201 lines
33 KiB
XML
201 lines
33 KiB
XML
<document id="DA3CD87F173459B824F90D73313A263E" ID-CLB-Dataset="86221" ID-DOI="10.11646/zootaxa.5159.3.1" ID-GBIF-Dataset="e19dda1c-86af-404d-9595-c08d8c848abd" ID-ISSN="1175-5326" ID-Zenodo-Dep="6781669" ID-ZooBank="13308111-EF49-4710-9C45-CF69DABE2C5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656566366326" checkinUser="plazi" docAuthor="Kataev, Boris M., Hongbin, Liang & Wrase, David W." docDate="2022" docId="D364737A24291733FF60B6F211FE1DF6" docLanguage="en" docName="zootaxa.5159.3.1.pdf" docOrigin="Zootaxa 5159 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus hamulipenis Kataev & Hongbin & Wrase 2022, sp. n." docType="treatment" docVersion="7" lastPageNumber="340" masterDocId="2F5D0B02240C1714FFF7B578133B1A36" masterDocTitle="New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies" masterLastPageNumber="353" masterPageNumber="301" pageNumber="338" updateTime="1699353733348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="926FED90CAF00DA67D3F9C7AF97C582E">
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<mods:title id="582D3C3805124BE46422B84860DE14B8">New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies</mods:title>
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<mods:name id="5ABFB98F8196C5DD962225B4C97BE401" type="personal">
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<mods:namePart id="2B983E6438E5F194E38EB708ED0A49C5">Kataev, Boris M.</mods:namePart>
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<mods:affiliation id="F54FBE4373B67685C5F7EA08C93311E3">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:name id="1C9902221D0949208B1B2BAC1AFA8B5D" type="personal">
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<mods:roleTerm id="AFE1EE509809221EFC5E63FA4F6A9F7E">Author</mods:roleTerm>
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<mods:namePart id="4CB506E9B2146A681FEE12783BB698F5">Hongbin, Liang</mods:namePart>
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<mods:nameIdentifier id="6B4A14BB041FEBA85256A1EB8F04076B" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
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<mods:affiliation id="8DB328C8DC18BF99EF18B81E90B07685">Key laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. lianghb @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 9668 - 1167</mods:affiliation>
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<mods:nameIdentifier id="45F40919650BEAE77D26CCD28710819F" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:namePart id="DC6DBD0568EDDDF371D43169CCDBECE2">Wrase, David W.</mods:namePart>
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<mods:affiliation id="BA28D072B2E1D7785218456DDF560A4C">Oderstr. 2, D- 15306 Gusow-Platkow, Germany.</mods:affiliation>
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<mods:title id="CE51B92BB1A23C8E811EC4972F5F05B0">Zootaxa</mods:title>
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<mods:date id="4A551A7EECCD4923ED6EBFEB5E4BB102">2022</mods:date>
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<mods:number id="0D8CD93960EC6983962FBC19B1EDA295">2022-06-30</mods:number>
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<treatment id="D364737A24291733FF60B6F211FE1DF6" ID-DOI="http://doi.org/10.5281/zenodo.6781719" ID-GBIF-Taxon="196373184" ID-Zenodo-Dep="6781719" LSID="urn:lsid:plazi:treatment:D364737A24291733FF60B6F211FE1DF6" httpUri="http://treatment.plazi.org/id/D364737A24291733FF60B6F211FE1DF6" lastPageId="39" lastPageNumber="340" pageId="37" pageNumber="338">
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<subSubSection id="13D791E724291731FF60B6F211BF1993" box="[151,644,906,933]" pageId="37" pageNumber="338" type="nomenclature">
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<paragraph id="5B72C26C24291731FF60B6F211BF1993" blockId="37.[151,644,906,969]" box="[151,644,906,933]" pageId="37" pageNumber="338">
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<heading id="003A750024291731FF60B6F211BF1993" bold="true" box="[151,644,906,933]" fontSize="11" level="1" pageId="37" pageNumber="338" reason="1">
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<emphasis id="69B91E7E24291731FF60B6F211BF1993" bold="true" box="[151,644,906,933]" italics="true" pageId="37" pageNumber="338">
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<taxonomicName id="9CCDB9EF24291731FF60B6F211011993" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[151,570,906,933]" class="Insecta" family="Carabidae" genus="Trichotichnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="hamulipenis" status="sp. nov.">Trichotichnus (s. str.) hamulipenis</taxonomicName>
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<taxonomicNameLabel id="728AA30524291731FDB5B6F311BF1993" box="[578,644,907,933]" pageId="37" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="13D791E724291733FF60B6D711641C26" lastPageId="39" lastPageNumber="340" pageId="37" pageNumber="338" type="description">
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<paragraph id="5B72C26C24291731FF60B6D7125D19FF" blockId="37.[151,644,906,969]" box="[151,358,942,969]" pageId="37" pageNumber="338">
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(
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<figureCitation id="C3F6DEE924291731FF68B6D713C519FF" box="[159,254,943,969]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="37" pageNumber="338">Figs. 20</figureCitation>
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,
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<figureCitation id="C3F6DEE924291731FEFEB6D6126519FF" box="[265,350,942,969]" captionStart-0="FIGURES 94–95" captionStart-1="FIGURES 96–97" captionStart-2="FIGURES 98–105" captionStartId-0="38.[151,264,1747,1771]" captionStartId-1="39.[151,264,1168,1192]" captionStartId-2="40.[151,264,1637,1661]" captionTargetBox-0="[174,1411,469,1715]" captionTargetBox-1="[153,1425,777,1137]" captionTargetBox-2="[158,1430,964,1605]" captionTargetId-0="figure-151@38.[158,1429,453,1724]" captionTargetId-1="figure-266@39.[153,1434,777,1145]" captionTargetId-2="figure-323@40.[151,1436,957,1614]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionTargetPageId-2="40" captionText-0="FIGURES 94–95. Trichotichnus hamulipenis sp. n., habitus. 94, nominotypical subspecies (above Bapo); 95, T. hamulipenis nepalensis ssp. n. (holotype). Scale bar = 1.0 mm." captionText-1="FIGURES 96–97. Trichotichnus hamulipenis sp. n., elytral apex. 96, nominotypical subspecies; 97, T. hamulipenis nepalensis ssp. n. Scale bar = 0.5 mm." captionText-2="FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105)." figureDoi-0="http://doi.org/10.5281/zenodo.6781755" figureDoi-1="http://doi.org/10.5281/zenodo.6781757" figureDoi-2="http://doi.org/10.5281/zenodo.6781759" httpUri-0="https://zenodo.org/record/6781755/files/figure.png" httpUri-1="https://zenodo.org/record/6781757/files/figure.png" httpUri-2="https://zenodo.org/record/6781759/files/figure.png" pageId="37" pageNumber="338">94–105</figureCitation>
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)
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</paragraph>
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<paragraph id="5B72C26C24291731FF60B68E160B1E27" blockId="37.[151,1437,1014,2013]" box="[151,1328,1014,1041]" pageId="37" pageNumber="338">
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<emphasis id="69B91E7E24291731FF60B68E121A1E26" bold="true" box="[151,289,1014,1040]" pageId="37" pageNumber="338">Description</emphasis>
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(
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<specimenCount id="4DCB09E524291731FEC6B68F12AD1E27" box="[305,406,1015,1041]" pageId="37" pageNumber="338" type="male">10 males</specimenCount>
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and
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<specimenCount id="4DCB09E524291731FE3AB68E11021E27" box="[461,569,1014,1041]" pageId="37" pageNumber="338" type="female">5 females</specimenCount>
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measured). Body length
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<quantity id="9C356F8924291731FCA4B68F10D01E27" box="[851,1003,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.2" metricValueMin="8.8" pageId="37" pageNumber="338" unit="mm" value="9.5" valueMax="10.2" valueMin="8.8">8.8–10.2 mm</quantity>
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. Habitus as in
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<figureCitation id="C3F6DEE924291731FB78B68F16171E27" box="[1167,1324,1014,1041]" captionStart="FIGURES 94–95" captionStartId="38.[151,264,1747,1771]" captionTargetBox="[174,1411,469,1715]" captionTargetId="figure-151@38.[158,1429,453,1724]" captionTargetPageId="38" captionText="FIGURES 94–95. Trichotichnus hamulipenis sp. n., habitus. 94, nominotypical subspecies (above Bapo); 95, T. hamulipenis nepalensis ssp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781755" httpUri="https://zenodo.org/record/6781755/files/figure.png" pageId="37" pageNumber="338">Figs. 94 & 95</figureCitation>
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.
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</paragraph>
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<paragraph id="5B72C26C24291731FF30B163174C1E4B" blockId="37.[151,1437,1014,2013]" pageId="37" pageNumber="338">Body dark brown to black, shiny on dorsum, very slightly iridescent on elytra; labrum and mandibles along lower outer margin, occasionally also base of mandibles and narrow lateral margins of pronotum reddish brown. Palpi, antennae, tibiae and tarsi brownish yellow or light brown; femora blackish brown.</paragraph>
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<paragraph id="5B72C26C24291731FF30B1FF127F1CD7" blockId="37.[151,1437,1014,2013]" pageId="37" pageNumber="338">
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Head moderately sized (HWmax/PWmax 0.70–0.76, HWmin/PWmax 0.55– 0.58). Eyes large and convex, almost hemispherical (HWmax/HWmin 1.25–1.33), in lateral view almost round. Genae narrow, about as wide as antennomere 2 apically. Tempora short, glabrous. Fronto-clypeal suture superficial or slightly impressed, very widely rounded. Frontal foveae wide and shallow. Fronto-ocular furrows reaching supraorbital furrows, distinct along entire length, deepened at clypeus. Supraorbital furrows narrow, touching upper margin of eye. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eye and removed from supraorbital furrows by distance approximately equal to width of antennomere 2 apically; in many specimens one or two very small unsetigerous punctures present medially of each supraorbital seta. Labrum very shallowly concave anteriorly. Clypeus slightly convex, very shallowly emarginate along anterior margin, with a pair of setigerous pores each located at anterior angle.
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<collectingRegion id="99090C8E24291731FE44B0B311291FD3" box="[435,530,1483,1509]" country="Myanmar" name="Chin" pageId="37" pageNumber="338">Mentum</collectingRegion>
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separated from submentum, with moderately wide median tooth; epilobes markedly widened apically; submentum with one pair of long setae. Ligular sclerite widened apically, truncate at apex, with one pair of ventroapical setae. Paraglossae very narrow, shorter than ligular sclerite and separated from it by wide notches. Labial penultimate palpomere about as long as ultimate palpomere. Left mandible sharply bent in the apical quarter, blunted at tip. Dorsal microsculpture very fine, consisting of obliterated, more or less isodiametric meshes. Antennae slender and long, surpassing pronotal basal edge in female by three, in male almost by four apical antennomeres, with antennomeres 4–8 about 3.5 times as long as wide. Basal antennomere very slightly longer than antennomere 3.
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</paragraph>
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<paragraph id="5B72C26C24291732FF30B39312C41B9B" blockId="37.[151,1437,1014,2013]" lastBlockId="38.[151,1436,151,429]" lastPageId="38" lastPageNumber="339" pageId="37" pageNumber="338">Pronotum transverse (PWmax/PL 1.34–1.48), widest slightly before the middle (PWmax/PWmin 1.24–1.35), more strongly narrowed apically than basally, with one lateral seta somewhat removed from lateral edge to internal margin of lateral depression and inserted slightly before widest point of pronotum. Sides rounded along entire length or with very short sinuation just before basal angles; lateral bead narrow, slightly widened basally. Apical margin very shallowly emarginate, very narrowly bordered along entire length, but occasionally border obliterated at the middle. Apical angles widely rounded, almost not protruding ahead. Basal margin more or less straight, bordered along entire length, slightly longer than apical margin and markedly shorter than base of elytra between humeral angles. Basal angles very obtuse, with small obtuse denticle at tip protruding laterally. Pronotal disc convex medially. Lateral depression beginning from apical angles, moderately narrow in apical third, rather strongly widened in the basal two third and fused with wide and deep laterobasal depressions; basal foveae located within laterobasal depressions, shallow, occasionally indistinct; basal area convex medially. Median line distinct, superficial or slightly impressed. Anterior transverse depression shallow or deepened. Surface either densely punctate almost throughout, with coarsest punctures in laterobasal depressions and with finest punctures in central part of disc, or punctation restricted only to rather wide area along margins. Microsculpture highly obliterated, indistinct transverse meshes recognizable mostly along base.</paragraph>
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<caption id="0FB292E4242A1732FF60B3AB11AE1D39" ID-DOI="http://doi.org/10.5281/zenodo.6781755" ID-Zenodo-Dep="6781755" httpUri="https://zenodo.org/record/6781755/files/figure.png" pageId="38" pageNumber="339" startId="38.[151,264,1747,1771]" targetBox="[174,1411,469,1715]" targetPageId="38">
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<paragraph id="5B72C26C242A1732FF60B3AB11AE1D39" blockId="38.[151,1436,1747,1808]" pageId="38" pageNumber="339">
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<emphasis id="69B91E7E242A1732FF60B3AB126E1CDD" bold="true" box="[151,341,1747,1771]" pageId="38" pageNumber="339">FIGURES 94–95.</emphasis>
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<taxonomicName id="9CCDB9EF242A1732FEABB3AB11501CDD" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[348,619,1747,1771]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="hamulipenis" status="sp. nov.">
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<emphasis id="69B91E7E242A1732FEABB3AB11501CDD" box="[348,619,1747,1771]" italics="true" pageId="38" pageNumber="339">Trichotichnus hamulipenis</emphasis>
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</taxonomicName>
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<emphasis id="69B91E7E242A1732FD84B3AC11961CDA" bold="true" box="[627,685,1748,1772]" pageId="38" pageNumber="339">
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<taxonomicNameLabel id="728AA305242A1732FD84B3AC11961CDA" box="[627,685,1748,1772]" pageId="38" pageNumber="339" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, habitus. 94, nominotypical subspecies (above Bapo); 95,
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<taxonomicName id="9CCDB9EF242A1732FAF1B3AC12391D39" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="subSpecies" species="hamulipenis" subSpecies="nepalensis">
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<emphasis id="69B91E7E242A1732FAF1B3AC12391D39" italics="true" pageId="38" pageNumber="339">T. hamulipenis nepalensis</emphasis>
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</taxonomicName>
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<emphasis id="69B91E7E242A1732FEFFB38012701D26" bold="true" box="[264,331,1784,1808]" pageId="38" pageNumber="339">ssp. n.</emphasis>
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(holotype). Scale bar = 1.0 mm.
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</paragraph>
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</caption>
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<paragraph id="5B72C26C242A1733FF30B24610031B2B" blockId="38.[151,1436,1854,2024]" lastBlockId="39.[151,1437,151,753]" lastPageId="39" lastPageNumber="340" pageId="38" pageNumber="339">Elytra convex, elongate oval (EL/EW 1.43–1.55, EL/PL 2.59–3.310, EW/PWmax 1.24–1.38), widest in the apical third. Humeri angularly rounded, at most with indistinct denticle poorly recognizable in caudal view. Subapical sinuation distinct but shallow. Sutural angle blunted or narrowly rounded at tip. Basal edge evenly arched or oblique; internal humeral angle very obtuse, almost indistinct. Striae impunctate, reaching anteriorly basal edge, impressed shallowly on disc and slightly more deeply apically. Intervals faintly convex or somewhat flat on disc, markedly narrowed and moderately convex apically. Parascutellar (abbreviate) striole long, with a small setigerous pore basally. Interval 3 with a very small discal setigerous pore adjacent to stria 2 behind the middle. Marginal umbilicate series without distinct gap at middle. Microsculpture strongly obliterated, consisting of indistinct transverse lines (recognizable at least apically and on lateral intervals).</paragraph>
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<paragraph id="5B72C26C242B1733FF30B45F11091B77" blockId="39.[151,1437,151,753]" box="[199,562,295,321]" pageId="39" pageNumber="340">Wings wholly or partly reduced.</paragraph>
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<paragraph id="5B72C26C242B1733FF30B43312581BBF" blockId="39.[151,1437,151,753]" pageId="39" pageNumber="340">Prosternum with very fine short setae. Metepisternum short, 1.18–1.25 times as long as wide, markedly narrowed posteriorly.</paragraph>
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<paragraph id="5B72C26C242B1733FF30B4EB116218C7" blockId="39.[151,1437,151,753]" pageId="39" pageNumber="340">
|
||
Legs relatively long. Metacoxae without additional setae. Metafemur ventrally with two setae at posterior margin and without setae at anterior margin. Protibia on dorsal side without longitudinal sulcus, outer margin preapically with one stouter and two very thin spines in male (
|
||
<figureCitation id="C3F6DEE9242B1733FCA0B4A310921BC3" box="[855,937,475,501]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="39" pageNumber="340">Fig. 20</figureCitation>
|
||
) and with three comparatively stout, similar sized spines in female. Tarsi dorsally glabrous; pro- and mesotarsomeres 1–4, particularly in male, with dense long ventro-lateral setae (as in
|
||
<figureCitation id="C3F6DEE9242B1733FE4BB75B112B180B" box="[444,528,547,573]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="39" pageNumber="340">Fig. 20</figureCitation>
|
||
); tarsomere 5 with three or four pairs of ventro-lateral setae. Metatarsus slender, about as long as (occasionally slightly shorter or longer than) HWmax, with tarsomeres somewhat weakly widened distally; metatarsomere 1 slightly shorter than metatarsomeres 2 and 3 combined; metatarsomeres 1–4 moderately densely setose ventrally. In male, pro- and mesotarsomeres 1–4 moderately widened and with adhesive scales ventrally; pro- and mesotarsomeres 2 and 3 longer than wide. In both sexes, mesotarsomere 1 elongate, slightly shorter than mesotarsomeres 2 and 3 combined.
|
||
</paragraph>
|
||
<caption id="0FB292E4242B1733FF60B1E812951EFA" ID-DOI="http://doi.org/10.5281/zenodo.6781757" ID-Zenodo-Dep="6781757" httpUri="https://zenodo.org/record/6781757/files/figure.png" pageId="39" pageNumber="340" startId="39.[151,264,1168,1192]" targetBox="[153,1425,777,1137]" targetPageId="39">
|
||
<paragraph id="5B72C26C242B1733FF60B1E812951EFA" blockId="39.[151,1436,1168,1228]" pageId="39" pageNumber="340">
|
||
<emphasis id="69B91E7E242B1733FF60B1E8126F1E9E" bold="true" box="[151,340,1168,1192]" pageId="39" pageNumber="340">FIGURES 96–97.</emphasis>
|
||
<taxonomicName id="9CCDB9EF242B1733FEADB1E811531E9E" authority="Kataev & Hongbin & Wrase, 2022" authorityName="Kataev & Hongbin & Wrase" authorityYear="2022" box="[346,616,1168,1192]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="hamulipenis" status="sp. nov.">
|
||
<emphasis id="69B91E7E242B1733FEADB1E811531E9E" box="[346,616,1168,1192]" italics="true" pageId="39" pageNumber="340">Trichotichnus hamulipenis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E242B1733FD99B1E8119C1E9E" bold="true" box="[622,679,1168,1192]" pageId="39" pageNumber="340">
|
||
<taxonomicNameLabel id="728AA305242B1733FD99B1E8119C1E9E" box="[622,679,1168,1192]" pageId="39" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, elytral apex. 96, nominotypical subspecies; 97,
|
||
<taxonomicName id="9CCDB9EF242B1733FB61B1E916A71E9E" box="[1174,1436,1168,1192]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="subSpecies" species="hamulipenis" subSpecies="nepalensis">
|
||
<emphasis id="69B91E7E242B1733FB61B1E916A71E9E" box="[1174,1436,1168,1192]" italics="true" pageId="39" pageNumber="340">T. hamulipenis nepalensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="69B91E7E242B1733FF60B1CC13E11EFA" bold="true" box="[151,218,1204,1228]" pageId="39" pageNumber="340">ssp. n.</emphasis>
|
||
Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5B72C26C242B1733FF30B18217EE1F0F" blockId="39.[151,1436,1274,1985]" pageId="39" pageNumber="340">Abdominal sternites without additional long setae; apex of last visible abdominal sternite (VII) in male subtruncate and with one pair of marginal setae, in female rounded and with two pairs of marginal setae.</paragraph>
|
||
<paragraph id="5B72C26C242B1733FF30B03A174A1FFE" blockId="39.[151,1436,1274,1985]" pageId="39" pageNumber="340">
|
||
Median lobe of aedeagus (
|
||
<figureCitation id="C3F6DEE9242B1733FE19B03A11C81F6B" box="[494,755,1346,1373]" captionStart="FIGURES 98–105" captionStartId="40.[151,264,1637,1661]" captionTargetBox="[158,1430,964,1605]" captionTargetId="figure-323@40.[151,1436,957,1614]" captionTargetPageId="40" captionText="FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105)." figureDoi="http://doi.org/10.5281/zenodo.6781759" httpUri="https://zenodo.org/record/6781759/files/figure.png" pageId="39" pageNumber="340">Figs. 98–100, 103–105</figureCitation>
|
||
) stout, in lateral view arcuate; ventral side flattened in apical portion; terminal lamella in dorsal view wide, wider than long, not concave, strongly narrowed posteriorly, with very narrowly rounded apex; in lateral view sharply hooked ventrally. Apical orifice mostly in dorsal position, but slightly shifted to the left. Internal sac with large apical spine and twisted spiny patches.
|
||
</paragraph>
|
||
<paragraph id="5B72C26C242B1733FF30B0AA11641C26" blockId="39.[151,1436,1274,1985]" pageId="39" pageNumber="340">
|
||
Female genitalia (
|
||
<figureCitation id="C3F6DEE9242B1733FE66B0AA11761FDB" box="[401,589,1490,1517]" captionStart="FIGURES 98–105" captionStartId="40.[151,264,1637,1661]" captionTargetBox="[158,1430,964,1605]" captionTargetId="figure-323@40.[151,1436,957,1614]" captionTargetPageId="40" captionText="FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105)." figureDoi="http://doi.org/10.5281/zenodo.6781759" httpUri="https://zenodo.org/record/6781759/files/figure.png" pageId="39" pageNumber="340">Figs. 101 & 105</figureCitation>
|
||
): gonocoxite comparatively narrow in ventral view, about 0.6 times as long as gonosubcoxite, with rather wide base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13D791E7242B1733FF30B36311F31D4F" pageId="39" pageNumber="340" type="etymology">
|
||
<paragraph id="5B72C26C242B1733FF30B36312BF1C6E" blockId="39.[151,1436,1274,1985]" pageId="39" pageNumber="340">
|
||
<emphasis id="69B91E7E242B1733FF30B36312751C03" bold="true" box="[199,334,1563,1589]" pageId="39" pageNumber="340">Etymology.</emphasis>
|
||
The specific name is a Latin compound noun, referring to the sharp hook on the apex of the median lobe of the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="5B72C26C242B1733FF30B31A11F31D4F" blockId="39.[151,1436,1274,1985]" pageId="39" pageNumber="340">
|
||
<emphasis id="69B91E7E242B1733FF30B31A12591C4A" bold="true" box="[199,354,1634,1660]" pageId="39" pageNumber="340">Comparison.</emphasis>
|
||
This new species is similar to members of the
|
||
<emphasis id="69B91E7E242B1733FC8AB31A10CE1C4A" box="[893,1013,1634,1660]" italics="true" pageId="39" pageNumber="340">longitarsis</emphasis>
|
||
species group sensu
|
||
<bibRefCitation id="3F5CBF9D242B1733FB10B31A16501C4A" author="Habu, A." box="[1255,1387,1634,1660]" pageId="39" pageNumber="340" pagination="127 - 169" refId="ref37923" refString="Habu, A. (1961) Revisional study of the species of the Trichotichni, the subtribe of the tribe Harpalini from Japan. Bulletin of the National Institute of Agricultural Sciences (Series C), 13, 127 - 169." type="journal article" year="1961">Habu, 1961</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="3F5CBF9D242B1733FF60B3FF13F41C97" author="Habu, A." box="[151,207,1671,1697]" pageId="39" pageNumber="340" refId="ref37967" refString="Habu, A. (1973) Fauna Japonica. Carabidae: Harpalini (Insecta, Coleoptera). Keigaku Publishing Co., Tokyo, xiii + 430 pp." type="book" year="1973">1973</bibRefCitation>
|
||
(but excluding
|
||
<taxonomicName id="9CCDB9EF242B1733FE73B3FF11051C96" authorityName="Habu" authorityYear="1954" box="[388,574,1670,1696]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="noctuabundus">
|
||
<emphasis id="69B91E7E242B1733FE73B3FF11051C96" box="[388,574,1670,1696]" italics="true" pageId="39" pageNumber="340">T. noctuabundus</emphasis>
|
||
</taxonomicName>
|
||
) in many characters, including elongate body with relatively long legs, one pair of marginal setae on last abdominal sternite of male, densely setose tarsi ventrally (
|
||
<figureCitation id="C3F6DEE9242B1733FB98B3D217FA1CF3" box="[1135,1217,1706,1733]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 & 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 & 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="39" pageNumber="340">Fig. 20</figureCitation>
|
||
) and elongate male pro- and mesotarsomeres, but differing in having protibia not sulcate and wings reduced. In general habitus and male genitalia, it is most similar to
|
||
<taxonomicName id="9CCDB9EF242B1733FDC4B38B10641D3B" authority="Habu, 1954" authorityName="Habu" authorityYear="1954" box="[563,863,1778,1805]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="vespertinus">
|
||
<emphasis id="69B91E7E242B1733FDC4B38B11EA1D3A" box="[563,721,1779,1804]" italics="true" pageId="39" pageNumber="340">T. vespertinus</emphasis>
|
||
<bibRefCitation id="3F5CBF9D242B1733FD2DB38A10641D3B" author="Habu, A." box="[730,863,1778,1805]" pageId="39" pageNumber="340" pagination="53 - 59" refId="ref37858" refString="Habu, A. (1954) On four new species of Trichotichnus from Mt. Hiko, Kyushu (Coleoptera) (The Carabidae fauna of Mt. Hiko, II). Mushi, 26, 53 - 59." type="journal article" year="1954">Habu, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="23DA82FC242B1733FC5FB38B10D01D3B" box="[936,1003,1779,1805]" name="Japan" pageId="39" pageNumber="340">Japan</collectingCountry>
|
||
; the latter species, however, is distinguished by having femora brownish yellow, not infuscate, protibia with a longitudinal sulcus ventrally, and median lobe of aedeagus hooked at apex and with terminal lamella in dorsal view acute at tip; in addition, both species have a different pattern of spiny patches in internal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13D791E7242B1733FF30B2FA100A1DAA" box="[199,817,1922,1949]" pageId="39" pageNumber="340" type="distribution">
|
||
<paragraph id="5B72C26C242B1733FF30B2FA100A1DAA" blockId="39.[151,1436,1274,1985]" box="[199,817,1922,1949]" pageId="39" pageNumber="340">
|
||
<emphasis id="69B91E7E242B1733FF30B2FA125A1DAA" bold="true" box="[199,353,1922,1948]" pageId="39" pageNumber="340">Distribution.</emphasis>
|
||
Known from
|
||
<collectingCountry id="23DA82FC242B1733FE09B2FA117A1DAA" box="[510,577,1922,1948]" name="China" pageId="39" pageNumber="340">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="99090C8E242B1733FDA7B2FB11921DAB" box="[592,681,1923,1949]" country="China" name="Yunnan" pageId="39" pageNumber="340">Yunnan</collectingRegion>
|
||
) and
|
||
<collectingCountry id="23DA82FC242B1733FD10B2FA10151DAA" box="[743,814,1922,1948]" name="Nepal" pageId="39" pageNumber="340">Nepal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="13D791E7242B1733FF30B2DF11FE1DF6" box="[199,709,1958,1985]" pageId="39" pageNumber="340" type="discussion">
|
||
<paragraph id="5B72C26C242B1733FF30B2DF11FE1DF6" blockId="39.[151,1436,1274,1985]" box="[199,709,1958,1985]" pageId="39" pageNumber="340">
|
||
<emphasis id="69B91E7E242B1733FF30B2DF12001DF7" bold="true" box="[199,315,1959,1985]" pageId="39" pageNumber="340">Remarks.</emphasis>
|
||
The species forms two subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |