5C44FFD75FF1928D1622EFB1A" ID-DOI="http://doi.org/10.5281/zenodo.5598867" ID-Zenodo-Dep="5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="12" pageNumber="515" startId="12.[151,250,1161,1185]" targetBox="[154,1434,181,1137]" targetPageId="12">
|
||
<paragraph id="4028527DC44FFD75FF1928D1622EFB1A" blockId="12.[151,1437,1161,1221]" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF1928D1627DFB7E" bold="true" box="[151,274,1161,1185]" pageId="12" pageNumber="515">FIGURE 4.</emphasis>
|
||
<taxonomicName id="879729FEC44FFD75FE9628D1629DFB7E" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[280,498,1161,1185]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="12" pageNumber="515" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44FFD75FE9628D1629DFB7E" box="[280,498,1161,1185]" italics="true" pageId="12" pageNumber="515">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44FFD75FE7628D16127FB7E" bold="true" box="[504,584,1161,1185]" pageId="12" pageNumber="515">
|
||
<taxonomicNameLabel id="69D03314C44FFD75FE7628D16127FB7E" box="[504,584,1161,1185]" pageId="12" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life, adult male holotype MZB. Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="088D01F6C44FFD75FF4928AC6191FAA6" pageId="12" pageNumber="515" type="etymology">
|
||
<paragraph id="4028527DC44FFD75FF4928AC61EFFAEE" blockId="12.[151,1437,1267,2014]" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF4928AC6221FAD1" bold="true" box="[199,334,1268,1294]" pageId="12" pageNumber="515">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="879729FEC44FFD75FDBC28AB61F0FAD2" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[562,671,1267,1293]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="12" pageNumber="515" phylum="Chordata" rank="species" species="acehensis">
|
||
<emphasis id="72E38E6FC44FFD75FDBC28AB61F0FAD2" box="[562,671,1267,1293]" italics="true" pageId="12" pageNumber="515">acehensis</emphasis>
|
||
</taxonomicName>
|
||
is derived from the province of
|
||
<collectingRegion id="82539C9FC44FFD75FB9828AB673EFAD2" box="[1046,1105,1267,1293]" country="Indonesia" name="Aceh" pageId="12" pageNumber="515">Aceh</collectingRegion>
|
||
in northern
|
||
<collectingRegion id="82539C9FC44FFD75FB6C28AC6650FAD1" box="[1250,1343,1268,1294]" country="Indonesia" name="Sumatera Utara" pageId="12" pageNumber="515">Sumatra</collectingRegion>
|
||
and the Latin suffix–
|
||
<emphasis id="72E38E6FC44FFD75FEA62940620FFAEE" box="[296,352,1304,1329]" italics="true" pageId="12" pageNumber="515">ensis</emphasis>
|
||
meaning from that place.
|
||
</paragraph>
|
||
<paragraph id="4028527DC44FFD75FF4929636059FA89" blockId="12.[151,1437,1267,2014]" box="[199,822,1339,1366]" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF4929636132FA89" bold="true" box="[199,605,1339,1366]" pageId="12" pageNumber="515">Suggested English common name.</emphasis>
|
||
<collectingRegion id="82539C9FC44FFD75FDEA296361F0FA8A" box="[612,671,1339,1365]" country="Indonesia" name="Aceh" pageId="12" pageNumber="515">Aceh</collectingRegion>
|
||
Horned Frog
|
||
</paragraph>
|
||
<paragraph id="4028527DC44FFD75FF4929076191FAA6" blockId="12.[151,1437,1267,2014]" box="[199,766,1375,1402]" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF4929076175FAA5" bold="true" box="[199,538,1375,1402]" pageId="12" pageNumber="515">Suggested Indonesian name.</emphasis>
|
||
Katak–tanduk
|
||
<collectingRegion id="82539C9FC44FFD75FD4D29076191FAA6" box="[707,766,1375,1401]" country="Indonesia" name="Aceh" pageId="12" pageNumber="515">Aceh</collectingRegion>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="088D01F6C44FFD75FF4929DB6159F8AE" pageId="12" pageNumber="515" type="diagnosis">
|
||
<paragraph id="4028527DC44FFD75FF4929DB6159F8AE" blockId="12.[151,1437,1267,2014]" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF4929DB622DFA42" bold="true" box="[199,322,1411,1437]" pageId="12" pageNumber="515">Diagnosis.</emphasis>
|
||
The new species was assigned to the genus
|
||
<taxonomicName id="879729FEC44FFD75FCB629DB60D9FA42" box="[824,950,1411,1437]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="12" pageNumber="515" phylum="Chordata" rank="genus">
|
||
<emphasis id="72E38E6FC44FFD75FCB629DB60D9FA42" box="[824,950,1411,1437]" italics="true" pageId="12" pageNumber="515">Megophrys</emphasis>
|
||
</taxonomicName>
|
||
based on the combination of the following morphological characters, as defined by Kuhl and van Hasselt (1822) and
|
||
<bibRefCitation id="24062F8CC44FFD75FC5729FF67BEFA1D" author="Delorme, M. & Dubois, A. & Grosjean, S. & Ohler, A." box="[985,1233,1447,1474]" pageId="12" pageNumber="515" pagination="6 - 21" refId="ref15873" refString="Delorme, M., Dubois, A., Grosjean, S. & Ohler, A. (2006) Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes, 24 (1 - 4), 6 - 21." type="journal article" year="2006">
|
||
Delorme
|
||
<emphasis id="72E38E6FC44FFD75FBCD29F0671BFA1E" box="[1091,1140,1447,1473]" italics="true" pageId="12" pageNumber="515">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
: (1) pointed snout profile, bearing a pointed projection, protruding laterally beyond the lower jaw; (2) broad and flattened eyelid with palpebral projection; (3) possession of a broad and depressed head; (4) conical spine at the corner of mouth; (5) vertical pupil; (6) presence of maxillary and vomerine teeth.
|
||
<taxonomicName id="879729FEC44FFD75FCCD2A4B675AF9F2" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[835,1077,1555,1581]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="12" pageNumber="515" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44FFD75FCCD2A4B675AF9F2" box="[835,1077,1555,1581]" italics="true" pageId="12" pageNumber="515">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44FFD75FBB22A4C67F9F9F1" bold="true" box="[1084,1174,1556,1582]" pageId="12" pageNumber="515">
|
||
<taxonomicNameLabel id="69D03314C44FFD75FBB22A4C67F9F9F1" box="[1084,1174,1556,1582]" pageId="12" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be diagnosed from its geographically relevant congeners in the Sunda Shelf and the
|
||
<collectingCountry id="388012EDC44FFD75FCE32A6F6084F98E" box="[877,1003,1591,1617]" name="Philippines" pageId="12" pageNumber="515">Philippines</collectingCountry>
|
||
by the following combination of characters: medium body size, stocky (SVLh
|
||
<quantity id="876FFF98C44FFD75FDD02A0461AEF9A9" box="[606,705,1628,1654]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="12" pageNumber="515" unit="mm" value="38.1">38.1 mm</quantity>
|
||
in adult male); snout pointed with a short, acute rostral appendage (RSAL 1.3% in adult male); relatively short triangular palpebral projection with acute tip (EHL 25.1% UEWh in adult male); head length relatively short (RHLh 37.2% in adult male); head wider than its long (HW 113.6% HLh in adult male); tympanum distinct, about one-third of eye diameter (TDH 35.6% ED in adult male), nearly rounded (TDH 85.9 % TDV in adult male); vomerine teeth present; a pair of dorsolateral folds, extending from shoulder, above axilla to groin; dorsal and lateral skin tuberculate, shagreened on the throat to belly; short lower arm (RLAL 45.8% in adult male); foot nearly as long as thigh (FL 96.6% TL in adult male); tibiotarsal articulation reaching posterior of eye; toe webbing absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="088D01F6C44FFD68FF492B23612CFE9E" lastPageId="17" lastPageNumber="520" pageId="12" pageNumber="515" type="description">
|
||
<paragraph id="4028527DC44FFD74FF492B2360DFFDBE" blockId="12.[151,1437,1267,2014]" lastBlockId="13.[151,1437,151,609]" lastPageId="13" lastPageNumber="516" pageId="12" pageNumber="515">
|
||
<emphasis id="72E38E6FC44FFD75FF492B236191F84A" bold="true" box="[199,766,1915,1942]" pageId="12" pageNumber="515">
|
||
Description of
|
||
<typeStatus id="9F2CECDFC44FFD75FEF92B2462B1F849" box="[375,478,1916,1942]" pageId="12" pageNumber="515" type="holotype">holotype</typeStatus>
|
||
(measurements in mm).
|
||
</emphasis>
|
||
Adult male, medium body size (SVLh 38.1, SVL 37.4) and habitus stocky; head depressed and broad, wider (HW 16.4, 42.9% SVLh) than long (HL 13.3, 34.9% SVLh); snout short (SL 4.2, 11.1% SVLh), pointed at tip with acute short rostral appendage (SAL 0.7, 1.8% SVLh), laterally protruding and projecting beyond lower jaw; nostril positioned laterally, near to snout than to eye; eye positioned laterally, large, nearly three times of tympanum horizontal diameter (ED 5.5, 280.5% TDH), eye diameter slightly wider (ED 5.5, 14.4% SVLh) than snout–horn length (SLh 5.4, 14.2% SVLh), about two and three-quarter times of nostril–eye length (ED 275% NEL), pupil vertical elliptical; canthus rostralis with sharp, angular ridge, lore sloping and concave; internarial distance (IND 4.0, 10.4% SVLh) about two-thirds of interorbital distance (IND 68.4% IOD); palpebral projection length about one-quarter of total upper eyelid width (EHL 1.2, 25.1% UEWh), tip acute, surface smooth and scattered with small and low tubercles; tympanum distinct, smooth, oval, slightly rounded (TDV 2.3, 6.0% SVLh; TDH 2.0, 5.1% SVLh; TDH 85.9% TDV); angular supratympanic fold, distinct, widened anteriorly, narrowed posteriorly, extending from behind eye, curving down around upper border of tympanum and ending above axilla; white conical tubercles behind the supratympanic fold and anterior to axilla; spinous gland on corner of mouth on jaw angle; single row of maxillary teeth present; vomerine teeth in two widely separated groups, at level posterior borders of choana; tongue lanceolate, notched posteriorly, without papillae; median subgular vocal sac present, having slit–like opening on each side of jaw commissures.
|
||
</paragraph>
|
||
<caption id="14E802F5C44EFD74FF192B2C6769F870" ID-DOI="http://doi.org/10.5281/zenodo.5598871" ID-Zenodo-Dep="5598871" httpUri="https://zenodo.org/record/5598871/files/figure.png" pageId="13" pageNumber="516" startId="13.[151,250,1908,1932]" targetBox="[204,1418,671,1823]" targetPageId="13">
|
||
<paragraph id="4028527DC44EFD74FF192B2C6769F870" blockId="13.[151,1437,1908,1968]" pageId="13" pageNumber="516">
|
||
<emphasis id="72E38E6FC44EFD74FF192B2C627CF853" bold="true" box="[151,275,1908,1932]" pageId="13" pageNumber="516">FIGURE 5.</emphasis>
|
||
<taxonomicName id="879729FEC44EFD74FE942B2C629AF853" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[282,501,1908,1932]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="13" pageNumber="516" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44EFD74FE942B2C629AF853" box="[282,501,1908,1932]" italics="true" pageId="13" pageNumber="516">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44EFD74FE732B2C6121F853" bold="true" box="[509,590,1908,1932]" pageId="13" pageNumber="516">
|
||
<taxonomicNameLabel id="69D03314C44EFD74FE732B2C6121F853" box="[509,590,1908,1932]" pageId="13" pageNumber="516" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, adult male holotype (MZB. Amph 26098; SVL 37.4 mm): dorsal (A), ventral (B), lateral head (C), ventral right hand (D), and ventral right foot (E) views, scale = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="14E802F5C44DFD77FF192ED066F0FD7D" ID-Table-UUID="14E802F5C44DFD77FF192ED066F0FD7D" box="[151,1439,648,674]" httpUri="http://table.plazi.org/id/14E802F5C44DFD77FF192ED066F0FD7D" pageId="14" pageNumber="517" startId="14.[151,239,648,674]" targetBox="[165,1969,689,1358]" targetIsTable="true" targetPageId="14">
|
||
<paragraph id="4028527DC44DFD77FF192ED066F0FD7D" blockId="14.[151,1439,648,674]" box="[151,1439,648,674]" pageId="14" pageNumber="517">
|
||
<emphasis id="72E38E6FC44DFD77FF192ED06265FD7D" bold="true" box="[151,266,648,674]" pageId="14" pageNumber="517">TABLE 3.</emphasis>
|
||
Morphological variation between Sunda Shelf and Philippine
|
||
<taxonomicName id="879729FEC44DFD77FC2B2ED06772FD7D" box="[933,1053,648,674]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="genus">
|
||
<emphasis id="72E38E6FC44DFD77FC2B2ED06772FD7D" box="[933,1053,648,674]" italics="true" pageId="14" pageNumber="517">Megophrys</emphasis>
|
||
</taxonomicName>
|
||
(modified from
|
||
<bibRefCitation id="24062F8CC44DFD77FB402ED066FBFD7D" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1230,1428,648,674]" pageId="14" pageNumber="517" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44DFD77FA992ED0663CFD7D" box="[1303,1363,648,674]" italics="true" pageId="14" pageNumber="517">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4028527DC44DFD77FF2B2EE96435FA91" pageId="14" pageNumber="517">
|
||
<table id="3297A0DDC44D0286FF2B2EE964DEFA91" box="[165,1969,689,1358]" gridcols="6" gridrows="18" pageId="14" pageNumber="517">
|
||
<tr id="FEA7503FC44D0286FF2B2EE964DEFD15" box="[165,1969,689,714]" gridrow="0" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2EE9629FFD15" box="[165,496,689,714]" gridcol="0" gridrow="0" pageId="14" pageNumber="517">Species</th>
|
||
<th id="BD763943C44D0286FDC32EE96058FD15" box="[589,823,689,714]" gridcol="1" gridrow="0" pageId="14" pageNumber="517">Parietoscapular ridge *</th>
|
||
<th id="BD763943C44D0286FC0F2EE96752FD15" box="[897,1085,689,714]" gridcol="2" gridrow="0" pageId="14" pageNumber="517">Vomerine teeth</th>
|
||
<th id="BD763943C44D0286FBE62EE96631FD15" box="[1128,1374,689,714]" gridcol="3" gridrow="0" pageId="14" pageNumber="517">Dorsolateral fold **</th>
|
||
<th id="BD763943C44D0286FA072EE9650BFD15" box="[1417,1636,689,714]" gridcol="4" gridrow="0" pageId="14" pageNumber="517">Rostral Appendage</th>
|
||
<th id="BD763943C44D0286F9212EE964DEFD15" box="[1711,1969,689,714]" gridcol="5" gridrow="0" pageId="14" pageNumber="517">Eyelid Appendage</th>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2E8F64DEFD2F" box="[165,1969,727,752]" gridrow="1" pageId="14" pageNumber="517" rowspan-0="1">
|
||
<td id="BD763943C44D0286FDC32E8F6058FD2F" box="[589,823,727,752]" gridcol="1" gridrow="1" pageId="14" pageNumber="517">present (+), absent (−)</td>
|
||
<td id="BD763943C44D0286FC0F2E8F6752FD2F" box="[897,1085,727,752]" gridcol="2" gridrow="1" pageId="14" pageNumber="517">present (+), absent</td>
|
||
<td id="BD763943C44D0286FBE62E8F6631FD2F" box="[1128,1374,727,752]" gridcol="3" gridrow="1" pageId="14" pageNumber="517">Type I, II, III, IV, absent</td>
|
||
<td id="BD763943C44D0286FA072E8F650BFD2F" box="[1417,1636,727,752]" gridcol="4" gridrow="1" pageId="14" pageNumber="517">present (+), less (−−),</td>
|
||
<td id="BD763943C44D0286F9212E8F64DEFD2F" box="[1711,1969,727,752]" gridcol="5" gridrow="1" pageId="14" pageNumber="517">absent (−), triangular (+),</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2EA364DEFCCB" box="[165,1969,763,788]" gridrow="2" pageId="14" pageNumber="517" rowspan-0="1" rowspan-1="1">
|
||
<td id="BD763943C44D0286FC0F2EA36752FCCB" box="[897,1085,763,788]" gridcol="2" gridrow="2" pageId="14" pageNumber="517">(−)</td>
|
||
<td id="BD763943C44D0286FBE62EA36631FCCB" box="[1128,1374,763,788]" gridcol="3" gridrow="2" pageId="14" pageNumber="517">(−)</td>
|
||
<td id="BD763943C44D0286FA072EA3650BFCCB" box="[1417,1636,763,788]" gridcol="4" gridrow="2" pageId="14" pageNumber="517">absent (−)</td>
|
||
<td id="BD763943C44D0286F9212EA364DEFCCB" box="[1711,1969,763,788]" gridcol="5" gridrow="2" pageId="14" pageNumber="517">tubercle (++)</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2F7C64DEFCE2" box="[165,1969,804,829]" gridrow="3" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2F7C629FFCE2" box="[165,496,804,829]" gridcol="0" gridrow="3" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2F7C62F5FCE2" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[165,410,804,829]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="selatanensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2F7C62F5FCE2" box="[165,410,804,829]" italics="true" pageId="14" pageNumber="517">Megophrys selatanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44DFD77FE2E2F7C629FFCE2" bold="true" box="[416,496,804,829]" pageId="14" pageNumber="517">
|
||
<taxonomicNameLabel id="69D03314C44DFD77FE2E2F7C629FFCE2" box="[416,496,804,829]" pageId="14" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32F7C6058FCE2" box="[589,823,804,829]" gridcol="1" gridrow="3" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2F7C6752FCE2" box="[897,1085,804,829]" gridcol="2" gridrow="3" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62F7C6631FCE2" box="[1128,1374,804,829]" gridcol="3" gridrow="3" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072F7C650BFCE2" box="[1417,1636,804,829]" gridcol="4" gridrow="3" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286F9212F7C64DEFCE2" box="[1711,1969,804,829]" gridcol="5" gridrow="3" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2F1164DEFCBD" box="[165,1969,841,866]" gridrow="4" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2F11629FFCBD" box="[165,496,841,866]" gridcol="0" gridrow="4" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2F1162EFFCBD" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[165,384,841,866]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2F1162EFFCBD" box="[165,384,841,866]" italics="true" pageId="14" pageNumber="517">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44DFD77FE092F1162B8FCBD" bold="true" box="[391,471,841,866]" pageId="14" pageNumber="517">
|
||
<taxonomicNameLabel id="69D03314C44DFD77FE092F1162B8FCBD" box="[391,471,841,866]" pageId="14" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32F116058FCBD" box="[589,823,841,866]" gridcol="1" gridrow="4" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2F116752FCBD" box="[897,1085,841,866]" gridcol="2" gridrow="4" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62F116631FCBD" box="[1128,1374,841,866]" gridcol="3" gridrow="4" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072F11650BFCBD" box="[1417,1636,841,866]" gridcol="4" gridrow="4" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286F9212F1164DEFCBD" box="[1711,1969,841,866]" gridcol="5" gridrow="4" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2F3764DEFC57" box="[165,1969,879,904]" gridrow="5" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2F37629FFC57" box="[165,496,879,904]" gridcol="0" gridrow="5" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2F376232FC57" box="[165,349,879,904]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="lancip">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2F376232FC57" box="[165,349,879,904]" italics="true" pageId="14" pageNumber="517">Megophrys lancip</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32F376058FC57" box="[589,823,879,904]" gridcol="1" gridrow="5" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2F376752FC57" box="[897,1085,879,904]" gridcol="2" gridrow="5" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62F376631FC57" box="[1128,1374,879,904]" gridcol="3" gridrow="5" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072F37650BFC57" box="[1417,1636,879,904]" gridcol="4" gridrow="5" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286F9212F3764DEFC57" box="[1711,1969,879,904]" gridcol="5" gridrow="5" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2FCD64DEFC71" box="[165,1969,917,942]" gridrow="6" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2FCD629FFC71" box="[165,496,917,942]" gridcol="0" gridrow="6" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2FCD6218FC71" box="[165,375,917,942]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="montana">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2FCD6218FC71" box="[165,375,917,942]" italics="true" pageId="14" pageNumber="517">Megophrys montana</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32FCD6058FC71" box="[589,823,917,942]" gridcol="1" gridrow="6" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2FCD6752FC71" box="[897,1085,917,942]" gridcol="2" gridrow="6" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62FCD6631FC71" box="[1128,1374,917,942]" gridcol="3" gridrow="6" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072FCD650BFC71" box="[1417,1636,917,942]" gridcol="4" gridrow="6" pageId="14" pageNumber="517">−− / +</td>
|
||
<td id="BD763943C44D0286F9212FCD64DEFC71" box="[1711,1969,917,942]" gridcol="5" gridrow="6" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2FE364DEFC0B" box="[165,1969,955,980]" gridrow="7" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2FE3629FFC0B" box="[165,496,955,980]" gridcol="0" gridrow="7" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2FE36214FC0B" box="[165,379,955,980]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="parallela">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2FE36214FC0B" box="[165,379,955,980]" italics="true" pageId="14" pageNumber="517">Megophrys parallela</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32FE36058FC0B" box="[589,823,955,980]" gridcol="1" gridrow="7" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2FE36752FC0B" box="[897,1085,955,980]" gridcol="2" gridrow="7" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62FE36631FC0B" box="[1128,1374,955,980]" gridcol="3" gridrow="7" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072FE3650BFC0B" box="[1417,1636,955,980]" gridcol="4" gridrow="7" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F9212FE364DEFC0B" box="[1711,1969,955,980]" gridcol="5" gridrow="7" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B2FB964DEFC25" box="[165,1969,993,1018]" gridrow="8" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2FB9629FFC25" box="[165,496,993,1018]" gridcol="0" gridrow="8" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2FB9620DFC25" box="[165,354,993,1018]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="nasuta">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2FB9620DFC25" box="[165,354,993,1018]" italics="true" pageId="14" pageNumber="517">Megophrys nasuta</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC32FB96058FC25" box="[589,823,993,1018]" gridcol="1" gridrow="8" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F2FB96752FC25" box="[897,1085,993,1018]" gridcol="2" gridrow="8" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE62FB96631FC25" box="[1128,1374,993,1018]" gridcol="3" gridrow="8" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072FB9650BFC25" box="[1417,1636,993,1018]" gridcol="4" gridrow="8" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286F9212FB964DEFC25" box="[1711,1969,993,1018]" gridcol="5" gridrow="8" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B285E64DEFBC0" box="[165,1969,1030,1055]" gridrow="9" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B285E629FFBC0" box="[165,496,1030,1055]" gridcol="0" gridrow="9" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B285E62EDFBC0" box="[165,386,1030,1055]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="stejnegeri">
|
||
<emphasis id="72E38E6FC44DFD77FF2B285E62EDFBC0" box="[165,386,1030,1055]" italics="true" pageId="14" pageNumber="517">Megophrys stejnegeri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC3285E6058FBC0" box="[589,823,1030,1055]" gridcol="1" gridrow="9" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F285E6752FBC0" box="[897,1085,1030,1055]" gridcol="2" gridrow="9" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FBE6285E6631FBC0" box="[1128,1374,1030,1055]" gridcol="3" gridrow="9" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA07285E650BFBC0" box="[1417,1636,1030,1055]" gridcol="4" gridrow="9" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F921285E64DEFBC0" box="[1711,1969,1030,1055]" gridcol="5" gridrow="9" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B287464DEFB9A" box="[165,1969,1068,1093]" gridrow="10" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2874629FFB9A" box="[165,496,1068,1093]" gridcol="0" gridrow="10" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B287462ACFB9A" box="[165,451,1068,1093]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="kalimantanensis">
|
||
<emphasis id="72E38E6FC44DFD77FF2B287462ACFB9A" box="[165,451,1068,1093]" italics="true" pageId="14" pageNumber="517">Megophrys kalimantanensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC328746058FB9A" box="[589,823,1068,1093]" gridcol="1" gridrow="10" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F28746752FB9A" box="[897,1085,1068,1093]" gridcol="2" gridrow="10" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE628746631FB9A" box="[1128,1374,1068,1093]" gridcol="3" gridrow="10" pageId="14" pageNumber="517">I</td>
|
||
<td id="BD763943C44D0286FA072874650BFB9A" box="[1417,1636,1068,1093]" gridcol="4" gridrow="10" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286F921287464DEFB9A" box="[1711,1969,1068,1093]" gridcol="5" gridrow="10" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B280A64DEFBB4" box="[165,1969,1106,1131]" gridrow="11" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B280A629FFBB4" box="[165,496,1106,1131]" gridcol="0" gridrow="11" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B280A62E3FBB4" box="[165,396,1106,1131]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="kobayashii">
|
||
<emphasis id="72E38E6FC44DFD77FF2B280A62E3FBB4" box="[165,396,1106,1131]" italics="true" pageId="14" pageNumber="517">Megophrys kobayashii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC3280A6058FBB4" box="[589,823,1106,1131]" gridcol="1" gridrow="11" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F280A6752FBB4" box="[897,1085,1106,1131]" gridcol="2" gridrow="11" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE6280A6631FBB4" box="[1128,1374,1106,1131]" gridcol="3" gridrow="11" pageId="14" pageNumber="517">II</td>
|
||
<td id="BD763943C44D0286FA07280A650BFBB4" box="[1417,1636,1106,1131]" gridcol="4" gridrow="11" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F921280A64DEFBB4" box="[1711,1969,1106,1131]" gridcol="5" gridrow="11" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B282064DEFB4E" box="[165,1969,1144,1169]" gridrow="12" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2820629FFB4E" box="[165,496,1144,1169]" gridcol="0" gridrow="12" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B28206206FB4E" authorityName="Taylor" authorityYear="1920" box="[165,361,1144,1169]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="ligayae">
|
||
<emphasis id="72E38E6FC44DFD77FF2B28206206FB4E" box="[165,361,1144,1169]" italics="true" pageId="14" pageNumber="517">Megophrys ligayae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC328206058FB4E" box="[589,823,1144,1169]" gridcol="1" gridrow="12" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F28206752FB4E" box="[897,1085,1144,1169]" gridcol="2" gridrow="12" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE628206631FB4E" box="[1128,1374,1144,1169]" gridcol="3" gridrow="12" pageId="14" pageNumber="517">II</td>
|
||
<td id="BD763943C44D0286FA072820650BFB4E" box="[1417,1636,1144,1169]" gridcol="4" gridrow="12" pageId="14" pageNumber="517">−−</td>
|
||
<td id="BD763943C44D0286F921282064DEFB4E" box="[1711,1969,1144,1169]" gridcol="5" gridrow="12" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B28C664DEFB68" box="[165,1969,1182,1207]" gridrow="13" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B28C6629FFB68" box="[165,496,1182,1207]" gridcol="0" gridrow="13" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B28C66211FB68" box="[165,382,1182,1207]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="72E38E6FC44DFD77FF2B28C66211FB68" box="[165,382,1182,1207]" italics="true" pageId="14" pageNumber="517">Megophrys baluensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC328C66058FB68" box="[589,823,1182,1207]" gridcol="1" gridrow="13" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F28C66752FB68" box="[897,1085,1182,1207]" gridcol="2" gridrow="13" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE628C66631FB68" box="[1128,1374,1182,1207]" gridcol="3" gridrow="13" pageId="14" pageNumber="517">III</td>
|
||
<td id="BD763943C44D0286FA0728C6650BFB68" box="[1417,1636,1182,1207]" gridcol="4" gridrow="13" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F92128C664DEFB68" box="[1711,1969,1182,1207]" gridcol="5" gridrow="13" pageId="14" pageNumber="517">++</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B289B64DEFB03" box="[165,1969,1219,1244]" gridrow="14" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B289B629FFB03" box="[165,496,1219,1244]" gridcol="0" gridrow="14" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B289B62FAFB03" authorityName="Inger" authorityYear="1989" box="[165,405,1219,1244]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="edwardinae">
|
||
<emphasis id="72E38E6FC44DFD77FF2B289B62FAFB03" box="[165,405,1219,1244]" italics="true" pageId="14" pageNumber="517">Megophrys edwardinae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC3289B6058FB03" box="[589,823,1219,1244]" gridcol="1" gridrow="14" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FC0F289B6752FB03" box="[897,1085,1219,1244]" gridcol="2" gridrow="14" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FBE6289B6631FB03" box="[1128,1374,1219,1244]" gridcol="3" gridrow="14" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FA07289B650BFB03" box="[1417,1636,1219,1244]" gridcol="4" gridrow="14" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F921289B64DEFB03" box="[1711,1969,1219,1244]" gridcol="5" gridrow="14" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B28B164DEFADD" box="[165,1969,1257,1282]" gridrow="15" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B28B1629FFADD" box="[165,496,1257,1282]" gridcol="0" gridrow="15" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B28B16233FADD" box="[165,348,1257,1282]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="72E38E6FC44DFD77FF2B28B16233FADD" box="[165,348,1257,1282]" italics="true" pageId="14" pageNumber="517">Megophrys dringi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC328B16058FADD" box="[589,823,1257,1282]" gridcol="1" gridrow="15" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FC0F28B16752FADD" box="[897,1085,1257,1282]" gridcol="2" gridrow="15" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286FBE628B16631FADD" box="[1128,1374,1257,1282]" gridcol="3" gridrow="15" pageId="14" pageNumber="517">IV</td>
|
||
<td id="BD763943C44D0286FA0728B1650BFADD" box="[1417,1636,1257,1282]" gridcol="4" gridrow="15" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F92128B164DEFADD" box="[1711,1969,1257,1282]" gridcol="5" gridrow="15" pageId="14" pageNumber="517">++</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B295764DEFAF7" box="[165,1969,1295,1320]" gridrow="16" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B2957629FFAF7" box="[165,496,1295,1320]" gridcol="0" gridrow="16" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B2957620DFAF7" authorityName="Boulenger" authorityYear="1903" box="[165,354,1295,1320]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="aceras">
|
||
<emphasis id="72E38E6FC44DFD77FF2B2957620DFAF7" box="[165,354,1295,1320]" italics="true" pageId="14" pageNumber="517">Megophrys aceras</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC329576058FAF7" box="[589,823,1295,1320]" gridcol="1" gridrow="16" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FC0F29576752FAF7" box="[897,1085,1295,1320]" gridcol="2" gridrow="16" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE629576631FAF7" box="[1128,1374,1295,1320]" gridcol="3" gridrow="16" pageId="14" pageNumber="517">IV</td>
|
||
<td id="BD763943C44D0286FA072957650BFAF7" box="[1417,1636,1295,1320]" gridcol="4" gridrow="16" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F921295764DEFAF7" box="[1711,1969,1295,1320]" gridcol="5" gridrow="16" pageId="14" pageNumber="517">+</td>
|
||
</tr>
|
||
<tr id="FEA7503FC44D0286FF2B296D64DEFA91" box="[165,1969,1333,1358]" gridrow="17" pageId="14" pageNumber="517">
|
||
<th id="BD763943C44D0286FF2B296D629FFA91" box="[165,496,1333,1358]" gridcol="0" gridrow="17" pageId="14" pageNumber="517">
|
||
<taxonomicName id="879729FEC44DFD77FF2B296D621BFA91" authorityName="Boulenger" authorityYear="1886" box="[165,372,1333,1358]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="14" pageNumber="517" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="72E38E6FC44DFD77FF2B296D621BFA91" box="[165,372,1333,1358]" italics="true" pageId="14" pageNumber="517">Megophrys longipes</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="BD763943C44D0286FDC3296D6058FA91" box="[589,823,1333,1358]" gridcol="1" gridrow="17" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FC0F296D6752FA91" box="[897,1085,1333,1358]" gridcol="2" gridrow="17" pageId="14" pageNumber="517">+</td>
|
||
<td id="BD763943C44D0286FBE6296D6631FA91" box="[1128,1374,1333,1358]" gridcol="3" gridrow="17" pageId="14" pageNumber="517">IV</td>
|
||
<td id="BD763943C44D0286FA07296D650BFA91" box="[1417,1636,1333,1358]" gridcol="4" gridrow="17" pageId="14" pageNumber="517">−</td>
|
||
<td id="BD763943C44D0286F921296D64DEFA91" box="[1711,1969,1333,1358]" gridcol="5" gridrow="17" pageId="14" pageNumber="517">++</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="4028527DC44DFD77FF1929236158F9DE" blockId="14.[151,2041,1403,1537]" pageId="14" pageNumber="517">
|
||
* Fold forming Y, X or H on the parietoscapular region to the level of axilla; ** Dorsolateral fold shape: dorsolateral folds are elongated and extend from the parietoscapular region to the groin (
|
||
<typeStatus id="9F2CECDFC44DFD77FED829C762E2FA66" box="[342,397,1439,1465]" pageId="14" pageNumber="517">Type</typeStatus>
|
||
I); dorsolateral folds extend from the central of parietoscapular region to mid-body (
|
||
<typeStatus id="9F2CECDFC44DFD77FA9929C76620FA66" box="[1303,1359,1439,1465]" pageId="14" pageNumber="517">Type</typeStatus>
|
||
II); multiple dorsolateral folds - at least three or four - and they are discontinuous, formed by a series of elongated tubercles (
|
||
<typeStatus id="9F2CECDFC44DFD77FCA3299B600BFA02" box="[813,868,1475,1501]" pageId="14" pageNumber="517">Type</typeStatus>
|
||
III); dorsolateral folds are elongated and curve from the axillary region towards (and reaching) the posterior dorsal margin of tympanum (
|
||
<typeStatus id="9F2CECDFC44DFD77FE4329BF616BF9DE" box="[461,516,1511,1537]" pageId="14" pageNumber="517">Type</typeStatus>
|
||
IV).
|
||
</paragraph>
|
||
<paragraph id="4028527DC44CFD76FF492CCF610DFE9D" blockId="15.[151,1437,150,2014]" pageId="15" pageNumber="518">Forelimb slender and short, hand length about half of arm length (HAL 9.5, 54.3% LAL), lower arm proximally enlarged, wider than upper arm; fingers moderately slender, with rounded and swollen tips, unwebbed and lacking of lateral fringes; finger length formula I<II<IV<III (fin1L 2.6, 6.9% SVLh; fin2L 3.2, 8.4% SVLh; fin4L 3.5, 9.2% SVLh; and fin3L 4.9, 12.8% SVLh); subarticular tubercles absent; outer palmar tubercle smaller than inner (OPTL 1.9, 5.0% SVLh; IPTL 2.5, 6.6% SVLh).</paragraph>
|
||
<paragraph id="4028527DC44CFD76FF492D1360CAFDC6" blockId="15.[151,1437,150,2014]" pageId="15" pageNumber="518">Hindlimb slender, moderately long (HLL 52.0, 136.5% SVLh), thigh (FML 17.1, 44.8% SVLh) slightly longer than tibia (TL 15.4, 40.4% SVLh), about twice of tarsus length (TSL 8.5, 22.3% SVLh), and nearly as long as foot (FL 14.9, 39.1% SVLh); toe length formula I<II<V<III<IV (Toe1L 2.1, 5.4% SVLh; Toe2L 3.3, 8.6% SVLh; Toe5L 4.5, 11.8% SVLh; Toe3L 4.6, 12.0% SVLh; and Toe4L 7.8, 20.6% SVLh); tibiotarsal articulation reaching to posterior corner of eye; inner metatarsal tubercle moderately large (IMTL 2.0, 5.1% SVLh), outer absent; subarticular tubercles absent, toes with rounded and swollen tips, toes web absent.</paragraph>
|
||
<paragraph id="4028527DC44CFD76FF492E7B67E0FCE6" blockId="15.[151,1437,150,2014]" pageId="15" pageNumber="518">Dorsal skin surface tuberculate, low dense tubercles in entire dorsal and larger-sized tubercles mostly on forelimb, hindlimb, flanks and groin, shagreened in preservative; a pair of dorsolateral folds present, one on each side, extending from shoulder above the axilla to groin; vent with dorsal dermal accessory extension; transverse folds on limbs present, three on lower arm, three on thigh and three on tibia; limbs skin laterally tuberculate, larger size of tubercles than on dorsal side; ventral skin smooth shagreened on throat to chest, but smooth on belly and limbs, wrinkled in preservative; a pair of white conical pectoral glands, at base of axilla; white conical femoral glands, at the mid flanks of the posterior thigh; dark brown microspinules nuptial pads covering dorsal and median surfaces of the distal half of metacarpal and proximal half of basal phalanx of first and second fingers.</paragraph>
|
||
<paragraph id="4028527DC44CFD76FF492F1A62B0FB36" blockId="15.[151,1437,150,2014]" pageId="15" pageNumber="518">
|
||
<emphasis id="72E38E6FC44CFD76FF492F1A6220FC83" bold="true" box="[199,335,834,860]" pageId="15" pageNumber="518">Coloration.</emphasis>
|
||
In life (
|
||
<figureCitation id="D8AC4EF8C44CFD76FE272F1B614DFC82" box="[425,546,835,861]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="15" pageNumber="518">Fig. 4A–B</figureCitation>
|
||
,
|
||
<figureCitation id="D8AC4EF8C44CFD76FDA32F1A611BFC82" box="[557,628,834,861]" captionStart="FIGURE 5" captionStartId="13.[151,250,1908,1932]" captionTargetBox="[204,1418,671,1823]" captionTargetId="figure-228@13.[163,1424,633,1884]" captionTargetPageId="13" captionText="FIGURE 5. Megophrys acehensis sp. nov., adult male holotype (MZB. Amph 26098; SVL 37.4 mm): dorsal (A), ventral (B), lateral head (C), ventral right hand (D), and ventral right foot (E) views, scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5598871" httpUri="https://zenodo.org/record/5598871/files/figure.png" pageId="15" pageNumber="518">5A–B</figureCitation>
|
||
), pupil dark, iris golden–brown, base of dorsum immaculate dark brown with indistinct dark brown inverted triangle pattern on parieto-orbital to scapular region and inverted “V” like pattern on back, positioned closer to vent than to parietal region; dorsal surface of head, forelimb and hindlimb darker than lateral body; dark brown transverse folds present on dorsal surface of forearm, thigh and tibia, two on the forearm, three on thigh and three on tibia, fingers with dark brown crossbars, one on first and second fingers, two on third and fourth fingers; dorsomedial surfaces of first and second fingers with dark brown microspinular nuptial pads; knee with irregular dark brown spots; dorsal surface and lateral sides of head dark brown, irregular lighter brown pattern below canthus rostralis, light brown on underside of eyelids; body flanks dark brown and unmarked, lighter than dorsum; forelimb flanks dark brown, as dorsum; hindlimb flanks cream with dark brown blotches; ventral surface from throat to chest dark brown with heavy dark longitudinal marking; belly and underside of fore- and hindlimbs light cream with dark brown blotches; in preservative, pattern remains, but dorsal, lateral, and ventral surfaces are darker (
|
||
<figureCitation id="D8AC4EF8C44CFD76FEF4289762BCFB36" box="[378,467,1230,1257]" captionStart="FIGURE 5" captionStartId="13.[151,250,1908,1932]" captionTargetBox="[204,1418,671,1823]" captionTargetId="figure-228@13.[163,1424,633,1884]" captionTargetPageId="13" captionText="FIGURE 5. Megophrys acehensis sp. nov., adult male holotype (MZB. Amph 26098; SVL 37.4 mm): dorsal (A), ventral (B), lateral head (C), ventral right hand (D), and ventral right foot (E) views, scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5598871" httpUri="https://zenodo.org/record/5598871/files/figure.png" pageId="15" pageNumber="518">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4028527DC44CFD76FF4928AA671CFA3A" blockId="15.[151,1437,150,2014]" pageId="15" pageNumber="518">
|
||
<emphasis id="72E38E6FC44CFD76FF4928AA6250FAD3" bold="true" box="[199,319,1266,1292]" pageId="15" pageNumber="518">Variation.</emphasis>
|
||
The
|
||
<specimenCount id="569199F4C44CFD76FEF428AB62B4FAD2" box="[378,475,1267,1293]" pageId="15" pageNumber="518" type="generic">two type</specimenCount>
|
||
specimens are morphometrically similar. The
|
||
<typeStatus id="9F2CECDFC44CFD76FC6928AB6726FAD2" box="[999,1097,1267,1293]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
has slightly longer rostral appendage, head, and snout-nostril and snout lengths, but a slightly shorter palpebral projection, and a thicker and narrower head. Morphometric variation is shown in
|
||
<tableCitation id="0D1567C6C44CFD76FD3929636063FA8A" box="[695,780,1339,1365]" captionStart="TABLE 2" captionStartId="8.[151,239,474,500]" captionTargetPageId="8" captionText="TABLE 2. Measurements (in mm) of Megophrys selatanensis sp. nov. and Megophrys acehensis sp. nov." httpUri="http://table.plazi.org/id/14E802F5C44BFD71FF192D82666FFE2B" pageId="15" pageNumber="518" tableUuid="14E802F5C44BFD71FF192D82666FFE2B">Table 2</tableCitation>
|
||
. The transverse fold in the occipital region varies in degree of development, being distinct on the
|
||
<typeStatus id="9F2CECDFC44CFD76FDC8290761C7FAA6" box="[582,680,1375,1401]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
but indistinct on the
|
||
<typeStatus id="9F2CECDFC44CFD76FC1529076090FAA6" box="[923,1023,1375,1401]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
|
||
. The dorsal coloration in life is dark brown on the
|
||
<typeStatus id="9F2CECDFC44CFD76FEBA29DB62F9FA42" box="[308,406,1411,1437]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
and orange-brown on the
|
||
<typeStatus id="9F2CECDFC44CFD76FD3129DB604CFA42" box="[703,803,1411,1437]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
|
||
. The ventral color of the
|
||
<specimenCount id="569199F4C44CFD76FBB129DB6789FA42" box="[1087,1254,1411,1437]" pageId="15" pageNumber="518" type="generic">two specimens</specimenCount>
|
||
is marbled with brown in both specimens but, the background color of the
|
||
<typeStatus id="9F2CECDFC44CFD76FCBA29FF60F9FA1E" box="[820,918,1447,1473]" pageId="15" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
is cream anteriorly and light grey posteriorly while that of the
|
||
<typeStatus id="9F2CECDFC44CFD76FEDB299362DAFA3A" box="[341,437,1483,1509]" pageId="15" pageNumber="518" type="paratype">paratype</typeStatus>
|
||
is dark orange anteriorly and whitish posteriorly (
|
||
<figureCitation id="D8AC4EF8C44CFD76FC6929936745FA3A" box="[999,1066,1483,1509]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="15" pageNumber="518">Fig. 4</figureCitation>
|
||
A−D).
|
||
</paragraph>
|
||
<paragraph id="4028527DC44CFD69FF4929B66249FE2A" blockId="15.[151,1437,150,2014]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="519" pageId="15" pageNumber="518">
|
||
<emphasis id="72E38E6FC44CFD76FF4929B66202F9D7" bold="true" box="[199,365,1518,1544]" pageId="15" pageNumber="518">Comparisons.</emphasis>
|
||
<taxonomicName id="879729FEC44CFD76FEF829B76106F9D6" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[374,617,1519,1545]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44CFD76FEF829B76106F9D6" box="[374,617,1519,1545]" italics="true" pageId="15" pageNumber="518">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44CFD76FDFC29B761A2F9D6" bold="true" box="[626,717,1519,1545]" pageId="15" pageNumber="518">
|
||
<taxonomicNameLabel id="69D03314C44CFD76FDFC29B761A2F9D6" box="[626,717,1519,1545]" pageId="15" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from its most similar congener,
|
||
<taxonomicName id="879729FEC44CFD76FB1B29B7664CF9D6" box="[1173,1315,1519,1545]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="parallela">
|
||
<emphasis id="72E38E6FC44CFD76FB1B29B7664CF9D6" box="[1173,1315,1519,1545]" italics="true" pageId="15" pageNumber="518">M. parallela</emphasis>
|
||
</taxonomicName>
|
||
by having dorsolateral folds extending to the groin (vs. maximum of two-thirds length of trunk:
|
||
<bibRefCitation id="24062F8CC44CFD76FBC42A4B660EF9F2" author="Inger, R. F. & Iskandar, D. T." box="[1098,1377,1554,1581]" pageId="15" pageNumber="518" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC44CFD76FAE22A4B622FF98E" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FF302A6F6392F98E" box="[190,253,1591,1617]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), a rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC44CFD76FC9A2A6F6740F98E" author="Inger, R. F. & Iskandar, D. T." box="[788,1071,1590,1617]" pageId="15" pageNumber="518" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC44CFD76FBB52A6F6664F98E" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1083,1291,1591,1617]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FB062A6F67A8F98E" box="[1160,1223,1591,1617]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), shagreened ventral surface from throat to belly (vs. smooth:
|
||
<bibRefCitation id="24062F8CC44CFD76FD432A036081F9AA" author="Inger, R. F. & Iskandar, D. T." box="[717,1006,1626,1653]" pageId="15" pageNumber="518" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC44CFD76FC732A0367BBF9AA" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1021,1236,1627,1653]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FBC32A0367E1F9AA" box="[1101,1166,1627,1653]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), and a relatively shorter foot to thigh ratio—FL 92.1−96.6% TL (vs. 105.4−112.5% TL:
|
||
<bibRefCitation id="24062F8CC44CFD76FC522A2767C2F946" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[988,1197,1663,1689]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FBA32A27671FF946" box="[1069,1136,1663,1689]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). From
|
||
<emphasis id="72E38E6FC44CFD76FA842A2766F3F947" box="[1290,1436,1663,1688]" italics="true" pageId="15" pageNumber="518">
|
||
<taxonomicName id="879729FEC44CFD76FA842A2766F7F947" box="[1290,1432,1663,1688]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="montana">M. montana</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species differs by having tuberculate dorsal skin surface (vs. smooth:
|
||
<bibRefCitation id="24062F8CC44CFD76FBA32AFB6668F962" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1069,1287,1699,1725]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FBF02AFB67AEF962" box="[1150,1217,1699,1725]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), shagreened skin on throat (vs. smooth:
|
||
<bibRefCitation id="24062F8CC44CFD76FE5C2A9F61CAF93E" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[466,677,1735,1761]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FDAE2A9F610EF93E" box="[544,609,1735,1761]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), a relatively short foot to thigh ratio—FL 92.1−96.6% TL (vs. FL 106.2−115.6% TL:
|
||
<bibRefCitation id="24062F8CC44CFD76FEFD2AB3612DF8DA" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[371,578,1771,1797]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FE4E2AB36290F8DA" box="[448,511,1771,1797]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), and absent toe webbing (vs. rather developed webbing from second to fifth toes:
|
||
<bibRefCitation id="24062F8CC44CFD76FF5D2B5762CDF8F6" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[211,418,1807,1833]" pageId="15" pageNumber="518" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC44CFD76FEAE2B576231F8F6" box="[288,350,1807,1833]" italics="true" pageId="15" pageNumber="518">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). From
|
||
<taxonomicName id="879729FEC44CFD76FE792B57606AF8F6" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[503,773,1807,1833]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="selatanensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44CFD76FE792B57606AF8F6" box="[503,773,1807,1833]" italics="true" pageId="15" pageNumber="518">Megophrys selatanensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44CFD76FC822B57600AF8F6" bold="true" box="[780,869,1807,1833]" pageId="15" pageNumber="518">
|
||
<taxonomicNameLabel id="69D03314C44CFD76FC822B57600AF8F6" box="[780,869,1807,1833]" pageId="15" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="879729FEC44CFD76FCFD2B57670BF8F6" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[883,1124,1807,1833]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="15" pageNumber="518" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC44CFD76FCFD2B57670BF8F6" box="[883,1124,1807,1833]" italics="true" pageId="15" pageNumber="518">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC44CFD76FBE52B5767ABF8F6" bold="true" box="[1131,1220,1807,1833]" pageId="15" pageNumber="518">
|
||
<taxonomicNameLabel id="69D03314C44CFD76FBE52B5767ABF8F6" box="[1131,1220,1807,1833]" pageId="15" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by having a longer rostral appendage—SAL 1.3–1.8% SVLh (vs. SAL 0.3−0.6% SVLh), a slightly longer palpebral projection —EHL 25.1–28.4% UEWh (vs. EHL 21.3–24.3% UEWh), slightly narrower ratio of head width to its length—HW 114.1−115.3% HLh (vs. HW 119.3−126.4% HLh), wider ratio of tympanum to eye diameter—TDH 35.6−37.5% ED (vs. TDH 27.3−28.5% ED), tympanum nearly rounded—TDH 85.9–93.8 TDV (vs. vertically elongated, TDH 46.5−61.9% TDV), slightly shorter lower arms—LAL 44.8–45.8% SVLh (vs. LAL 48.0–49.2% SVLh), tuberculate dorsal skin surface (vs. smooth), shagreened skin on throat (vs. smooth), and absent toe webbing (vs. toes webbed at base).
|
||
<taxonomicName id="879729FEC453FD69FF732CE3629FFF0A" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[253,496,187,213]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FF732CE3629FFF0A" box="[253,496,187,213]" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FE792CE3613EFF0A" bold="true" box="[503,593,187,213]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FE792CE3613EFF0A" box="[503,593,187,213]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="879729FEC453FD69FD692CE3603CFF0A" box="[743,851,187,213]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="lancip">
|
||
<emphasis id="72E38E6FC453FD69FD692CE3603CFF0A" box="[743,851,187,213]" italics="true" pageId="16" pageNumber="519">M. lancip</emphasis>
|
||
</taxonomicName>
|
||
by males having a shorter rostral appendage—SAL 1.3−1.8% SVLh (vs. SAL 2.9−3.1% SVLh in males, but overlapped with females SAL 1.4−4.1% SVLh:
|
||
<bibRefCitation id="24062F8CC453FD69FAB42C87626FFEC2" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FA062C8763D4FEC2" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), shorter head length—HLh 37.2–37.8% SVLh (vs. HLh 42.1–44.6% SVLh;
|
||
<bibRefCitation id="24062F8CC453FD69FBFD2D5B6628FEC2" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1139,1351,259,285]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FB4C2D5B666CFEC2" box="[1218,1283,259,285]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), wider head in males—HW 114.1–115.3% HLh (vs. HW 103.2–105.8% HLh in males, but overlapped with females HW 110.6–116.0% HLh:
|
||
<bibRefCitation id="24062F8CC453FD69FE092D136130FEBA" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[391,607,331,357]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FE592D136176FEBA" box="[471,537,331,357]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), wider tympanum relative to eye diameter—TDH 35.6−37.5% ED (vs. TDH 25.7−28.0 ED:
|
||
<bibRefCitation id="24062F8CC453FD69FE022D37610EFE56" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[396,609,367,393]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FE552D376173FE56" box="[475,540,367,393]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), tympanum nearly rounded—TDH 85.9–93.8 TDV (vs. vertically elongated, TDH 61.1−69.1% TDV), relatively short foot to thigh ratio—FL 92.1−96.6% TL (vs. FL 102.5−109.8% TL;
|
||
<bibRefCitation id="24062F8CC453FD69FF192DEF6209FE0E" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[151,358,439,465]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FF6A2DEF624CFE0E" box="[228,291,439,465]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), and toe webbing absent (vs. rather developed on first to fifth toes, see
|
||
<figureCitation id="D8AC4EF8C453FD69FB022DEF666AFE0E" box="[1164,1285,439,465]" captionStart="FIGURE 4" captionStartId="12.[151,250,1161,1185]" captionTargetBox="[154,1434,181,1137]" captionTargetId="figure-22@12.[154,1434,181,1137]" captionTargetPageId="12" captionText="FIGURE 4. Megophrys acehensis sp. nov. in life, adult male holotype MZB.Amph 26098 (A−B), subadult male paratype UTA A-66178 (C−D)." figureDoi="http://doi.org/10.5281/zenodo.5598867" httpUri="https://zenodo.org/record/5598867/files/figure.png" pageId="16" pageNumber="519">Figs. 4D, I</figureCitation>
|
||
,
|
||
<figureCitation id="D8AC4EF8C453FD69FA9C2DEE665DFE0E" box="[1298,1330,438,465]" captionStart="FIGURE 5" captionStartId="13.[151,250,1908,1932]" captionTargetBox="[204,1418,671,1823]" captionTargetId="figure-228@13.[163,1424,633,1884]" captionTargetPageId="13" captionText="FIGURE 5. Megophrys acehensis sp. nov., adult male holotype (MZB. Amph 26098; SVL 37.4 mm): dorsal (A), ventral (B), lateral head (C), ventral right hand (D), and ventral right foot (E) views, scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5598871" httpUri="https://zenodo.org/record/5598871/files/figure.png" pageId="16" pageNumber="519">5B</figureCitation>
|
||
in
|
||
<bibRefCitation id="24062F8CC453FD69FAD82DEF6275FE2A" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FF192D8363B9FE2A" box="[151,214,475,501]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4028527DC453FD69FF492DA76727FC15" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FF492DA762D6FDC6" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[199,441,511,537]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FF492DA762D6FDC6" box="[199,441,511,537]" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FE4E2DA76175FDC6" bold="true" box="[448,538,511,537]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FE4E2DA76175FDC6" box="[448,538,511,537]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="879729FEC453FD69FD202DA76070FDC7" box="[686,799,511,536]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="nasuta">
|
||
<emphasis id="72E38E6FC453FD69FD202DA76070FDC7" box="[686,799,511,536]" italics="true" pageId="16" pageNumber="519">M. nasuta</emphasis>
|
||
</taxonomicName>
|
||
by the absence of additional lateral flank folds (vs. presence:
|
||
<bibRefCitation id="24062F8CC453FD69FF542E7B62C3FDE2" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[218,428,547,573]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FEA72E7B6206FDE2" box="[297,361,547,573]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="24062F8CC453FD69FE362E7B629BFDE2" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[440,500,547,573]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">2019</bibRefCitation>
|
||
), males being smaller—known adult male
|
||
<quantity id="876FFF98C453FD69FC542E7B6750FDE2" box="[986,1087,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL 66.0–
|
||
<quantity id="876FFF98C453FD69FABE2E7B66F6FDE2" box="[1328,1433,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="9.34" pageId="16" pageNumber="519" unit="mm" value="93.4">93.4 mm</quantity>
|
||
:
|
||
<bibRefCitation id="24062F8CC453FD69FF192E1F6209FDBE" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[151,358,583,609]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FF6A2E1F624CFDBE" box="[228,291,583,609]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), shorter acute rostral appendage—SAL 1.3−1.8% SVLh (vs. acuminate rostral appendage, SAL 1.8−9.2% SVLh:
|
||
<bibRefCitation id="24062F8CC453FD69FED22E336145FD5A" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[348,554,619,645]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FE272E336288FD5A" box="[425,487,619,645]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), and shorter acute palpebral projection—EHL 25.1−28.4% UEWh (vs. acuminate palpebral projection, EHL 32.7−61.4% UEWh:
|
||
<bibRefCitation id="24062F8CC453FD69FD6C2ED760DFFD76" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[738,944,655,681]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FCA02ED76002FD76" box="[814,877,655,681]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
). From
|
||
<emphasis id="72E38E6FC453FD69FB8B2ED76788FD76" box="[1029,1255,655,681]" italics="true" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FB8B2ED7678CFD76" box="[1029,1251,655,681]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="kalimantanensis">M. kalimantanensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species differs by the absence of additional flank folds (vs. present:
|
||
<bibRefCitation id="24062F8CC453FD69FCA12EEB6093FD12" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[815,1020,691,717]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FCF52EEB60D5FD12" box="[891,954,691,717]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), smaller body size in males—known adult male
|
||
<quantity id="876FFF98C453FD69FE982E8F6213FD2E" box="[278,380,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL
|
||
<quantity id="876FFF98C453FD69FDA02E8F618BFD2E" box="[558,740,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="8.245" metricValueMax="10.059999999999999" metricValueMin="6.43" pageId="16" pageNumber="519" unit="mm" value="82.44999999999999" valueMax="100.6" valueMin="64.3">64.3–100.6 mm</quantity>
|
||
:
|
||
<bibRefCitation id="24062F8CC453FD69FD7E2E8F60ABFD2E" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[752,964,727,753]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FCB12E8F6010FD2E" box="[831,895,727,753]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), slightly longer rostral appendage—SAL 1.3−1.8% SVLh (vs. SAL 0.1−0.8% SVLh:
|
||
<bibRefCitation id="24062F8CC453FD69FDF02EA36026FCCA" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[638,841,763,789]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FD442EA36068FCCA" box="[714,775,763,789]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), and shorter palpebral projection—EHL 25.1−28.4% UEWh (vs. 30.9−53.2% UEWh:
|
||
<bibRefCitation id="24062F8CC453FD69FD852F4761B3FCE6" author="Munir, M. & Hamidy, A. & Matsui, M. & Iskandar, D. T. & Sidik, I. & Shimada, T." box="[523,732,799,825]" pageId="16" pageNumber="519" pagination="1 - 24" refId="ref18787" refString="Munir, M., Hamidy, A., Matsui, M., Iskandar, D. T., Sidik, I. & Shimada, T. (2019) A new species of Megophrys Kuhl & van Hasselt (Amphibia: Megophryidae) from Borneo allied to M. nasuta (Schlegel, 1858). Zootaxa, 4679 (1), 1 - 24. https: // doi. org / 10.11646 / zootaxa. 4679.1.1" type="journal article" year="2019">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FDD62F4761F7FCE6" box="[600,664,799,825]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
). From
|
||
<emphasis id="72E38E6FC453FD69FCBD2F4760A1FCE7" box="[819,974,799,824]" italics="true" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FCBD2F4760A5FCE7" box="[819,970,799,824]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="stejnegeri">M. stejnegeri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species differs by having dorsolateral folds extending to the groin (vs. maximum of two-thirds the length of the trunk:
|
||
<bibRefCitation id="24062F8CC453FD69FB0E2F1B6790FC82" author="Inger, R. F." box="[1152,1279,834,861]" pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">Inger 1954</bibRefCitation>
|
||
), males being smaller—known adult male
|
||
<quantity id="876FFF98C453FD69FE532F3F612CFC5E" box="[477,579,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. adult male 40.6−60.0 mm:
|
||
<bibRefCitation id="24062F8CC453FD69FC7B2F3F67E7FC5E" author="Taylor, E. H." box="[1013,1160,871,897]" pageId="16" pageNumber="519" pagination="213 - 359" refId="ref20101" refString="Taylor, E. H. (1920) Philippine Amphibia. The Philippine Journal of Science, 16 (3), 213 - 359. https: // doi. org / 10.5962 / bhl. part. 4751" type="journal article" year="1920">Taylor, 1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FB182F3F6672FC5E" author="Inger, R. F." box="[1174,1309,870,897]" pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">Inger, 1954</bibRefCitation>
|
||
), vomerine teeth present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FE212FD36150FC7A" author="Taylor, E. H." box="[431,575,907,933]" pageId="16" pageNumber="519" pagination="213 - 359" refId="ref20101" refString="Taylor, E. H. (1920) Philippine Amphibia. The Philippine Journal of Science, 16 (3), 213 - 359. https: // doi. org / 10.5962 / bhl. part. 4751" type="journal article" year="1920">Taylor, 1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FDC42FD36063FC7A" author="Inger, R. F." box="[586,780,906,933]" pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FD052FD361A5FC7A" box="[651,714,907,933]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1954
|
||
</bibRefCitation>
|
||
), and smooth skin on the posterior of tympanum (vs. tuberculate, see figure 37 by
|
||
<bibRefCitation id="24062F8CC453FD69FE2D2FF761E5FC16" author="Diesmos, A. C. & Watters, J. L. & Huron, N. A. & Davis, D. R. & Alcala, A. C. & Crombie R. I. & Afuang, L. E. & Sison, R. V. & Sanguila, M. B. & Penrod, M. L. & Labonte, M. J. & Davey C. S. & Leone E. A. & Diesmos M. L. & Sy, M. Y. & Welton, L. J. & Brown R. M. & Siler C. D." box="[419,650,942,969]" pageId="16" pageNumber="519" pagination="457 - 539" refId="ref15919" refString="Diesmos, A. C., Watters, J. L., Huron, N. A., Davis, D. R., Alcala, A. C., Crombie R. I., Afuang, L. E., Gee-das, G., Sison, R. V., Sanguila, M. B., Penrod, M. L., Labonte, M. J., Davey C. S., Leone E. A., Diesmos M. L., Sy, M. Y., Welton, L. J., Brown R. M. & Siler C. D. (2015) Amphibians of the Philippines, Part I: Checklist of the Species. Proceedings of the California Academy of Science, 62 (20), 457 - 539." type="journal article" year="2015">
|
||
Diesmos
|
||
<emphasis id="72E38E6FC453FD69FD822FF76124FC16" box="[524,587,943,969]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
and figure 18 by
|
||
<bibRefCitation id="24062F8CC453FD69FCDE2FF76753FC15" author="Sanguila, M. B. & Cobb, K. & Siller C. D. & Diesmos, A. C. & Alcala A. C. & Brown R. M." box="[848,1084,943,970]" pageId="16" pageNumber="519" pagination="1 - 32" refId="ref19568" refString="Sanguila, M. B., Cobb, K. A, Siller C. D., Diesmos, A. C., Alcala A. C. & Brown R. M. (2016) The amphibians and reptiles of Mindanao Island, southern Philippines, II: the herpetofauna of northeast Mindanao and adjacent islands. ZooKeys, 624, 1 - 32. https: // doi. org / 10.3897 / zookeys. 624.9814" type="journal article" year="2016">
|
||
Sanguila
|
||
<emphasis id="72E38E6FC453FD69FC372FF76097FC16" box="[953,1016,943,969]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4028527DC453FD69FF492F8B638FF98E" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FF492F8B62D3FC32" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[199,444,979,1005]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FF492F8B62D3FC32" box="[199,444,979,1005]" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FE482F8B614DFC32" bold="true" box="[454,546,979,1005]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FE482F8B614DFC32" box="[454,546,979,1005]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="879729FEC453FD69FCEF2F8B676DFC32" box="[865,1026,979,1005]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="kobayashii">
|
||
<emphasis id="72E38E6FC453FD69FCEF2F8B676DFC32" box="[865,1026,979,1005]" italics="true" pageId="16" pageNumber="519">M. kobayashii</emphasis>
|
||
</taxonomicName>
|
||
by having
|
||
<typeStatus id="9F2CECDFC453FD69FB072F8B67D6FC32" box="[1161,1209,979,1005]" pageId="16" pageNumber="519">type</typeStatus>
|
||
I dorsolateral folds (vs.
|
||
<typeStatus id="9F2CECDFC453FD69FF472FAF6396FBCE" box="[201,249,1015,1041]" pageId="16" pageNumber="519">type</typeStatus>
|
||
II, see
|
||
<tableCitation id="0D1567C6C453FD69FEDF2FAF62C8FBCE" box="[337,423,1015,1041]" captionStart="TABLE 3" captionStartId="14.[151,239,648,674]" captionTargetPageId="14" captionText="TABLE 3. Morphological variation between Sunda Shelf and Philippine Megophrys (modified from Munir et al., 2018)." httpUri="http://table.plazi.org/id/14E802F5C44DFD77FF192ED066F0FD7D" pageId="16" pageNumber="519" tableUuid="14E802F5C44DFD77FF192ED066F0FD7D">Table 3</tableCitation>
|
||
), absent additional lateral flank folds (vs. present:
|
||
<bibRefCitation id="24062F8CC453FD69FC612FAF6644FBCE" author="Malkmus, R. & Matsui, M." box="[1007,1323,1015,1041]" pageId="16" pageNumber="519" pagination="31 - 37" refId="ref18635" refString="Malkmus, R. & Matsui, M. (1997) Megophrys kobayashii, ein neuer pelobatider frosch vom Mount Kinabalu, Sauria, 19, 31 - 37." type="journal article" year="1997">Malkmus and Matsui, 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FAB72FAF6391FBEA" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FA062FAF63D4FBEA" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), males being smaller—known adult male
|
||
<quantity id="876FFF98C453FD69FD5228436050FBEA" box="[732,831,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. 93.0−101.0 mm:
|
||
<bibRefCitation id="24062F8CC453FD69FBFF284366FFFBEA" author="Inger, R. F. & Stuebing, R. B." box="[1137,1424,1050,1077]" pageId="16" pageNumber="519" refId="ref17466" refString="Inger, R. F. & Stuebing, R. B. (2005) A field guide to the frogs of Borneo. 2 nd Edition. Natural History Publication Borneo Sdn. Bhd. Kota Kinabalu, Sabah, 201 pp." type="book" year="2005">Inger and Stuebing, 2005</bibRefCitation>
|
||
), rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FDC8286760EFFB86" author="Malkmus, R. & Matsui, M." box="[582,896,1087,1113]" pageId="16" pageNumber="519" pagination="31 - 37" refId="ref18635" refString="Malkmus, R. & Matsui, M. (1997) Megophrys kobayashii, ein neuer pelobatider frosch vom Mount Kinabalu, Sauria, 19, 31 - 37." type="journal article" year="1997">Malkmus and Matsui, 1997</bibRefCitation>
|
||
), and smaller and fewer tubercles on flanks (vs. larger and numerous:
|
||
<bibRefCitation id="24062F8CC453FD69FE03283B61A8FBA2" author="Malkmus, R. & Matsui, M." box="[397,711,1123,1149]" pageId="16" pageNumber="519" pagination="31 - 37" refId="ref18635" refString="Malkmus, R. & Matsui, M. (1997) Megophrys kobayashii, ein neuer pelobatider frosch vom Mount Kinabalu, Sauria, 19, 31 - 37." type="journal article" year="1997">Malkmus and Matsui, 1997</bibRefCitation>
|
||
). From
|
||
<taxonomicName id="879729FEC453FD69FC91283B60F8FBA2" authorityName="Taylor" authorityYear="1920" box="[799,919,1123,1149]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="ligayae">
|
||
<emphasis id="72E38E6FC453FD69FC91283B60F8FBA2" box="[799,919,1123,1149]" italics="true" pageId="16" pageNumber="519">M. ligayae</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs by having
|
||
<typeStatus id="9F2CECDFC453FD69FAAB283B663AFBA2" box="[1317,1365,1123,1149]" pageId="16" pageNumber="519">type</typeStatus>
|
||
I dorsolateral folds, extended to the groin (vs.
|
||
<typeStatus id="9F2CECDFC453FD69FDE228DF61F3FB7E" box="[620,668,1159,1185]" pageId="16" pageNumber="519">type</typeStatus>
|
||
II, reaching a maximum of two-thirds of trunk:
|
||
<bibRefCitation id="24062F8CC453FD69FB3028DF6620FB7E" author="Taylor, E. H." box="[1214,1359,1159,1185]" pageId="16" pageNumber="519" pagination="213 - 359" refId="ref20101" refString="Taylor, E. H. (1920) Philippine Amphibia. The Philippine Journal of Science, 16 (3), 213 - 359. https: // doi. org / 10.5962 / bhl. part. 4751" type="journal article" year="1920">Taylor, 1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FAD528DF63BDFB1A" author="Inger, R. F." pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">Inger, 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FF5328F362C5FB1A" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[221,426,1195,1221]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FEA428F36207FB1A" box="[298,360,1195,1221]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), absent additional lateral folds (vs. present:
|
||
<bibRefCitation id="24062F8CC453FD69FC1428F36745FB1A" author="Taylor, E. H." box="[922,1066,1195,1221]" pageId="16" pageNumber="519" pagination="213 - 359" refId="ref20101" refString="Taylor, E. H. (1920) Philippine Amphibia. The Philippine Journal of Science, 16 (3), 213 - 359. https: // doi. org / 10.5962 / bhl. part. 4751" type="journal article" year="1920">Taylor, 1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FBBB28F367D7FB1A" author="Inger, R. F." box="[1077,1208,1194,1221]" pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">Inger, 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FB4D28F366FFFB1A" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[1219,1424,1195,1221]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FA8128F36621FB1A" box="[1295,1358,1195,1221]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), males being smaller—known adult male
|
||
<quantity id="876FFF98C453FD69FDE5289761BEFB36" box="[619,721,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL 60.4−69.0 mm:
|
||
<bibRefCitation id="24062F8CC453FD69FBB4289767A3FB36" author="Taylor, E. H." box="[1082,1228,1231,1257]" pageId="16" pageNumber="519" pagination="213 - 359" refId="ref20101" refString="Taylor, E. H. (1920) Philippine Amphibia. The Philippine Journal of Science, 16 (3), 213 - 359. https: // doi. org / 10.5962 / bhl. part. 4751" type="journal article" year="1920">Taylor, 1920</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FB5728976630FB36" author="Inger, R. F." box="[1241,1375,1230,1257]" pageId="16" pageNumber="519" pagination="183 - 531" refId="ref17254" refString="Inger, R. F. (1954) The systematics and zoogeography of Philippine Amphibia. Fieldiana Zoology, 33 (4), 183 - 531. https: // doi. org / 10.5962 / bhl. title. 5571" type="journal article" year="1954">Inger, 1954</bibRefCitation>
|
||
), and toe webbing absent (vs. rather developed web on third, fourth, and fifth toes:
|
||
<bibRefCitation id="24062F8CC453FD69FB9928AB6665FAD2" author="Diesmos, A. C. & Watters, J. L. & Huron, N. A. & Davis, D. R. & Alcala, A. C. & Crombie R. I. & Afuang, L. E. & Sison, R. V. & Sanguila, M. B. & Penrod, M. L. & Labonte, M. J. & Davey C. S. & Leone E. A. & Diesmos M. L. & Sy, M. Y. & Welton, L. J. & Brown R. M. & Siler C. D." box="[1047,1290,1266,1293]" pageId="16" pageNumber="519" pagination="457 - 539" refId="ref15919" refString="Diesmos, A. C., Watters, J. L., Huron, N. A., Davis, D. R., Alcala, A. C., Crombie R. I., Afuang, L. E., Gee-das, G., Sison, R. V., Sanguila, M. B., Penrod, M. L., Labonte, M. J., Davey C. S., Leone E. A., Diesmos M. L., Sy, M. Y., Welton, L. J., Brown R. M. & Siler C. D. (2015) Amphibians of the Philippines, Part I: Checklist of the Species. Proceedings of the California Academy of Science, 62 (20), 457 - 539." type="journal article" year="2015">
|
||
Diesmos
|
||
<emphasis id="72E38E6FC453FD69FB0C28AB67ACFAD2" box="[1154,1219,1267,1293]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="879729FEC453FD69FA9028AB626BFAEE" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FA9028AB626BFAEE" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FE85294F620BFAEE" bold="true" box="[267,356,1303,1329]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FE85294F620BFAEE" box="[267,356,1303,1329]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="879729FEC453FD69FE79294F61F0FAEE" authorityName="Inger" authorityYear="1989" box="[503,671,1303,1329]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="edwardinae">
|
||
<emphasis id="72E38E6FC453FD69FE79294F61F0FAEE" box="[503,671,1303,1329]" italics="true" pageId="16" pageNumber="519">M. edwardinae</emphasis>
|
||
</taxonomicName>
|
||
by having
|
||
<typeStatus id="9F2CECDFC453FD69FC95294F6024FAEE" box="[795,843,1303,1329]" pageId="16" pageNumber="519">type</typeStatus>
|
||
I dorsolateral folds (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FB3F294F6640FAEE" author="Inger, R. F." box="[1201,1327,1303,1329]" pageId="16" pageNumber="519" pagination="229 - 243" refId="ref17339" refString="Inger, R. F. (1989) Four new species of frogs from Borneo. Malayan Nature Journal, 42, 229 - 243." type="journal article" year="1989">Inger 1989</bibRefCitation>
|
||
), a rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FE7329636112FA8A" author="Inger, R. F." box="[509,637,1339,1365]" pageId="16" pageNumber="519" pagination="229 - 243" refId="ref17339" refString="Inger, R. F. (1989) Four new species of frogs from Borneo. Malayan Nature Journal, 42, 229 - 243." type="journal article" year="1989">Inger 1989</bibRefCitation>
|
||
), vomerine teeth present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FBA4296367C5FA8A" author="Inger, R. F." box="[1066,1194,1339,1365]" pageId="16" pageNumber="519" pagination="229 - 243" refId="ref17339" refString="Inger, R. F. (1989) Four new species of frogs from Borneo. Malayan Nature Journal, 42, 229 - 243." type="journal article" year="1989">Inger 1989</bibRefCitation>
|
||
), and slightly shorter thighs—TL 40.4−41.8% SVLh (vs. TL 45.0−50.0% SVL:
|
||
<bibRefCitation id="24062F8CC453FD69FCAE290760F1FAA6" author="Inger, R. F." box="[800,926,1375,1401]" pageId="16" pageNumber="519" pagination="229 - 243" refId="ref17339" refString="Inger, R. F. (1989) Four new species of frogs from Borneo. Malayan Nature Journal, 42, 229 - 243." type="journal article" year="1989">Inger 1989</bibRefCitation>
|
||
).
|
||
<taxonomicName id="879729FEC453FD69FC3E290767CDFAA6" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[944,1186,1375,1401]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FC3E290767CDFAA6" box="[944,1186,1375,1401]" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FB262907666EFAA6" bold="true" box="[1192,1281,1375,1401]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FB262907666EFAA6" box="[1192,1281,1375,1401]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="879729FEC453FD69FEBC29DB62AFFA42" box="[306,448,1411,1437]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="72E38E6FC453FD69FEBC29DB62AFFA42" box="[306,448,1411,1437]" italics="true" pageId="16" pageNumber="519">M. baluensis</emphasis>
|
||
</taxonomicName>
|
||
by having
|
||
<typeStatus id="9F2CECDFC453FD69FDB229DB6103FA42" box="[572,620,1411,1437]" pageId="16" pageNumber="519">type</typeStatus>
|
||
I dorsolateral folds (vs.
|
||
<typeStatus id="9F2CECDFC453FD69FCF529DB60C4FA42" box="[891,939,1411,1437]" pageId="16" pageNumber="519">type</typeStatus>
|
||
III:
|
||
<bibRefCitation id="24062F8CC453FD69FC5229DB67C6FA42" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[988,1193,1411,1437]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FBA629DB6708FA42" box="[1064,1127,1411,1437]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
), males being slightly smaller—known adult male
|
||
<quantity id="876FFF98C453FD69FE5A29FF6157FA1E" box="[468,568,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL 41.0−45.0 mm:
|
||
<bibRefCitation id="24062F8CC453FD69FC1829FF6737FA1E" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[918,1112,1446,1473]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FC5929FF6779FA1E" box="[983,1046,1447,1473]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), a rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FE93299362B6FA3A" author="Boulenger, G. A." box="[285,473,1483,1509]" pageId="16" pageNumber="519" pagination="451 - 454" refId="ref15237" refString="Boulenger, G. A. (1899) Description of three new reptiles and a new batrachian from Mount Kinabalu, North Borneo. Annals and Magazine of Natural History, 7 (4), 451 - 454. https: // doi. org / 10.1080 / 00222939908678228" type="journal article" year="1899">Boulenger, 1899</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FE6A29936109FA3A" author="Inger, R. F." box="[484,614,1483,1509]" pageId="16" pageNumber="519" pagination="1 - 402" refId="ref17297" refString="Inger, R. F. (1966) The systematic and zoogeography of the amphibians of Borneo. Fieldiana Zoology, 52, 1 - 402. https: // doi. org / 10.5962 / bhl. title. 3147" type="journal article" year="1966">Inger, 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FDFF29936043FA3A" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[625,812,1482,1509]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FD3D2993619EFA3A" box="[691,753,1483,1509]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), having a triangular palpebral projection (vs. small like tubercle:
|
||
<bibRefCitation id="24062F8CC453FD69FE8E29B762ECF9D6" author="Inger, R. F." box="[256,387,1519,1545]" pageId="16" pageNumber="519" pagination="229 - 243" refId="ref17339" refString="Inger, R. F. (1989) Four new species of frogs from Borneo. Malayan Nature Journal, 42, 229 - 243." type="journal article" year="1989">Inger, 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FE0129B7613CF9D6" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[399,595,1518,1545]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FE5F29B7617EF9D6" box="[465,529,1519,1545]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), having smaller and fewer tubercles on body flanks (vs. larger and numerous:
|
||
<bibRefCitation id="24062F8CC453FD69FF412A4B62F6F9F2" author="Inger, R. F. & Stuebing, R. B. & Grafe, T. U. & Dehling, J. M." box="[207,409,1555,1581]" pageId="16" pageNumber="519" refId="ref17511" refString="Inger, R. F., Stuebing, R. B., Grafe, T. U. & Dehling, J. M. (2017). A field guide to the Frogs of Borneo. Third edition. Natural History Publication Borneo. Kinabalu, 236 pp." type="book" year="2017">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FE9D2A4B623BF9F2" box="[275,340,1555,1581]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
), and adpressed tibiotarsal articulation reaching posterior of eye (vs. shoulder:
|
||
<bibRefCitation id="24062F8CC453FD69FAAC2A4B63BCF98E" author="Boulenger, G. A." pageId="16" pageNumber="519" pagination="451 - 454" refId="ref15237" refString="Boulenger, G. A. (1899) Description of three new reptiles and a new batrachian from Mount Kinabalu, North Borneo. Annals and Magazine of Natural History, 7 (4), 451 - 454. https: // doi. org / 10.1080 / 00222939908678228" type="journal article" year="1899">Boulenger, 1899</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4028527DC453FD68FF492A03612CFE9E" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,150,681]" lastPageId="17" lastPageNumber="520" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FF492A0362D6F9AA" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[199,441,1627,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC453FD69FF492A0362D6F9AA" box="[199,441,1627,1653]" italics="true" pageId="16" pageNumber="519">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC453FD69FE4F2A036175F9AA" bold="true" box="[449,538,1627,1653]" pageId="16" pageNumber="519">
|
||
<taxonomicNameLabel id="69D03314C453FD69FE4F2A036175F9AA" box="[449,538,1627,1653]" pageId="16" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="879729FEC453FD69FCC22A0360DAF9AA" box="[844,949,1627,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="72E38E6FC453FD69FCC22A0360DAF9AA" box="[844,949,1627,1653]" italics="true" pageId="16" pageNumber="519">M. dringi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="879729FEC453FD69FC4A2A03675AF9AB" authorityName="Boulenger" authorityYear="1903" box="[964,1077,1627,1652]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="aceras">
|
||
<emphasis id="72E38E6FC453FD69FC4A2A03675AF9AB" box="[964,1077,1627,1652]" italics="true" pageId="16" pageNumber="519">M. aceras</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="879729FEC453FD69FBE52A03679FF9AA" authorityName="Boulenger" authorityYear="1886" box="[1131,1264,1627,1653]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="longipes">
|
||
<emphasis id="72E38E6FC453FD69FBE52A03679FF9AA" box="[1131,1264,1627,1653]" italics="true" pageId="16" pageNumber="519">M. longipes</emphasis>
|
||
</taxonomicName>
|
||
by the absence of Y, X,>–<or H shaped folds on dorsal (vs. presence:
|
||
<bibRefCitation id="24062F8CC453FD69FCAE2A2760B1F946" author="Boulenger, G. A." box="[800,990,1662,1689]" pageId="16" pageNumber="519" pagination="850 - 850" refId="ref15178" refString="Boulenger, G. A. (1885) Description of a new frog of the genus Megalophrys. Proceedings of the scientific meeting of the Zoological Society of London, 53 (4), 850 - 850. https: // doi. org / 10.1111 / j. 1096 - 3642.1885. tb 02927. x" type="journal article" year="1885">Boulenger, 1885</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="24062F8CC453FD69FC652A276749F946" author="Boulenger, G. A." box="[1003,1062,1663,1689]" pageId="16" pageNumber="519" pagination="131 - 176" refId="ref15288" refString="Boulenger, G. A. (1903) Report on the batrachians and reptiles. In: Annandale, N. & Robinson, H. C. (Eds.), Fasciculi Malayenses. Zoology. Part I. Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States 1901 - 1902. Longmans, Green & Co, London, pp. 131 - 176." type="book chapter" year="1903">1903</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC453FD69FBBA2A276791F946" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[1076,1278,1662,1689]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FBF62A2767D5F946" box="[1144,1210,1663,1689]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
, see
|
||
<tableCitation id="0D1567C6C453FD69FAB92A2766E1F946" box="[1335,1422,1663,1689]" captionStart="TABLE 3" captionStartId="14.[151,239,648,674]" captionTargetPageId="14" captionText="TABLE 3. Morphological variation between Sunda Shelf and Philippine Megophrys (modified from Munir et al., 2018)." httpUri="http://table.plazi.org/id/14E802F5C44DFD77FF192ED066F0FD7D" pageId="16" pageNumber="519" tableUuid="14E802F5C44DFD77FF192ED066F0FD7D">Table 3</tableCitation>
|
||
). Furthermore, from
|
||
<emphasis id="72E38E6FC453FD69FEFF2AFB628CF962" box="[369,483,1699,1725]" italics="true" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FEFF2AFB62B0F962" box="[369,479,1699,1725]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="dringi">M. dringi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species being smaller in male—known adult male
|
||
<quantity id="876FFF98C453FD69FB032AFB679DF962" box="[1165,1266,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL 43.0−47.0 mm:
|
||
<bibRefCitation id="24062F8CC453FD69FEC62A9F6167F93E" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[328,520,1734,1761]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FE072A9F62A8F93E" box="[393,455,1735,1761]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), having a stocky body (vs. slender:
|
||
<bibRefCitation id="24062F8CC453FD69FC152A9F6734F93E" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[923,1115,1734,1761]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FC522A9F6775F93E" box="[988,1050,1735,1761]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), a rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FE902AB3628CF8DA" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[286,483,1770,1797]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FEEF2AB362CFF8DA" box="[353,416,1771,1797]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), having a triangular palpebral projection (vs. small like tubercle:
|
||
<bibRefCitation id="24062F8CC453FD69FB452AB366E0F8DA" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[1227,1423,1770,1797]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FA832AB36622F8DA" box="[1293,1357,1771,1797]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), vomerine teeth present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FD912B5761B0F8F6" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[543,735,1806,1833]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FDEE2B5761F1F8F6" box="[608,670,1807,1833]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
), distinct tympanum (vs. partially obscured:
|
||
<bibRefCitation id="24062F8CC453FD69FB412B5766FFF8F6" author="Inger, R. F. & Stuebing, R. B. & Tan, F. L." box="[1231,1424,1806,1833]" pageId="16" pageNumber="519" pagination="115 - 131" refId="ref17366" refString="Inger, R. F., Stuebing, R. B. & Tan, F. L. (1995) New species and new records of anurans from Borneo. The Raffles Bulletin of Zoology, 43 (1), 115 - 131." type="journal article" year="1995">
|
||
Inger
|
||
<emphasis id="72E38E6FC453FD69FA9E2B576621F8F6" box="[1296,1358,1807,1833]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
). From
|
||
<emphasis id="72E38E6FC453FD69FF532B6B623AF893" box="[221,341,1843,1868]" italics="true" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FF532B6B623EF893" authorityName="Boulenger" authorityYear="1903" box="[221,337,1843,1868]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="aceras">M. aceras</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species differs by having smaller body size in male—known adult male
|
||
<quantity id="876FFF98C453FD69FB742B6B660FF892" box="[1274,1376,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="16" pageNumber="519" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL 48.0−
|
||
<quantity id="876FFF98C453FD69FECF2B0F62C7F8AE" box="[321,424,1879,1905]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" pageId="16" pageNumber="519" unit="mm" value="62.4">62.4 mm</quantity>
|
||
:
|
||
<bibRefCitation id="24062F8CC453FD69FE3F2B0F61A6F8AE" author="Inger, R. F. & Iskandar, D. T." box="[433,713,1878,1905]" pageId="16" pageNumber="519" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
), a stocky body (vs. slender:
|
||
<bibRefCitation id="24062F8CC453FD69FB862B0F67ABF8AE" author="Boulenger, G. A." box="[1032,1220,1879,1905]" pageId="16" pageNumber="519" pagination="131 - 176" refId="ref15288" refString="Boulenger, G. A. (1903) Report on the batrachians and reptiles. In: Annandale, N. & Robinson, H. C. (Eds.), Fasciculi Malayenses. Zoology. Part I. Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States 1901 - 1902. Longmans, Green & Co, London, pp. 131 - 176." type="book chapter" year="1903">Boulenger, 1903</bibRefCitation>
|
||
), rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC453FD69FEFD2B236140F84A" author="Boulenger, G. A." box="[371,559,1915,1941]" pageId="16" pageNumber="519" pagination="131 - 176" refId="ref15288" refString="Boulenger, G. A. (1903) Report on the batrachians and reptiles. In: Annandale, N. & Robinson, H. C. (Eds.), Fasciculi Malayenses. Zoology. Part I. Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States 1901 - 1902. Longmans, Green & Co, London, pp. 131 - 176." type="book chapter" year="1903">Boulenger, 1903</bibRefCitation>
|
||
), tibiotarsal articulation reaching to posterior corner of eye (vs. shoulder, angle of jaws or temporal area:
|
||
<bibRefCitation id="24062F8CC453FD69FE372BC76124F865" author="Taylor, E. H." box="[441,587,1951,1978]" pageId="16" pageNumber="519" pagination="265 - 599" refId="ref20142" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. The University of Kansas Science Bulletin, 43 (8), 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor, 1962</bibRefCitation>
|
||
), toes web absent (vs. rather developed web on third, fourth, and fifth toes:
|
||
<bibRefCitation id="24062F8CC453FD69FF192B9B6248F802" author="Taylor, E. H." box="[151,295,1987,2013]" pageId="16" pageNumber="519" pagination="265 - 599" refId="ref20142" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. The University of Kansas Science Bulletin, 43 (8), 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor, 1962</bibRefCitation>
|
||
, see figure
|
||
<quantity id="876FFF98C453FD69FE2D2B9A62A2F802" box="[419,461,1986,2013]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="16" pageNumber="519" unit="in" value="5.0">5 in</quantity>
|
||
<bibRefCitation id="24062F8CC453FD69FE5D2B9B61CFF802" author="Munir, M. & Hamidy, A. & Farajallah, A. & Smith, E. N." box="[467,672,1987,2013]" pageId="16" pageNumber="519" pagination="389 - 412" refId="ref18722" refString="Munir, M., Hamidy, A., Farajallah, A. & Smith, E. N. (2018) A new Megophrys Kuhl and van Hasselt (Amphibia: Megophryidae) from southwestern Sumatra, Indonesia. Zootaxa, 4442 (3), 389 - 412. https: // doi. org / 10.11646 / zootaxa. 4442.3.3" type="journal article" year="2018">
|
||
Munir
|
||
<emphasis id="72E38E6FC453FD69FD912B9B6132F802" box="[543,605,1987,2013]" italics="true" pageId="16" pageNumber="519">et al.,</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). From
|
||
<emphasis id="72E38E6FC453FD69FD7D2B9B6012F802" box="[755,893,1987,2013]" italics="true" pageId="16" pageNumber="519">
|
||
<taxonomicName id="879729FEC453FD69FD7D2B9B6015F802" authorityName="Boulenger" authorityYear="1886" box="[755,890,1987,2013]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="16" pageNumber="519" phylum="Chordata" rank="species" species="longipes">M. longipes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the new species differs by having a smaller body size in male—known adult male
|
||
<quantity id="876FFF98C452FD68FD8B2CCF6107FF6E" box="[517,616,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="3.7399999999999998" pageId="17" pageNumber="520" unit="mm" value="37.4">37.4 mm</quantity>
|
||
SVL (vs. SVL
|
||
<quantity id="876FFF98C452FD68FC9F2CCE60D6FF6E" box="[785,953,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.205" metricValueMax="4.52" metricValueMin="3.8899999999999997" pageId="17" pageNumber="520" unit="mm" value="42.05" valueMax="45.2" valueMin="38.9">38.9−45.2 mm</quantity>
|
||
:
|
||
<bibRefCitation id="24062F8CC452FD68FC4D2CCF67B4FF6E" author="Inger, R. F. & Iskandar, D. T." box="[963,1243,150,177]" pageId="17" pageNumber="520" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
), having a stocky body (vs. slender:
|
||
<bibRefCitation id="24062F8CC452FD68FEE82CE3614CFF0A" author="Boulenger, G. A." box="[358,547,186,213]" pageId="17" pageNumber="520" pagination="850 - 850" refId="ref15178" refString="Boulenger, G. A. (1885) Description of a new frog of the genus Megalophrys. Proceedings of the scientific meeting of the Zoological Society of London, 53 (4), 850 - 850. https: // doi. org / 10.1111 / j. 1096 - 3642.1885. tb 02927. x" type="journal article" year="1885">Boulenger, 1885</bibRefCitation>
|
||
), a rostral appendage present (vs. absent:
|
||
<bibRefCitation id="24062F8CC452FD68FC792CE367DBFF0A" author="Boulenger, G. A." box="[1015,1204,186,213]" pageId="17" pageNumber="520" pagination="850 - 850" refId="ref15178" refString="Boulenger, G. A. (1885) Description of a new frog of the genus Megalophrys. Proceedings of the scientific meeting of the Zoological Society of London, 53 (4), 850 - 850. https: // doi. org / 10.1111 / j. 1096 - 3642.1885. tb 02927. x" type="journal article" year="1885">Boulenger, 1885</bibRefCitation>
|
||
), having a triangular palpebral projection (vs. small like tubercle:
|
||
<bibRefCitation id="24062F8CC452FD68FD092C87602CFF26" author="Boulenger, G. A." box="[647,835,222,249]" pageId="17" pageNumber="520" pagination="850 - 850" refId="ref15178" refString="Boulenger, G. A. (1885) Description of a new frog of the genus Megalophrys. Proceedings of the scientific meeting of the Zoological Society of London, 53 (4), 850 - 850. https: // doi. org / 10.1111 / j. 1096 - 3642.1885. tb 02927. x" type="journal article" year="1885">Boulenger, 1885</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="24062F8CC452FD68FCC32C8760B1FF25" author="Taylor, E. H." box="[845,990,223,250]" pageId="17" pageNumber="520" pagination="265 - 599" refId="ref20142" refString="Taylor, E. H. (1962) The amphibian fauna of Thailand. The University of Kansas Science Bulletin, 43 (8), 265 - 599. https: // doi. org / 10.5962 / bhl. part. 13347" type="journal article" year="1962">Taylor, 1962</bibRefCitation>
|
||
), shorter thigh—TL 0.41−0.42 SVL (vs. TL 0.55−0.60 SVL:
|
||
<bibRefCitation id="24062F8CC452FD68FEF12D5B61F4FEC2" author="Inger, R. F. & Iskandar, D. T." box="[383,667,258,285]" pageId="17" pageNumber="520" pagination="133 - 142" refId="ref17414" refString="Inger, R. F. & Iskandar, D. T. (2005) A collection of amphibians from West Sumatra, with description of new species of Megophrys (Amphibia: Anura). The Raffles Bulletin of Zoology, 53 (1), 133 - 142." type="journal article" year="2005">Inger and Iskandar, 2005</bibRefCitation>
|
||
) and tibiotarsal articulation reaching posterior corner of eye (vs. far beyond tip of snout;
|
||
<bibRefCitation id="24062F8CC452FD68FEF12D7F6158FE9E" author="Boulenger, G. A." box="[383,567,294,321]" pageId="17" pageNumber="520" pagination="850 - 850" refId="ref15178" refString="Boulenger, G. A. (1885) Description of a new frog of the genus Megalophrys. Proceedings of the scientific meeting of the Zoological Society of London, 53 (4), 850 - 850. https: // doi. org / 10.1111 / j. 1096 - 3642.1885. tb 02927. x" type="journal article" year="1885">Boulenger 1885</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="088D01F6C452FD68FF492D12600AFD76" pageId="17" pageNumber="520" type="distribution">
|
||
<paragraph id="4028527DC452FD68FF492D12600AFD76" blockId="17.[151,1437,150,681]" pageId="17" pageNumber="520">
|
||
<emphasis id="72E38E6FC452FD68FF492D126134FEBB" bold="true" box="[199,603,330,357]" pageId="17" pageNumber="520">Distribution and Natural History.</emphasis>
|
||
The
|
||
<typeStatus id="9F2CECDFC452FD68FD1B2D136198FEBA" box="[661,759,331,357]" pageId="17" pageNumber="520" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="879729FEC452FD68FC902D13677EFEBA" authority="Munir & Nishikawa & Hamidy & Smith, 2021" authorityName="Munir & Nishikawa & Hamidy & Smith" authorityYear="2021" box="[798,1041,331,357]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="acehensis" status="sp. nov.">
|
||
<emphasis id="72E38E6FC452FD68FC902D13677EFEBA" box="[798,1041,331,357]" italics="true" pageId="17" pageNumber="520">Megophrys acehensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="72E38E6FC452FD68FB962D13671DFEBA" bold="true" box="[1048,1138,331,357]" pageId="17" pageNumber="520">
|
||
<taxonomicNameLabel id="69D03314C452FD68FB962D13671DFEBA" box="[1048,1138,331,357]" pageId="17" pageNumber="520" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was collected on leaf litter in a sloping area at the edge of a primary forest near a stream, while the
|
||
<typeStatus id="9F2CECDFC452FD68FC4C2D37674DFE56" box="[962,1058,367,393]" pageId="17" pageNumber="520" type="paratype">paratype</typeStatus>
|
||
was collected from leaf litter in a palm oil plantation near the edge of an old secondary forest. Landslides, new road development, and monoculture forests have become major threats at the
|
||
<typeStatus id="9F2CECDFC452FD68FDE72DEF61A4FE0E" box="[617,715,439,465]" pageId="17" pageNumber="520" type="holotype">holotype</typeStatus>
|
||
locality, while the threats at the
|
||
<typeStatus id="9F2CECDFC452FD68FBB12DEF67F0FE0E" box="[1087,1183,439,465]" pageId="17" pageNumber="520" type="paratype">paratype</typeStatus>
|
||
locality were the land use changes and water pollution from palm oil fields. The precise distribution, population, habitat requirements, breeding behavior, call and tadpole information are unknown. The following anuran species have been found sympatrically with the new species, at
|
||
<typeStatus id="9F2CECDFC452FD68FD9A2E7B6119FDE2" box="[532,630,547,573]" pageId="17" pageNumber="520" type="holotype">holotype</typeStatus>
|
||
locality:
|
||
<taxonomicName id="879729FEC452FD68FD502E7B60E1FDE2" box="[734,910,547,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="72E38E6FC452FD68FD502E7B6003FDE3" box="[734,876,547,572]" italics="true" pageId="17" pageNumber="520">Limnonectes</emphasis>
|
||
sp
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FC162E7B6717FDE2" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[920,1144,547,573]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="larutensis">
|
||
<emphasis id="72E38E6FC452FD68FC162E7B6717FDE2" box="[920,1144,547,573]" italics="true" pageId="17" pageNumber="520">Philautus larutensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FB082E7B61BBFDBE" authority="Arifin, Smart, Hertwig, Smith, Iskandar, and Hass" authorityName="Arifin, Smart, Hertwig, Smith, Iskandar, and Hass" class="Amphibia" family="Ranidae" genus="Sumaterana" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="dabulescens">
|
||
<emphasis id="72E38E6FC452FD68FB082E7B66F3FDE2" box="[1158,1436,547,573]" italics="true" pageId="17" pageNumber="520">Sumaterana dabulescens</emphasis>
|
||
Arifin, Smart, Hertwig, Smith, Iskandar, and Hass
|
||
</taxonomicName>
|
||
; at the
|
||
<typeStatus id="9F2CECDFC452FD68FCA92E1F60E8FDBE" box="[807,903,583,609]" pageId="17" pageNumber="520" type="paratype">paratype</typeStatus>
|
||
locality:
|
||
<taxonomicName id="879729FEC452FD68FC7A2E1F6791FDBE" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1012,1278,583,609]" class="Amphibia" family="Ranidae" genus="Chalcorana" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="chalconota">
|
||
<emphasis id="72E38E6FC452FD68FC7A2E1F6791FDBE" box="[1012,1278,583,609]" italics="true" pageId="17" pageNumber="520">Chalcorana chalconota</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FA812E1F6265FD5A" baseAuthorityName="Tschudi" baseAuthorityYear="1838" class="Amphibia" family="Bufonidae" genus="Leptophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="borbonica">
|
||
<emphasis id="72E38E6FC452FD68FA812E1F6265FD5A" italics="true" pageId="17" pageNumber="520">Leptophryne borbonica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FE932E33629AFD5A" baseAuthorityName="Tschudi" baseAuthorityYear="1838" box="[285,501,619,645]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="72E38E6FC452FD68FE932E33629AFD5A" box="[285,501,619,645]" italics="true" pageId="17" pageNumber="520">Limnonectes kuhlii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FD862E3361F5FD5A" box="[520,666,619,645]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="macrodon">
|
||
<emphasis id="72E38E6FC452FD68FD862E3361F5FD5A" box="[520,666,619,645]" italics="true" pageId="17" pageNumber="520">L. macrodon</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FD232E336008FD5A" box="[685,871,619,645]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="72E38E6FC452FD68FD232E336054FD5B" box="[685,827,619,644]" italics="true" pageId="17" pageNumber="520">Limnonectes</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FCF62E3367EAFD5B" box="[888,1157,619,645]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="17" pageNumber="520" phylum="Arthropoda" rank="species" species="catamitus">
|
||
<emphasis id="72E38E6FC452FD68FCF62E3367EAFD5B" box="[888,1157,619,645]" italics="true" pageId="17" pageNumber="520">Rhacophorus catamitus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="879729FEC452FD68FB162E336641FD5A" box="[1176,1326,619,645]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="72E38E6FC452FD68FB162E33666CFD5A" box="[1176,1283,619,645]" italics="true" pageId="17" pageNumber="520">Philautus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="879729FEC452FD68FAE22E33600AFD76" authority="Arifin, Cahyadi, Smart, Jankowski, and Haas." authorityName="Arifin, Cahyadi, Smart, Jankowski, and Haas." class="Amphibia" family="Ranidae" genus="Pulchrana" kingdom="Animalia" order="Anura" pageId="17" pageNumber="520" phylum="Chordata" rank="species" species="fantastica">
|
||
<emphasis id="72E38E6FC452FD68FAE22E336233FD76" italics="true" pageId="17" pageNumber="520">Pulchrana fantastica</emphasis>
|
||
Arifin, Cahyadi, Smart, Jankowski, and Haas.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |