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<document id="D1F17211E2997CDF710B872828B0E095" ID-CLB-Dataset="86221" ID-DOI="10.11646/zootaxa.5159.3.1" ID-GBIF-Dataset="e19dda1c-86af-404d-9595-c08d8c848abd" ID-ISSN="1175-5326" ID-Zenodo-Dep="6781669" ID-ZooBank="13308111-EF49-4710-9C45-CF69DABE2C5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656566366326" checkinUser="plazi" docAuthor="Kataev, Boris M., Hongbin, Liang &amp; Wrase, David W." docDate="2022" docId="D364737A2407171AFF60B00512301D22" docLanguage="en" docName="zootaxa.5159.3.1.pdf" docOrigin="Zootaxa 5159 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus angustitarsis Kataev &amp; Hongbin &amp; Wrase 2022, sp. n." docType="treatment" docVersion="6" lastPageNumber="315" masterDocId="2F5D0B02240C1714FFF7B578133B1A36" masterDocTitle="New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies" masterLastPageNumber="353" masterPageNumber="301" pageNumber="312" updateTime="1699353733348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="47F9291087139A146E3B9976FA6C9F8B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="43AF477AE73CAA5B197DFC2DC425E155">
<mods:title id="9F6F49D24B09C69DF7C2A96A9CC2AA8A">New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="056778D30E8012A04D733FA138DC57C5" type="personal">
<mods:role id="AACF451FE61C12B8C3BE57E189CB1319">
<mods:roleTerm id="98EFCD0816552DA3AF47839ECFBCC430">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B03FCF97E108AA8923268916368F3F6">Kataev, Boris M.</mods:namePart>
<mods:affiliation id="9E50F67BB34BA6DE45EC2EE8A4B252E3">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
</mods:name>
<mods:name id="72F9E9CE51C1A51C9A2EA29D4F3ABAD1" type="personal">
<mods:role id="3391EEEFE3C645324BD856472F17A6EC">
<mods:roleTerm id="493FB527CD48663A78DD390FDCFFD517">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="769CAD8A53CCD6349AEAACEA482EA8FA">Hongbin, Liang</mods:namePart>
<mods:nameIdentifier id="682EF0DB201833C856A1BB48DC3A2E7F" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
<mods:affiliation id="1D5A10E11A05A1DBBBC2754C25A2FC1B">Key laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. lianghb @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 9668 - 1167</mods:affiliation>
<mods:nameIdentifier id="DB548858DA17932AB50667271048A351" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="5E2434A757286D5A7E14390F4FFC8F48" type="personal">
<mods:role id="11529DAE03FD891A94877F713885C59E">
<mods:roleTerm id="CB860468F6B88F3B97CD470B5666C523">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE55BDDBD694B4ACF427A58E454E57A8">Wrase, David W.</mods:namePart>
<mods:affiliation id="280DD6EF88465346293B4AEF33C2A4E5">Oderstr. 2, D- 15306 Gusow-Platkow, Germany.</mods:affiliation>
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<mods:typeOfResource id="D20632520E8B538D21E90B5F740C49EB">text</mods:typeOfResource>
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<mods:title id="30964A776884F1155E04D8F55FCE33DC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C31F52F5393773FC075564A3E24BC7A1">
<mods:date id="30F8A6B91C87DA670686CD0C06BDD671">2022</mods:date>
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<mods:number id="2843987333BF3CD223FD308AA2C8ACD3">2022-06-30</mods:number>
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<mods:classification id="A1DB587BC823B77213873A95B673D8C6">journal article</mods:classification>
<mods:identifier id="26FD914691979434A7644D5A00747412" type="CLB-Dataset">86221</mods:identifier>
<mods:identifier id="11AFA46E96590B0E4C895F6095B25675" type="DOI">10.11646/zootaxa.5159.3.1</mods:identifier>
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<treatment id="D364737A2407171AFF60B00512301D22" ID-DOI="http://doi.org/10.5281/zenodo.6781689" ID-GBIF-Taxon="196373196" ID-Zenodo-Dep="6781689" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22" httpUri="http://treatment.plazi.org/id/D364737A2407171AFF60B00512301D22" lastPageId="14" lastPageNumber="315" pageId="11" pageNumber="312">
<subSubSection id="13D791E72407171FFF60B00511B21FA1" box="[151,649,1405,1432]" pageId="11" pageNumber="312" type="nomenclature">
<paragraph id="5B72C26C2407171FFF60B00511B21FA1" blockId="11.[151,649,1405,1467]" box="[151,649,1405,1432]" pageId="11" pageNumber="312">
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<emphasis id="69B91E7E2407171FFF60B00511B21FA1" bold="true" box="[151,649,1405,1432]" pageId="11" pageNumber="312">
<taxonomicName id="9CCDB9EF2407171FFF60B00511041FA1" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[151,575,1405,1432]" class="Insecta" family="Carabidae" genus="Trichotichnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
<emphasis id="69B91E7E2407171FFF60B005127A1FAE" bold="true" box="[151,321,1405,1432]" italics="true" pageId="11" pageNumber="312">Trichotichnus</emphasis>
(s. str.)
<emphasis id="69B91E7E2407171FFE53B00511041FA1" bold="true" box="[420,575,1405,1431]" italics="true" pageId="11" pageNumber="312">angustitarsis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="728AA3052407171FFDB0B00511B21FA1" box="[583,649,1405,1431]" pageId="11" pageNumber="312" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13D791E72407171FFF60B0D912A51F8D" box="[151,414,1441,1467]" pageId="11" pageNumber="312" type="description">
<paragraph id="5B72C26C2407171FFF60B0D912A51F8D" blockId="11.[151,649,1405,1467]" box="[151,414,1441,1467]" pageId="11" pageNumber="312">
(
<figureCitation id="C3F6DEE92407171FFF68B0D913D41F8D" box="[159,239,1441,1467]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="11" pageNumber="312">Figs. 9</figureCitation>
,
<figureCitation id="C3F6DEE92407171FFF0CB0D912781F8D" box="[251,323,1441,1467]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 &amp; 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="11" pageNumber="312">19, 21</figureCitation>
,
<figureCitation id="C3F6DEE92407171FFEB8B0D912AD1F8D" box="[335,406,1441,1467]" captionStart-0="FIGURES 23–27" captionStart-1="FIGURE 28" captionStartId-0="14.[151,264,922,946]" captionStartId-1="15.[151,250,1700,1724]" captionTargetBox-0="[160,1428,186,890]" captionTargetBox-1="[151,1435,181,1677]" captionTargetId-0="figure-32@14.[151,1436,181,899]" captionTargetId-1="figure-19@15.[151,1435,181,1677]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 &amp; 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 &amp; 25), lateral (24 &amp; 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 &amp; 27), B = 1.0 mm (24 &amp; 25)." captionText-1="FIGURE 28. Trichotichnus angustitarsis sp. n., distribution." figureDoi-0="http://doi.org/10.5281/zenodo.6781705" figureDoi-1="http://doi.org/10.5281/zenodo.6781707" httpUri-0="https://zenodo.org/record/6781705/files/figure.png" httpUri-1="https://zenodo.org/record/6781707/files/figure.png" pageId="11" pageNumber="312">23–28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13D791E72407171FFF60B09C11701DC0" pageId="11" pageNumber="312" type="materials_examined">
<paragraph id="5B72C26C2407171FFF60B09C129E1C70" blockId="11.[151,1437,1508,2039]" pageId="11" pageNumber="312">
<emphasis id="69B91E7E2407171FFF60B09C12751FC8" bold="true" box="[151,334,1508,1534]" pageId="11" pageNumber="312">Type material.</emphasis>
<materialsCitation id="EBA5C8312407171FFEA9B09C129E1C70" ID-GBIF-Occurrence="3821894405" collectingDate="2004-11-03" collectionCode="IOZ" collectorName="Liang Hongbin &amp; California Academy" country="China" elevation="1412" latitude="27.73902" location="Male" longLatPrecision="1" longitude="98.34975" pageId="11" pageNumber="312" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="84767CCE2407171FFEA9B09C12F11FC8" box="[350,458,1508,1534]" pageId="11" pageNumber="312" type="holotype">Holotype</typeStatus>
.
<location id="5E1294B72407171FFE29B09C11211FC8" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFE29B09C11211FC8" box="[478,538,1508,1534]" country="China" latitude="27.73902" longLatPrecision="1" longitude="98.34975" name="Male" pageId="11" pageNumber="312" stateProvince="Yunnan">Male</location>
, “
<collectingCountry id="23DA82FC2407171FFDCFB09C11AD1FC8" box="[568,662,1508,1534]" name="China" pageId="11" pageNumber="312">CHINA</collectingCountry>
,
<collectingRegion id="99090C8E2407171FFD5EB09D11C41FC9" box="[681,767,1509,1535]" country="China" name="Yunnan" pageId="11" pageNumber="312">Yunnan</collectingRegion>
Provin., Gongshan, Dulongjiang Bapo, along roadside
<geoCoordinate id="3EF9A4AB2407171FFF60B37012181C14" box="[151,291,1544,1570]" degrees="27.73902" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="27.73902">27.73902°N</geoCoordinate>
,
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”, “
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,
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<collectingDate id="3F371D442407171FFDB5B37011891C14" box="[578,690,1544,1570]" pageId="11" pageNumber="312" value="2004-11-03">2004.11.3</collectingDate>
</date>
night
<collectorName id="F638A7BA2407171FFD0BB37010931C14" box="[764,936,1544,1570]" pageId="11" pageNumber="312">Liang Hongbin</collectorName>
collector
<collectorName id="F638A7BA2407171FFBECB37017C51C14" box="[1051,1278,1544,1570]" pageId="11" pageNumber="312">California Academy</collectorName>
&amp; IOZ., Chinese. Acad. Sci.” (
<collectionCode id="3DDC5AA92407171FFE92B35412A21C70" box="[357,409,1580,1606]" pageId="11" pageNumber="312">IOZ</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5B72C26C2407171FFF30B32916011D98" blockId="11.[151,1437,1508,2039]" pageId="11" pageNumber="312">
<materialsCitation id="EBA5C8312407171FFF30B32910171CB8" ID-GBIF-Occurrence="3821894469" collectingDate="2004-11-03" collectionCode="CAS" collectorName="D. Kavanaugh" country="China" county="Gongshan" elevation="1640" latitude="28.00287" location="Elidang village" longLatPrecision="1" longitude="98.32145" municipality="Dulongjiang" pageId="11" pageNumber="312" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="84767CCE2407171FFF30B32912031C5D" box="[199,312,1617,1643]" pageId="11" pageNumber="312" type="paratype">Paratypes</typeStatus>
.
<emphasis id="69B91E7E2407171FFEB4B32812A81C5C" bold="true" box="[323,403,1616,1642]" pageId="11" pageNumber="312">
<collectingCountry id="23DA82FC2407171FFEB4B32812B51C5C" box="[323,398,1616,1642]" name="China" pageId="11" pageNumber="312">China</collectingCountry>
.
</emphasis>
<collectingRegion id="99090C8E2407171FFE6DB32811281C5C" box="[410,531,1616,1642]" country="China" name="Yunnan" pageId="11" pageNumber="312">YUNNAN</collectingRegion>
:
<specimenCount id="4DCB09E52407171FFDD4B32811491C5C" box="[547,626,1616,1642]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
,
<specimenCount id="4DCB09E52407171FFD89B32811D81C5C" box="[638,739,1616,1642]" pageId="11" pageNumber="312" type="female">1 female</specimenCount>
,
<collectingCounty id="B213BAE02407171FFD18B32810581C5C" box="[751,867,1616,1642]" pageId="11" pageNumber="312">Gongshan</collectingCounty>
,
<collectingMunicipality id="BB1658162407171FFC98B32810C51C5C" box="[879,1022,1616,1642]" pageId="11" pageNumber="312">Dulongjiang</collectingMunicipality>
,
<location id="5E1294B72407171FFBFDB32817831C5C" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFBFDB32817831C5C" box="[1034,1208,1616,1642]" country="China" county="Gongshan" latitude="28.00287" longLatPrecision="1" longitude="98.32145" municipality="Dulongjiang" name="Elidang village" pageId="11" pageNumber="312" stateProvince="Yunnan">Elidang village</location>
, beach,
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<geoCoordinate id="3EF9A4AB2407171FFF60B30C121B1CB9" box="[151,288,1652,1679]" degrees="98.32145" direction="east" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="98.32145">98.32145°E</geoCoordinate>
,
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</quantity>
,
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<collectingDate id="3F371D442407171FFE7CB30C12C41CB8" box="[395,511,1652,1678]" pageId="11" pageNumber="312" value="2004-11-03">3.XI.2004</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFDFDB30D11901CB8" box="[522,683,1652,1679]" pageId="11" pageNumber="312">D. Kavanaugh</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFD1EB30C101E1CB8" box="[745,805,1652,1678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="312" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFCC0B30C10D21C84" ID-GBIF-Occurrence="3821894439" collectingDate="2006-08-29" collectionCode="IOZ" collectorName="Y. Liu" country="China" county="Gongshan" elevation="1814" latitude="27.68545" location="Maku" longLatPrecision="1" longitude="98.30419" municipality="Dulongjiang" pageId="11" pageNumber="312" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFCC0B30C10BE1CB8" box="[823,901,1652,1678]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
,
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,
<collectingCounty id="B213BAE02407171FFC0AB30C174A1CB8" box="[1021,1137,1652,1678]" pageId="11" pageNumber="312">Gongshan</collectingCounty>
,
<collectingMunicipality id="BB1658162407171FFB8BB30C16301CB8" box="[1148,1291,1652,1678]" pageId="11" pageNumber="312">Dulongjiang</collectingMunicipality>
,
<location id="5E1294B72407171FFAE2B30C16621CB8" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFAE2B30C16621CB8" box="[1301,1369,1652,1678]" country="China" county="Gongshan" latitude="27.68545" longLatPrecision="1" longitude="98.30419" municipality="Dulongjiang" name="Maku" pageId="11" pageNumber="312" stateProvince="Yunnan">Maku</location>
, road,
<collectingMethod id="828CBA7B2407171FFF60B3E013C41C85" box="[151,255,1688,1715]" pageId="11" pageNumber="312">light trap</collectingMethod>
,
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,
<quantity id="9C356F892407171FFDDCB3E011BA1C85" box="[555,641,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.814" pageId="11" pageNumber="312" unit="m" value="1814.0">
<elevation id="D0E0255F2407171FFDDCB3E011BA1C85" box="[555,641,1688,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.814" pageId="11" pageNumber="312" unit="m" value="1814.0">1814 m</elevation>
</quantity>
,
<date id="2F73E4AC2407171FFD7BB3E0101A1C84" box="[652,801,1688,1714]" pageId="11" pageNumber="312" value="2006-08-29">
<collectingDate id="3F371D442407171FFD7BB3E0101A1C84" box="[652,801,1688,1714]" pageId="11" pageNumber="312" value="2006-08-29">29.VIII.2006</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFCDDB3E110541C84" box="[810,879,1688,1715]" pageId="11" pageNumber="312">Y. Liu</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFC5AB3E010DA1C84" box="[941,993,1688,1714]" pageId="11" pageNumber="312">IOZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFC03B3E016071CE0" ID-GBIF-Occurrence="3821894438" collectingDate="2006-08-29" collectionCode="IOZ" collectorName="D. Kavanaugh &amp; J. A. Miller" country="China" elevation="1970" latitude="27.97775" location="Dulongjiang" longLatPrecision="1" longitude="98.29771" municipality="Gongshan" pageId="11" pageNumber="312" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFC03B3E0176C1C84" box="[1012,1111,1688,1714]" pageId="11" pageNumber="312" type="female">1 female</specimenCount>
,
<collectingMunicipality id="BB1658162407171FFB96B3E017EE1C84" box="[1121,1237,1688,1714]" pageId="11" pageNumber="312">Gongshan</collectingMunicipality>
,
<location id="5E1294B72407171FFB17B3E016541C84" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFB17B3E016541C84" box="[1248,1391,1688,1714]" country="China" latitude="27.97775" longLatPrecision="1" longitude="98.29771" municipality="Gongshan" name="Dulongjiang" pageId="11" pageNumber="312" stateProvince="Yunnan">Dulongjiang</location>
, 1.0 km
<locationDeviation id="4AC0DB4E2407171FFF34B3C412631CE0" box="[195,344,1724,1750]" location="Maku" pageId="11" pageNumber="312">SW of Maku</locationDeviation>
,
<geoCoordinate id="3EF9A4AB2407171FFE92B3C412D61CE1" box="[357,493,1724,1751]" degrees="27.97775" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="27.97775">27.97775°N</geoCoordinate>
<geoCoordinate id="3EF9A4AB2407171FFE02B3C411451CE0" box="[501,638,1724,1750]" degrees="98.29771" direction="east" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="98.29771">98.29771°E</geoCoordinate>
,
<quantity id="9C356F892407171FFD7DB3C411D91CE1" box="[650,738,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="11" pageNumber="312" unit="m" value="1970.0">
<elevation id="D0E0255F2407171FFD7DB3C411D91CE1" box="[650,738,1724,1751]" metricMagnitude="3" metricUnit="m" metricValue="1.97" pageId="11" pageNumber="312" unit="m" value="1970.0">1970 m</elevation>
</quantity>
,
<date id="2F73E4AC2407171FFD18B3C410BF1CE0" box="[751,900,1724,1750]" pageId="11" pageNumber="312" value="2006-08-29">
<collectingDate id="3F371D442407171FFD18B3C410BF1CE0" box="[751,900,1724,1750]" pageId="11" pageNumber="312" value="2006-08-29">29.VIII.2006</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFC67B3C5170D1CE0" box="[912,1078,1724,1751]" pageId="11" pageNumber="312">D. Kavanaugh</collectorName>
,
<collectorName id="F638A7BA2407171FFBB5B3C417851CE0" box="[1090,1214,1724,1750]" pageId="11" pageNumber="312">J.A. Miller</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFAF7B3C4160F1CE0" box="[1280,1332,1724,1750]" pageId="11" pageNumber="312">IOZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFABEB3C412AF1D28" ID-GBIF-Occurrence="3821894526" collectingDate="2006-08-30" collectionCode="CAS" collectorName="D. Kavanaugh &amp; J. A. Miller" country="China" elevation="1845" latitude="27.6831" location="Dulongjiang" longLatPrecision="1" longitude="98.30038" municipality="Gongshan" pageId="11" pageNumber="312" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFABEB3C416A31CE0" box="[1353,1432,1724,1750]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
,
<collectingMunicipality id="BB1658162407171FFF60B39812301CCC" box="[151,267,1760,1786]" pageId="11" pageNumber="312">Gongshan</collectingMunicipality>
,
<location id="5E1294B72407171FFEE1B398129E1CCC" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFEE1B398129E1CCC" box="[278,421,1760,1786]" country="China" latitude="27.6831" longLatPrecision="1" longitude="98.30038" municipality="Gongshan" name="Dulongjiang" pageId="11" pageNumber="312" stateProvince="Yunnan">Dulongjiang</location>
,
<locationDeviation id="4AC0DB4E2407171FFE58B39811B41CCC" box="[431,655,1760,1787]" location="Maku" pageId="11" pageNumber="312">
<quantity id="9C356F892407171FFE58B39812C71CCC" box="[431,508,1760,1787]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="312" unit="km" value="0.5">0.5 km</quantity>
WSW Maku
</locationDeviation>
, trail,
<geoCoordinate id="3EF9A4AB2407171FFD23B39810671CCC" box="[724,860,1760,1786]" degrees="27.68310" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="27.6831">27.68310°N</geoCoordinate>
<geoCoordinate id="3EF9A4AB2407171FFC95B39810D01CCC" box="[866,1003,1760,1786]" degrees="98.30038" direction="east" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="98.30038">98.30038°E</geoCoordinate>
,
<quantity id="9C356F892407171FFC03B39817711CCD" box="[1012,1098,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.845" pageId="11" pageNumber="312" unit="m" value="1845.0">
<elevation id="D0E0255F2407171FFC03B39817711CCD" box="[1012,1098,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.845" pageId="11" pageNumber="312" unit="m" value="1845.0">1845 m</elevation>
</quantity>
,
<date id="2F73E4AC2407171FFBA2B39817D11CCC" box="[1109,1258,1760,1786]" pageId="11" pageNumber="312" value="2006-08-30">
<collectingDate id="3F371D442407171FFBA2B39817D11CCC" box="[1109,1258,1760,1786]" pageId="11" pageNumber="312" value="2006-08-30">30.VIII.2006</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFB03B39916A31CCC" box="[1268,1432,1760,1787]" pageId="11" pageNumber="312">D. Kavanaugh</collectorName>
,
<collectorName id="F638A7BA2407171FFF60B27C122A1D28" box="[151,273,1796,1822]" pageId="11" pageNumber="312">J.A. Miller</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFEA6B27C12B61D28" box="[337,397,1796,1822]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="312" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFE57B27C12C61D74" ID-GBIF-Occurrence="3821894339" collectingDate="2006-08-27" collectionCode="CAS" collectorName="Y. Liu" country="China" county="Gongshan" elevation="1511" latitude="27.87755" location="Kongdang" longLatPrecision="1" longitude="98.33612" municipality="Dulongjiang" pageId="11" pageNumber="312" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFE57B27C113F1D28" box="[416,516,1796,1822]" pageId="11" pageNumber="312" type="female">1 female</specimenCount>
,
<collectingCounty id="B213BAE02407171FFDF8B27C11B81D28" box="[527,643,1796,1822]" pageId="11" pageNumber="312">Gongshan</collectingCounty>
,
<collectingMunicipality id="BB1658162407171FFD78B27C10251D28" box="[655,798,1796,1822]" pageId="11" pageNumber="312">Dulongjiang</collectingMunicipality>
,
<location id="5E1294B72407171FFCDEB27C109A1D28" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFCDEB27C109A1D28" box="[809,929,1796,1822]" country="China" county="Gongshan" latitude="27.87755" longLatPrecision="1" longitude="98.33612" municipality="Dulongjiang" name="Kongdang" pageId="11" pageNumber="312" stateProvince="Yunnan">Kongdang</location>
,
<collectingMethod id="828CBA7B2407171FFC5CB27C17281D29" box="[939,1043,1796,1823]" pageId="11" pageNumber="312">light trap</collectingMethod>
,
<geoCoordinate id="3EF9A4AB2407171FFBE8B27C179C1D29" box="[1055,1191,1796,1823]" degrees="27.87755" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="27.87755">27.87755°N</geoCoordinate>
<geoCoordinate id="3EF9A4AB2407171FFB59B27C160D1D28" box="[1198,1334,1796,1822]" degrees="98.33612" direction="east" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="98.33612">98.33612°E</geoCoordinate>
,
<quantity id="9C356F892407171FFAB5B27C16AC1D29" box="[1346,1431,1796,1823]" metricMagnitude="3" metricUnit="m" metricValue="1.511" pageId="11" pageNumber="312" unit="m" value="1511.0">
<elevation id="D0E0255F2407171FFAB5B27C16AC1D29" box="[1346,1431,1796,1823]" metricMagnitude="3" metricUnit="m" metricValue="1.511" pageId="11" pageNumber="312" unit="m" value="1511.0">1511 m</elevation>
</quantity>
,
<date id="2F73E4AC2407171FFF60B25012171D74" box="[151,300,1832,1858]" pageId="11" pageNumber="312" value="2006-08-27">
<collectingDate id="3F371D442407171FFF60B25012171D74" box="[151,300,1832,1858]" pageId="11" pageNumber="312" value="2006-08-27">27.VIII.2006</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFEC1B25112401D74" box="[310,379,1832,1859]" pageId="11" pageNumber="312">Y. Liu</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFE4EB25012CE1D74" box="[441,501,1832,1858]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="11" pageNumber="312" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFDFFB25010021D50" ID-GBIF-Occurrence="3821894559" collectingDate="2005-05-28" collectionCode="IOZ, ZIN" collectorName="H. M. Yan" country="China" county="Longling Co." elevation="2067" latitude="24.83671" location="Xiaoheishan" longLatPrecision="1" longitude="98.76185" municipality="Longjiang" pageId="11" pageNumber="312" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFDFFB250115A1D74" box="[520,609,1832,1858]" pageId="11" pageNumber="312" type="male">2 males</specimenCount>
,
<specimenCount id="4DCB09E52407171FFD9BB25011F41D74" box="[620,719,1832,1858]" pageId="11" pageNumber="312" type="female">1 female</specimenCount>
,
<collectingCounty id="B213BAE02407171FFD2DB250104A1D74" box="[730,881,1832,1858]" pageId="11" pageNumber="312">Longling Co.</collectingCounty>
,
<collectingMunicipality id="BB1658162407171FFC8BB25010C91D74" box="[892,1010,1832,1858]" pageId="11" pageNumber="312">Longjiang</collectingMunicipality>
,
<location id="5E1294B72407171FFC0BB25017B01D74" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFC0BB25017B01D74" box="[1020,1163,1832,1858]" country="China" county="Longling Co." latitude="24.83671" longLatPrecision="1" longitude="98.76185" municipality="Longjiang" name="Xiaoheishan" pageId="11" pageNumber="312" stateProvince="Yunnan">Xiaoheishan</location>
, tree &amp; log,
<geoCoordinate id="3EF9A4AB2407171FFAE3B25016A61D74" box="[1300,1437,1832,1858]" degrees="24.83671" direction="north" orientation="latitude" pageId="11" pageNumber="312" precision="1" value="24.83671">24.83671°N</geoCoordinate>
<geoCoordinate id="3EF9A4AB2407171FFF60B234121B1D51" box="[151,288,1868,1895]" degrees="98.76185" direction="east" orientation="longitude" pageId="11" pageNumber="312" precision="1" value="98.76185">98.76185°E</geoCoordinate>
,
<quantity id="9C356F892407171FFEDCB23412B81D51" box="[299,387,1868,1895]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="11" pageNumber="312" unit="m" value="2067.0">
<elevation id="D0E0255F2407171FFEDCB23412B81D51" box="[299,387,1868,1895]" metricMagnitude="3" metricUnit="m" metricValue="2.0669999999999997" pageId="11" pageNumber="312" unit="m" value="2067.0">2067 m</elevation>
</quantity>
,
<date id="2F73E4AC2407171FFE67B234113E1D51" box="[400,517,1868,1895]" pageId="11" pageNumber="312" value="2005-05-28">
<collectingDate id="3F371D442407171FFE67B234113E1D51" box="[400,517,1868,1895]" pageId="11" pageNumber="312" value="2005-05-28">28.V.2005</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFDE6B23511441D51" box="[529,639,1869,1895]" pageId="11" pageNumber="312">H.M. Yan</collectorName>
leg. (
<collectionCode id="3DDC5AA92407171FFD48B23411C91D50" box="[703,754,1868,1894]" pageId="11" pageNumber="312">IOZ</collectionCode>
,
<collectionCode id="3DDC5AA92407171FFD09B234100A1D50" box="[766,817,1868,1894]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="312" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFCB1B23412A11D98" ID-GBIF-Occurrence="3821894534" collectingDate="2007-06-02" collectorName="D. W. Wrase" country="China" elevation="1900" latitude="24.807777" location="Baoshan Pref." longLatPrecision="21" longitude="98.534164" pageId="11" pageNumber="312" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFCB1B23410AF1D50" box="[838,916,1868,1894]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
,
<specimenCount id="4DCB09E52407171FFC57B234172B1D50" box="[928,1040,1868,1894]" pageId="11" pageNumber="312" type="female">3 females</specimenCount>
,
<location id="5E1294B72407171FFBEBB23417861D50" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFBEBB23417861D50" box="[1052,1213,1868,1894]" country="China" latitude="24.807777" longLatPrecision="21" longitude="98.534164" name="Baoshan Pref." pageId="11" pageNumber="312" stateProvince="Yunnan">Baoshan Pref.</location>
, mountain range
<locationDeviation id="4AC0DB4E2407171FFA77B23412611DBC" location="Tengchong" pageId="11" pageNumber="312">
<quantity id="9C356F892407171FFA77B23413801DBC" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="312" unit="km" value="25.0">25 km</quantity>
S Tengchong
</locationDeviation>
,
<quantity id="9C356F892407171FFE90B20812841DBD" box="[359,447,1904,1931]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="312" unit="m" value="1900.0">
<elevation id="D0E0255F2407171FFE90B20812841DBD" box="[359,447,1904,1931]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="11" pageNumber="312" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<geoCoordinate id="3EF9A4AB2407171FFE3BB208116B1DBC" box="[460,592,1904,1930]" degrees="24" direction="north" minutes="48" orientation="latitude" pageId="11" pageNumber="312" precision="15" seconds="28" value="24.807777">24°48'28&quot;N</geoCoordinate>
<geoCoordinate id="3EF9A4AB2407171FFDAEB20811E71DBC" box="[601,732,1904,1930]" degrees="98" direction="east" minutes="32" orientation="longitude" pageId="11" pageNumber="312" precision="15" seconds="03" value="98.534164">98°32'03&quot;E</geoCoordinate>
, devasted primary decid. forest, litter, sifted,
<date id="2F73E4AC2407171FFB16B208166E1DBC" box="[1249,1365,1904,1930]" pageId="11" pageNumber="312" value="2007-06-02">
<collectingDate id="3F371D442407171FFB16B208166E1DBC" box="[1249,1365,1904,1930]" pageId="11" pageNumber="312" value="2007-06-02">2.VI.2007</collectingDate>
</date>
,
<collectorName id="F638A7BA2407171FFA95B20913E61D99" pageId="11" pageNumber="312">D.W. Wrase</collectorName>
leg. [17] (cWR)
</materialsCitation>
;
<materialsCitation id="EBA5C8312407171FFE52B2EC160D1D98" ID-GBIF-Occurrence="3821894319" box="[421,1334,1940,1967]" collectingDate="2007-06-02" collectionCode="ZIN" collectorName="M. Schulke" country="China" elevation="1900" latitude="24.807777" location="Baoshan Pref." longLatPrecision="21" longitude="98.534164" pageId="11" pageNumber="312" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="4DCB09E52407171FFE52B2EC12C81D98" box="[421,499,1940,1966]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
,
<specimenCount id="4DCB09E52407171FFE08B2EC11551D98" box="[511,622,1940,1966]" pageId="11" pageNumber="312" type="female">6 females</specimenCount>
, same data but litter, fungi, sifted,
<collectorName id="F638A7BA2407171FFC06B2ED17491D98" box="[1009,1138,1940,1967]" pageId="11" pageNumber="312">M. Schülke</collectorName>
leg. (cWR,
<collectionCode id="3DDC5AA92407171FFB0DB2EC16161D98" box="[1274,1325,1940,1966]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="11" pageNumber="312" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5B72C26C2407171FFF30B2C011701DC0" blockId="11.[151,1437,1508,2039]" pageId="11" pageNumber="312">
<materialsCitation id="EBA5C8312407171FFF30B2C0117C1DC0" ID-GBIF-Occurrence="3821894570" collectingDate="2011-05-01" collectingDateMax="2011-05-31" collectingDateMin="2011-05-01" collectionCode="NME" country="Laos" elevation="1600" latitude="20.2" location="Phou Pan" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleul" pageId="11" pageNumber="312" specimenCount="1" specimenCount-male="1" stateProvince="Houaphan">
<emphasis id="69B91E7E2407171FFF30B2C012331DE4" bold="true" box="[199,264,1976,2002]" pageId="11" pageNumber="312">
<collectingCountry id="23DA82FC2407171FFF30B2C012381DE4" box="[199,259,1976,2002]" name="Laos" pageId="11" pageNumber="312">Laos</collectingCountry>
.
</emphasis>
<specimenCount id="4DCB09E52407171FFEE5B2C012581DE4" box="[274,355,1976,2002]" pageId="11" pageNumber="312" type="male">1 male</specimenCount>
, “NE-LAOS Pr.
<collectingRegion id="99090C8E2407171FFDD0B2C111A31DE4" box="[551,664,1976,2003]" country="Laos" name="Houaphan" pageId="11" pageNumber="312">Hua Phan</collectingRegion>
<collectingMunicipality id="BB1658162407171FFD55B2C110191DE4" box="[674,802,1976,2003]" pageId="11" pageNumber="312">Ban Saleul</collectingMunicipality>
,
<location id="5E1294B72407171FFCD8B2C010A61DE5" LSID="urn:lsid:plazi:treatment:D364737A2407171AFF60B00512301D22:5E1294B72407171FFCD8B2C010A61DE5" box="[815,925,1976,2003]" country="Laos" latitude="20.2" longLatPrecision="1268" longitude="104.01667" municipality="Ban Saleul" name="Phou Pan" pageId="11" pageNumber="312" stateProvince="Houaphan">Phou Pan</location>
(Mt.), ~
<geoCoordinate id="3EF9A4AB2407171FFBFCB2C117501DE2" box="[1035,1131,1977,2004]" degrees="20" direction="north" minutes="12" orientation="latitude" pageId="11" pageNumber="312" precision="925" value="20.2">20°12′N</geoCoordinate>
,
<geoCoordinate id="3EF9A4AB2407171FFB8EB2C117DF1DE2" box="[1145,1252,1977,2004]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="11" pageNumber="312" precision="925" value="104.01667">104°01′E</geoCoordinate>
,
<date id="2F73E4AC2407171FFB04B2C016AC1DE4" box="[1267,1431,1976,2002]" pageId="11" pageNumber="312" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">
<collectingDate id="3F371D442407171FFB04B2C016AC1DE4" box="[1267,1431,1976,2002]" pageId="11" pageNumber="312" value="2011-05-01" valueMax="2011-05-31" valueMin="2011-05-01">01.–31.V.2011</collectingDate>
</date>
,
<quantity id="9C356F892407171FFF60B2A4120F1DC1" box="[151,308,2012,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="11" pageNumber="312" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">
<elevation id="D0E0255F2407171FFF60B2A4120F1DC1" box="[151,308,2012,2039]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="11" pageNumber="312" unit="m" value="1600.0" valueMax="1900.0" valueMin="1300.0">1300–1900 m</elevation>
</quantity>
, local collector” (
<collectionCode id="3DDC5AA92407171FFE0CB2A411041DC0" box="[507,575,2012,2038]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="11" pageNumber="312">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="13D791E72400171AFF30B5EE12C31FB1" lastPageId="14" lastPageNumber="315" pageId="12" pageNumber="313" type="description">
<paragraph id="5B72C26C24001718FF30B5EE163D1A86" blockId="12.[151,1437,150,2013]" box="[199,1286,150,177]" pageId="12" pageNumber="313">
<emphasis id="69B91E7E24001718FF30B5EE126A1A86" bold="true" box="[199,337,150,176]" pageId="12" pageNumber="313">Description</emphasis>
(
<specimenCount id="4DCB09E524001718FE95B5EE12831A86" box="[354,440,150,177]" pageId="12" pageNumber="313" type="male">5 males</specimenCount>
and
<specimenCount id="4DCB09E524001718FE18B5EE11601A86" box="[495,603,150,177]" pageId="12" pageNumber="313" type="female">5 females</specimenCount>
measured). Body length 10.0–
<quantity id="9C356F8924001718FC43B5EE17211A87" box="[948,1050,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="12" pageNumber="313" unit="mm" value="11.4">11.4 mm</quantity>
. Habitus as in
<figureCitation id="C3F6DEE924001718FB49B5EE16391A86" box="[1214,1282,150,176]" captionStart="FIGURES 8–10" captionStartId="9.[151,264,1060,1084]" captionTargetBox="[188,1401,191,1028]" captionTargetId="figure-19@9.[178,1410,181,1036]" captionTargetPageId="9" captionText="FIGURES 8–10. Trichotichnus spp., habitus. 8, T. fukuharai Habu, 1957 (Yunnan); 9, T. angustitarsis sp. n. (S Tengchong); 10, T. subrobustus sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781687" httpUri="https://zenodo.org/record/6781687/files/figure.png" pageId="12" pageNumber="313">Fig. 9</figureCitation>
.
</paragraph>
<paragraph id="5B72C26C24001718FF30B5C217461B2A" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">Dorsal side of body black or dark brown, shiny, slightly iridescent on elytra; ventral side blackish brown. Mandibles basally, labrum and narrow lateral margins of pronotum reddish or yellowish brown; palpi and antennae brownish yellow; legs brownish yellow or brown, occasionally femora slightly infuscate.</paragraph>
<paragraph id="5B72C26C24001718FF30B45E16AB190E" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">
Head moderately sized (HWmax/PWmax 0.67–0.70, HWmin/PWmax 0.51–0.55). Eyes large and very convex, almost hemispherical (HWmax/HWmin 1.26–1.33), in lateral view almost round. Genae very narrow, narrower than antennomere 2 basally. Tempora very short, glabrous. Fronto-clypeal suture fine, shallow, slightly rounded. Frontoocular furrows fine, impressed at clypeus, reaching supraorbital furrows. Supraorbital furrows narrow, touching upper margin of eye. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eye and removed from supraorbital furrows by distance approximately equal to width of antennomere 1; in some specimens one or two very small unsetigerous punctures present medially of each supraorbital seta. Labrum very shallowly concave anteriorly. Clypeus somewhat flat, slightly emarginate along anterior margin, with a pair of setigerous pores each located at anterior angle.
<collectingRegion id="99090C8E24001718FE04B73F11691857" box="[499,594,583,609]" country="Myanmar" name="Chin" pageId="12" pageNumber="313">Mentum</collectingRegion>
separated from submentum, with moderately wide median tooth; epilobes narrow; submentum with one pair of long setae. Ligular sclerite markedly widened apically, truncate at apex, with one pair of ventroapical setae. Paraglossae narrow, markedly longer than ligular sclerite and separated from it by wide notches. Penultimate labial palpomere about as long as ultimate palpomere. Left mandible sharply bent in the apical fifth, blunted at tip. Dorsal microsculpture very fine, consisting of more or less isodiametric meshes, in some specimens highly obliterated. Antennae slender, surpassing pronotal basal edge by two apical antennomeres, with antennomeres 4–8 about three times as long as wide. Basal antennomere very slightly longer than antennomere 3.
</paragraph>
<paragraph id="5B72C26C24001718FF30B63B112D1EDE" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">Pronotum transverse (PWmax/PL 1.45–1.55), widest at the end of the anterior third (PWmax/PWmin 1.14– 1.19), more strongly narrowed apically than basally, with one lateral seta inserted in widest point. Sides rounded along entire length; lateral bead narrow, complete. Apical margin shallowly or moderately emarginate, very narrowly bordered along entire length, but occasionally border obliterated at the middle. Apical angles widely rounded, slightly protruding ahead. Basal margin slightly rounded in middle portion, scarcely oblique laterally, bordered along entire length, slightly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles very obtuse, with very small, occasionally almost indistinct obtuse denticle at tip. Disc moderately convex, strongly flattened at basal angles. Lateral depression very narrow in apical third, strongly widened in the basal half and fused with wide and shallow basal foveae; area between basal foveae convex. Median line distinct, superficial. Surface densely and finely punctate laterobasally and along sides, with sparser and finer punctures at base medially and at apical margin laterally. Microsculpture very fine, consisting of isodiametric (mostly anteriorly) and transverse meshes.</paragraph>
<paragraph id="5B72C26C24001718FF30B18A113F1C1A" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">
Elytra convex, elongate oval (EL/EW 1.44–1.54, EL/PL 2.85–3.14, EW/PWmax 1.28–1.38), widest in the apical third; sides slightly concave before the middle. Humeri angularly rounded, with a tiny denticle visible in caudal view. Subapical sinuation very shallow. Sutural angle acutangular, sharp or blunted at tip. Basal edge evenly arched or oblique, forming a very obtuse angle (internal humeral angle) with lateral margin. Striae impunctate, impressed along entire length, reaching anteriorly basal edge. Intervals slightly convex, moderately narrowed apically. Parascutellar (abbreviate) striole long, with a small setigerous pore basally. Interval 3 with a very small discal setigerous pore adjacent to stria
<quantity id="9C356F8924001718FE7DB0B2128E1FD2" box="[394,437,1482,1508]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="12" pageNumber="313" unit="in" value="2.0">2 in</quantity>
the beginning of apical third (in
<specimenCount id="4DCB09E524001718FCDDB0B310F41FD2" box="[810,975,1482,1509]" pageId="12" pageNumber="313" type="generic">one specimens</specimenCount>
two such pores present on each elytron). Marginal umbilicate series without distinct gap at middle. Microsculpture on intervals strongly obliterated, consisting of indistinct transverse lines.
</paragraph>
<paragraph id="5B72C26C24001718FF30B34E12F61C66" blockId="12.[151,1437,150,2013]" box="[199,461,1590,1616]" pageId="12" pageNumber="313">Wings fully developed.</paragraph>
<paragraph id="5B72C26C24001718FF30B323122D1CAE" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">
Prosternum with very fine short setae. Metepisternum (
<figureCitation id="C3F6DEE924001718FCC4B32210BF1C42" box="[819,900,1626,1652]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 &amp; 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="12" pageNumber="313">Fig. 21</figureCitation>
) markedly longer than wide, strongly narrowed posteriorly.
</paragraph>
<paragraph id="5B72C26C24001718FF30B3DB17CE1DEA" blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="313">
Legs relatively short. Metacoxae without additional setae medially. Metafemur ventrally with two setae at posterior margin and without setae at anterior margin. Protibia on dorsal side without longitudinal sulcus, in both sexes with three similar sized preapical spines on outer margin (
<figureCitation id="C3F6DEE924001718FCE0B39210521D32" box="[791,873,1770,1796]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 &amp; 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="12" pageNumber="313">Fig. 19</figureCitation>
). Tarsi glabrous dorsally, pro- and mesotarsomeres
<quantity id="9C356F8924001718FF60B27613DB1D1E" box="[151,224,1806,1832]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="12" pageNumber="313" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
both sexes with comparatively sparse moderately long ventro-lateral setae; tarsomere 5 with three or four pairs of ventro-lateral setae. Metatarsus slender, almost as long as or slightly shorter than HWmax, with tarsomeres weakly widened distally; metatarsomere 1 slightly shorter than metatarsomeres 2 and 3 combined; metatarsomeres 1–4 moderately densely setose ventrally. In male, pro- and mesotarsomeres 1–4 comparatively weakly widened (only slightly wider than in female) and with adhesive scales ventrally; pro- and mesotarsomeres 2 and 3 longer than wide. In both sexes, mesotarsomere 1 elongate, about as long as mesotarsomeres 2 and 3 combined.
</paragraph>
<caption id="0FB292E424011719FF60B2D317891DD0" ID-DOI="http://doi.org/10.5281/zenodo.6781701" ID-Zenodo-Dep="6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="13" pageNumber="314" startId="13.[151,264,1963,1987]" targetBox="[176,1424,201,1937]" targetPageId="13">
<paragraph id="5B72C26C24011719FF60B2D317891DD0" blockId="13.[151,1436,1963,2023]" pageId="13" pageNumber="314">
<emphasis id="69B91E7E24011719FF60B2D3126F1DF5" bold="true" box="[151,340,1963,1987]" pageId="13" pageNumber="314">FIGURES 18–22.</emphasis>
<taxonomicName id="9CCDB9EF24011719FEADB2D3112E1DF4" box="[346,533,1963,1987]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="69B91E7E24011719FEADB2D312DE1DF5" box="[346,485,1963,1987]" italics="true" pageId="13" pageNumber="314">Trichotichnus</emphasis>
spp.
</taxonomicName>
(18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18,
<taxonomicName id="9CCDB9EF24011719FB69B2D416A21DF5" authority="Habu, 1957" authorityName="Habu" authorityYear="1957" box="[1182,1433,1963,1987]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="fukuharai">
<emphasis id="69B91E7E24011719FB69B2D416211DF5" box="[1182,1306,1963,1987]" italics="true" pageId="13" pageNumber="314">T. fukuharai</emphasis>
<bibRefCitation id="3F5CBF9D24011719FAE8B2D316A21DF5" author="Habu, A." box="[1311,1433,1963,1987]" pageId="13" pageNumber="314" pagination="76 - 78" refId="ref37900" refString="Habu, A. (1957) Description of a new Trichotichnus - species. Akitu, 6, 76 - 78." type="journal article" year="1957">Habu, 1957</bibRefCitation>
</taxonomicName>
; 19 &amp; 21,
<taxonomicName id="9CCDB9EF24011719FF00B2A812A81DD1" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[247,403,2000,2023]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
<emphasis id="69B91E7E24011719FF00B2A812A81DD1" box="[247,403,2000,2023]" italics="true" pageId="13" pageNumber="314">T. angustitarsis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24011719FE6EB2B712E91DD1" bold="true" box="[409,466,1999,2023]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="728AA30524011719FE6EB2B712E91DD1" box="[409,466,1999,2023]" pageId="13" pageNumber="314" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 20,
<taxonomicName id="9CCDB9EF24011719FDF2B2A811A11DD1" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[517,666,1999,2023]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="hamulipenis" status="sp. nov.">
<emphasis id="69B91E7E24011719FDF2B2A811A11DD1" box="[517,666,1999,2023]" italics="true" pageId="13" pageNumber="314">T. hamulipenis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24011719FD57B2B711E21DD1" bold="true" box="[672,729,1999,2023]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="728AA30524011719FD57B2B711E21DD1" box="[672,729,1999,2023]" pageId="13" pageNumber="314" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 22,
<taxonomicName id="9CCDB9EF24011719FCFAB2A8109B1DD1" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[781,928,1999,2023]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="subrobustus" status="sp. nov.">
<emphasis id="69B91E7E24011719FCFAB2A8109B1DD1" box="[781,928,1999,2023]" italics="true" pageId="13" pageNumber="314">T. subrobustus</emphasis>
</taxonomicName>
<emphasis id="69B91E7E24011719FC51B2B710E41DD1" bold="true" box="[934,991,1999,2023]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="728AA30524011719FC51B2B710E41DD1" box="[934,991,1999,2023]" pageId="13" pageNumber="314" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="0FB292E42402171AFF60B6E210EB19CD" ID-DOI="http://doi.org/10.5281/zenodo.6781705" ID-Zenodo-Dep="6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="14" pageNumber="315" startId="14.[151,264,922,946]" targetBox="[160,1428,186,890]" targetPageId="14">
<paragraph id="5B72C26C2402171AFF60B6E210EB19CD" blockId="14.[151,1437,922,1019]" pageId="14" pageNumber="315">
<emphasis id="69B91E7E2402171AFF60B6E212681985" bold="true" box="[151,339,922,947]" pageId="14" pageNumber="315">FIGURES 23–27.</emphasis>
<taxonomicName id="9CCDB9EF2402171AFEAEB6E311571984" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[345,620,923,947]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
<emphasis id="69B91E7E2402171AFEAEB6E311571984" box="[345,620,923,947]" italics="true" pageId="14" pageNumber="315">Trichotichnus angustitarsis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2402171AFD85B6E311911985" bold="true" box="[626,682,923,947]" pageId="14" pageNumber="315">
<taxonomicNameLabel id="728AA3052402171AFD85B6E311911985" box="[626,682,923,947]" pageId="14" pageNumber="315" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
23, terminal lamella of aedeagus; 24 &amp; 25, median lobe of aedeagus (holotype); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 &amp; 25), lateral (24 &amp; 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 &amp; 27), B = 1.0 mm (24 &amp; 25).
</paragraph>
</caption>
<paragraph id="5B72C26C2402171AFF30B13516141EBD" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">Abdominal sternites without additional long setae; last visible abdominal sternite (VII) with one pair of marginal setae in male and with two pairs in female, in both sexes rounded at apex, but in male more widely.</paragraph>
<paragraph id="5B72C26C2402171AFF30B1ED10AA1F09" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">
Median lobe of aedeagus (
<figureCitation id="C3F6DEE92402171AFE08B1ED11B11E86" box="[511,650,1173,1200]" captionStart="FIGURES 23–27" captionStartId="14.[151,264,922,946]" captionTargetBox="[160,1428,186,890]" captionTargetId="figure-32@14.[151,1436,181,899]" captionTargetPageId="14" captionText="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 &amp; 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 &amp; 25), lateral (24 &amp; 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 &amp; 27), B = 1.0 mm (24 &amp; 25)." figureDoi="http://doi.org/10.5281/zenodo.6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="14" pageNumber="315">Figs. 23–25</figureCitation>
) in lateral view arcuate on dorsal side, with ventral margin almost straight in middle portion and with apex angularly bent dorsally; ventral side edged laterally in apical part and flattened between margins; terminal lamella (
<figureCitation id="C3F6DEE92402171AFD9EB1A511821EC1" box="[617,697,1245,1271]" captionStart="FIGURES 23–27" captionStartId="14.[151,264,922,946]" captionTargetBox="[160,1428,186,890]" captionTargetId="figure-32@14.[151,1436,181,899]" captionTargetPageId="14" captionText="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 &amp; 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 &amp; 25), lateral (24 &amp; 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 &amp; 27), B = 1.0 mm (24 &amp; 25)." figureDoi="http://doi.org/10.5281/zenodo.6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="14" pageNumber="315">Fig. 23</figureCitation>
) in dorsal view wide, wider than long, concave, strongly narrowed posteriorly, with sides very slightly concave just before narrowly rounded apex. Apical orifice in dorsal position, wide apically. Internal sac without spines, with spiny patch apically.
</paragraph>
<paragraph id="5B72C26C2402171AFF30B03112C31FB1" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">
Female genitalia (
<figureCitation id="C3F6DEE92402171AFE66B031110B1F55" box="[401,560,1353,1379]" captionStart="FIGURES 23–27" captionStartId="14.[151,264,922,946]" captionTargetBox="[160,1428,186,890]" captionTargetId="figure-32@14.[151,1436,181,899]" captionTargetPageId="14" captionText="FIGURES 23–27. Trichotichnus angustitarsis sp. n. 23, terminal lamella of aedeagus; 24 &amp; 25, median lobe of aedeagus (holo- type); 26, gonocoxite; 27, laterotergite, gonosubcoxite, and gonocoxite (Kongdang). Dorsal (23 &amp; 25), lateral (24 &amp; 26) and ventral (27) views. Scale bars: A = 0.5 mm (23, 26 &amp; 27), B = 1.0 mm (24 &amp; 25)." figureDoi="http://doi.org/10.5281/zenodo.6781705" httpUri="https://zenodo.org/record/6781705/files/figure.png" pageId="14" pageNumber="315">Figs. 26 &amp; 27</figureCitation>
): gonocoxite comparatively narrow in ventral view, about 0.6 times as long as gonosubcoxite, with wide base.
</paragraph>
</subSubSection>
<subSubSection id="13D791E72402171AFF30B0E9128A1C91" pageId="14" pageNumber="315" type="etymology">
<paragraph id="5B72C26C2402171AFF30B0E913EE1FF9" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">
<emphasis id="69B91E7E2402171AFF30B0E912751F9D" bold="true" box="[199,334,1425,1451]" pageId="14" pageNumber="315">Etymology.</emphasis>
The specific name is a Latin compound adjective, referring to the narrow pro- and mesotarsi of male.
</paragraph>
<paragraph id="5B72C26C2402171AFF30B0A1128A1C91" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">
<emphasis id="69B91E7E2402171AFF30B0A112591FC5" bold="true" box="[199,354,1497,1523]" pageId="14" pageNumber="315">Comparison.</emphasis>
This new species is very similar in habitus to
<taxonomicName id="9CCDB9EF2402171AFC98B0A210C31FC5" authorityName="Habu" authorityYear="1957" box="[879,1016,1497,1523]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fukuharai">
<emphasis id="69B91E7E2402171AFC98B0A210C31FC5" box="[879,1016,1497,1523]" italics="true" pageId="14" pageNumber="315">T. fukuharai</emphasis>
</taxonomicName>
, but differs in having pronotum relatively smaller, with deeper laterobasal depressions and more distinct microsculpture, and elytra with wider apical portions of intervals. In addition, the males of
<taxonomicName id="9CCDB9EF2402171AFD5DB35A106C1C0D" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[682,855,1570,1595]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
<emphasis id="69B91E7E2402171AFD5DB35A106C1C0D" box="[682,855,1570,1595]" italics="true" pageId="14" pageNumber="315">T. angustitarsis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2402171AFCA8B359109B1C0D" bold="true" box="[863,928,1569,1595]" pageId="14" pageNumber="315">
<taxonomicNameLabel id="728AA3052402171AFCA8B359109B1C0D" box="[863,928,1569,1595]" pageId="14" pageNumber="315" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are distinguished from those of
<taxonomicName id="9CCDB9EF2402171AFAE5B35A16A71C0D" authorityName="Habu" authorityYear="1957" box="[1298,1436,1569,1595]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fukuharai">
<emphasis id="69B91E7E2402171AFAE5B35A16A71C0D" box="[1298,1436,1569,1595]" italics="true" pageId="14" pageNumber="315">T. fukuharai</emphasis>
</taxonomicName>
by having pro- and mesotarsi narrower (see
<figureCitation id="C3F6DEE92402171AFD70B33D101F1C69" box="[647,804,1605,1631]" captionStart="FIGURES 18–22" captionStartId="13.[151,264,1963,1987]" captionTargetBox="[176,1424,201,1937]" captionTargetId="figure-19@13.[162,1425,181,1939]" captionTargetPageId="13" captionText="FIGURES 18–22. Trichotichnus spp. (18–20, right male protarsus; 21 &amp; 22, left metepisternum). 18, T. fukuharai Habu, 1957; 19 &amp; 21, T. angustitarsis sp. n.; 20, T. hamulipenis sp. n.; 22, T. subrobustus sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781701" httpUri="https://zenodo.org/record/6781701/files/figure.png" pageId="14" pageNumber="315">Figs. 18 &amp; 19</figureCitation>
), last abdominal sternite with one pair of marginal setae and very different structure of median lobe of aedeagus (bent terminal lamella apex, no thick spine in internal sac, etc.), as described above.
</paragraph>
</subSubSection>
<subSubSection id="13D791E72402171AFF30B3C912301D22" pageId="14" pageNumber="315" type="distribution">
<paragraph id="5B72C26C2402171AFF30B3C912301D22" blockId="14.[151,1437,1101,1812]" pageId="14" pageNumber="315">
<emphasis id="69B91E7E2402171AFF30B3C912621CFD" bold="true" box="[199,345,1713,1739]" pageId="14" pageNumber="315">Distribution</emphasis>
(
<figureCitation id="C3F6DEE92402171AFE9BB3C912FF1CFD" box="[364,452,1713,1739]" captionStart="FIGURE 28" captionStartId="15.[151,250,1700,1724]" captionTargetBox="[151,1435,181,1677]" captionTargetId="figure-19@15.[151,1435,181,1677]" captionTargetPageId="15" captionText="FIGURE 28. Trichotichnus angustitarsis sp. n., distribution." figureDoi="http://doi.org/10.5281/zenodo.6781707" httpUri="https://zenodo.org/record/6781707/files/figure.png" pageId="14" pageNumber="315">Fig. 28</figureCitation>
). Known from western
<collectingRegion id="99090C8E2402171AFD2DB3CA100B1CFA" box="[730,816,1714,1740]" country="China" name="Yunnan" pageId="14" pageNumber="315">Yunnan</collectingRegion>
(
<collectingCountry id="23DA82FC2402171AFCB5B3C910B31CFD" box="[834,904,1713,1739]" name="China" pageId="14" pageNumber="315">China</collectingCountry>
) and North West
<collectingCountry id="23DA82FC2402171AFBACB3CA17AE1CFA" box="[1115,1173,1714,1740]" name="Laos" pageId="14" pageNumber="315">Laos</collectingCountry>
. The species occurs at altitudes of
<quantity id="9C356F892402171AFED7B3AD12FB1CC6" box="[288,448,1749,1776]" metricMagnitude="3" metricUnit="m" metricValue="1.635" metricValueMax="1.97" metricValueMin="1.3" pageId="14" pageNumber="315" unit="m" value="1635.0" valueMax="1970.0" valueMin="1300.0">1300–1970 m</quantity>
. The range of
<taxonomicName id="9CCDB9EF2402171AFD9EB3AE102C1CD9" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[617,791,1750,1775]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="angustitarsis" status="sp. nov.">
<emphasis id="69B91E7E2402171AFD9EB3AE102C1CD9" box="[617,791,1750,1775]" italics="true" pageId="14" pageNumber="315">T. angustitarsis</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2402171AFCD7B3AD10591CD9" bold="true" box="[800,866,1749,1775]" pageId="14" pageNumber="315">
<taxonomicNameLabel id="728AA3052402171AFCD7B3AD10591CD9" box="[800,866,1749,1775]" pageId="14" pageNumber="315" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is probably located to the west of the range of
<taxonomicName id="9CCDB9EF2402171AFA7FB3AE123F1D25" authorityName="Habu" authorityYear="1957" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="fukuharai">
<emphasis id="69B91E7E2402171AFA7FB3AE123F1D25" italics="true" pageId="14" pageNumber="315">T. fukuharai</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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