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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.927.49351" ID-GBIF-Dataset="90af2e77-90a3-49b7-8f20-ff414446f743" ID-PMC="PMC7180164" ID-Pensoft-Pub="1313-2970-927-37" ID-Pensoft-UUID="5D57229A06E85536AAA7BB2CBE72841E" ID-PubMed="32341673" ID-ZooBank="1CD8D18D48A140E08B3C090FC223BB95" ModsDocID="1313-2970-927-37" checkinTime="1587138262110" checkinUser="pensoft" docAuthor="Krajcovicova, Katarina, Matyukhin, Aleksandr Vladimirovich &amp; Jana Christophoryova," docDate="2020" docId="BE47A517201B557FB297A5EB8B63071D" docLanguage="en" docName="ZooKeys 927: 37-51" docOrigin="ZooKeys 927" docSource="http://dx.doi.org/10.3897/zookeys.927.49351" docTitle="Tyrannochthonius eua Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová 2020, sp. nov." docType="treatment" docUuid="F65825EA-F0DC-4D86-92E5-8B7D67C38383" docUuidSource="ZooBank" docVersion="4" id="5D57229A06E85536AAA7BB2CBE72841E" lastPageNumber="37" masterDocId="5D57229A06E85536AAA7BB2CBE72841E" masterDocTitle="Two new pseudoscorpion species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) from the Tonga Islands, Polynesia, with a redescription of the genus Nesocheiridium" masterLastPageNumber="51" masterPageNumber="37" pageNumber="37" updateTime="1668168731898" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new pseudoscorpion species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) from the Tonga Islands, Polynesia, with a redescription of the genus Nesocheiridium</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Krajcovicova, Katarina</mods:namePart>
<mods:affiliation>Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska dolina, Ilkovicova 6, SK- 842 15 Bratislava, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">krajcovic.katarina@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matyukhin, Aleksandr Vladimirovich</mods:namePart>
<mods:affiliation>Institute of Ecology and Evolution, Severtsov Russian Academy of Sciences, Federal State Institution of Science, Leninsky pr. 33, 117 071 Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jana Christophoryova,</mods:namePart>
<mods:affiliation>Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska dolina, Ilkovicova 6, SK- 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>927</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>37</mods:start>
<mods:end>51</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.927.49351</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.927.49351</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-927-37</mods:identifier>
<mods:identifier type="ZooBank">1CD8D18D48A140E08B3C090FC223BB95</mods:identifier>
<mods:identifier type="Pensoft-UUID">5D57229A06E85536AAA7BB2CBE72841E</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="163643528" LSID="urn:lsid:zoobank.org:act:F65825EA-F0DC-4D86-92E5-8B7D67C38383" httpUri="http://treatment.plazi.org/id/BE47A517201B557FB297A5EB8B63071D" lastPageNumber="37" pageId="0" pageNumber="37">
<subSubSection pageId="0" pageNumber="37" type="nomenclature">
<paragraph pageId="0" pageNumber="37">
<taxonomicName LSID="BE47A517-201B-557F-B297-A5EB8B63071D" authority="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová, 2020" authorityName="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius eua" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="eua" status="sp. nov.">Tyrannochthonius eua</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the studied species: Tyrannochthonius eua sp. nov. (yellow triangle), Nesocheiridium stellatum (white circle), N. onevai sp. nov. (red circle)." figureDoi="10.3897/zookeys.927.49351.figure1" httpUri="https://binary.pensoft.net/fig/400616" pageId="0" pageNumber="37">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tyrannochthonius eua sp. nov., holotype male, dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.927.49351.figure2" httpUri="https://binary.pensoft.net/fig/400617" pageId="0" pageNumber="37">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="materials_examined">
<paragraph pageId="0" pageNumber="37">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
: Polynesia • ♂; Tonga, Eua [
<geoCoordinate degrees="21.387" direction="south" orientation="latitude" precision="55" value="-21.387">-21.387</geoCoordinate>
,
<geoCoordinate degrees="174.930" direction="west" orientation="longitude" precision="55" value="-174.93">-174.930</geoCoordinate>
]; 215 m a.s.l.; 11 Jul. 1980; Galina Fedorovna Kurcheva leg.; moss;
<emphasis bold="true" pageId="0" pageNumber="37">NHMW</emphasis>
29197.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="description">
<paragraph pageId="0" pageNumber="37">Description.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Adult male</emphasis>
(Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tyrannochthonius eua sp. nov., holotype male, dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.927.49351.figure2" httpUri="https://binary.pensoft.net/fig/400617" pageId="0" pageNumber="37">2</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3A</figureCitation>
): 0.97
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad; with four corneate eyes; epistome present, triangular; with 18 setae arranged 6: 4: 4: 2: 2; without furrows; with two pairs of small lyrifissures, first pair situated in ocular row, second pair situated lateral to setae of posterior row.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Coxae</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3B</figureCitation>
): coxa I with rounded apical projection, not bearing microsetae; chaetotaxy of coxae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3B</figureCitation>
): palpal coxae 3; pedal coxae I 3, II 4, III 5, IV 5. Coxa II with eight terminally incised spines, set in an oblique row (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3B, C</figureCitation>
). Intercoxal tubercle absent.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Chelicera</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3D</figureCitation>
): 1.53
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad; five setae on hand, all acuminate; moveable finger with one medial seta; fixed finger with 11, moveable finger with nine teeth; one ventral and two dorsal lyrifissures on hand; galea absent; serrula exterior with 15 blades; rallum consisting of seven bipinnate blades.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Pedipalp</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3E</figureCitation>
): all setae acuminate, femur setal formula: 5: 2: 1: 3: 5; trochanter 1.44
<normalizedToken originalValue="×">x</normalizedToken>
, femur 4.11
<normalizedToken originalValue="×">x</normalizedToken>
, patella 2.11
<normalizedToken originalValue="×">x</normalizedToken>
, chela 5.18
<normalizedToken originalValue="×">x</normalizedToken>
, hand 2.18
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad. Hand without spine-like seta, dorsal surface with a single row of five chemosensory setae between trichobothria
<emphasis italics="true" pageId="0" pageNumber="37">esb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">ib</emphasis>
/
<emphasis italics="true" pageId="0" pageNumber="37">isb</emphasis>
; hand and fixed chelal finger together with eight trichobothria, moveable chelal finger with four trichobothria;
<emphasis italics="true" pageId="0" pageNumber="37">ib</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">isb</emphasis>
close together, submedially on dorsum of chelal hand;
<emphasis italics="true" pageId="0" pageNumber="37">eb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">esb</emphasis>
close together, at base of fixed finger;
<emphasis italics="true" pageId="0" pageNumber="37">ist</emphasis>
distal to
<emphasis italics="true" pageId="0" pageNumber="37">eb</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">esb</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="37">it</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">est</emphasis>
less than one areolar diameter apart,
<emphasis italics="true" pageId="0" pageNumber="37">it</emphasis>
slightly distal to
<emphasis italics="true" pageId="0" pageNumber="37">est</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="37">et</emphasis>
near tip of finger; trichobothrium
<emphasis italics="true" pageId="0" pageNumber="37">st</emphasis>
of moveable finger sub-basally;
<emphasis italics="true" pageId="0" pageNumber="37">sb</emphasis>
slightly closer to
<emphasis italics="true" pageId="0" pageNumber="37">st</emphasis>
than to
<emphasis italics="true" pageId="0" pageNumber="37">b</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="37">b</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="37">t</emphasis>
subdistally,
<emphasis italics="true" pageId="0" pageNumber="37">t</emphasis>
at same level as
<emphasis italics="true" pageId="0" pageNumber="37">it</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="37">b</emphasis>
slightly basal to
<emphasis italics="true" pageId="0" pageNumber="37">est.</emphasis>
Chelal teeth heterodentate: fixed finger with three small teeth followed by 17 large, erect, well-spaced teeth, decreasing in size towards base, distally alternating with six small intercalary teeth; moveable finger with nine large, erect, well-spaced teeth, without intercalary teeth.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Opisthosoma</emphasis>
: tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-IX: 4: 4: 4: 4: 4: 5: 6: 6: 6. Sternal chaetotaxy II-IX: 10: 28: 15: 10: 10: 8: 8: 8 (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tyrannochthonius eua sp. nov., holotype male, dorsal A carapace B coxae C coxal spines D right chelicera E right chela, showing trichobothrial pattern F chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger: t - terminal, b - basal, sb - subbasal, st - subterminal; trichobothria of fixed chelal finger: dx - duplex trichobothria, et - exterior terminal, it - interior terminal, est - exterior subterminal, ist - interior subterminal, esb - exterior subbasal, eb - exterior basal, isb - interior subbasal, ib - interior basal. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37">3F</figureCitation>
). Sternal lyrifissures II-IX: 2: 2: 0: 0: 0: 0: 0: 0. Genitalia: sternite III with narrow V-shaped opening. Genitalia not studied in detail.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Leg I</emphasis>
: trochanter 1.13
<normalizedToken originalValue="×">x</normalizedToken>
, femur 4.40
<normalizedToken originalValue="×">x</normalizedToken>
, patella 2.75
<normalizedToken originalValue="×">x</normalizedToken>
, tibia 3.25
<normalizedToken originalValue="×">x</normalizedToken>
, tarsus 5.67
<normalizedToken originalValue="×">x</normalizedToken>
deeper than broad.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Leg IV</emphasis>
: trochanter 1.22
<normalizedToken originalValue="×">x</normalizedToken>
, femoropatella 1.88
<normalizedToken originalValue="×">x</normalizedToken>
, tibia 3.43
<normalizedToken originalValue="×">x</normalizedToken>
, metatarsus 2.00
<normalizedToken originalValue="×">x</normalizedToken>
, tarsus 6.67
<normalizedToken originalValue="×">x</normalizedToken>
deeper than broad. Tactile seta present on metatarsus of leg IV; arolium slightly shorter than claws, not divided; claws simple.
</paragraph>
<caption doi="10.3897/zookeys.927.49351.figure1" httpUri="https://binary.pensoft.net/fig/400616" pageId="0" pageNumber="37" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 1.</emphasis>
Distribution of the studied species:
<taxonomicName authorityName="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius eua" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">Tyrannochthonius eua</emphasis>
</taxonomicName>
sp. nov. (yellow triangle),
<taxonomicName authorityName="Beier" authorityYear="1957" class="Arachnida" family="Cheiridiidae" genus="Nesocheiridium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesocheiridium stellatum" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="stellatum">
<emphasis italics="true" pageId="0" pageNumber="37">Nesocheiridium stellatum</emphasis>
</taxonomicName>
(white circle),
<taxonomicName lsidName="N. onevai" pageId="0" pageNumber="37" rank="species" species="onevai">
<emphasis italics="true" pageId="0" pageNumber="37">N. onevai</emphasis>
</taxonomicName>
sp. nov. (red circle).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.927.49351.figure2" httpUri="https://binary.pensoft.net/fig/400617" pageId="0" pageNumber="37" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 2.</emphasis>
<taxonomicName authorityName="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius eua" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">Tyrannochthonius eua</emphasis>
</taxonomicName>
sp. nov., holotype male, dorsal. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="37">
<taxonomicName genus="Dimensions" lsidName="Dimensions" pageId="0" pageNumber="37" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Dimensions</emphasis>
</taxonomicName>
(length/width or, in the case of the legs, length/depth) in mm. Body length 1.10. Pedipalp: trochanter 0.13/0.09, femur 0.37/0.09, patella 0.19/0.09, chela 0.57/0.11, hand 0.24/0.11, fixed finger 0.31, moveable finger 0.33. Chelicera 0.26/0.17, moveable finger 0.15. Carapace 0.38/0.39. Leg I: trochanter 0.09/0.08, femur 0.22/0.05, patella 0.11/0.04, tibia 0.13/0.04, tarsus 0.17/0.03. Leg IV: trochanter 0.11/0.09, femoropatella 0.32/0.17, tibia 0.24/0.07, metatarsus 0.12/0.06, tarsus 0.20/0.03.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="etymology">
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="37">The specific epithet refers to the island of Eua, on which this species occurs. It is used as a noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="remarks">
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="37">
The presence of intercalary teeth on the fixed chelal finger but not on the moveable chelal finger is unusual in
<taxonomicName authorityName="Chamberlin" authorityYear="1929" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="37">Tyrannochthonius</emphasis>
</taxonomicName>
species. However, a few other species possess this combination:
<taxonomicName lsidName="T. convivus" pageId="0" pageNumber="37" rank="species" species="convivus">
<emphasis italics="true" pageId="0" pageNumber="37">T. convivus</emphasis>
</taxonomicName>
Beier, 1974,
<taxonomicName lsidName="T. brasiliensis" pageId="0" pageNumber="37" rank="species" species="brasiliensis">
<emphasis italics="true" pageId="0" pageNumber="37">T. brasiliensis</emphasis>
</taxonomicName>
Mahnert, 1979,
<taxonomicName lsidName="T. amazonicus" pageId="0" pageNumber="37" rank="species" species="amazonicus">
<emphasis italics="true" pageId="0" pageNumber="37">T. amazonicus</emphasis>
</taxonomicName>
Mahnert, 1979,
<taxonomicName lsidName="T. rex" pageId="0" pageNumber="37" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="37">T. rex</emphasis>
</taxonomicName>
Harvey, 1989, and
<taxonomicName lsidName="T. swiftae" pageId="0" pageNumber="37" rank="species" species="swiftae">
<emphasis italics="true" pageId="0" pageNumber="37">T. swiftae</emphasis>
</taxonomicName>
Muchmore, 1993 (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.76057" author="Beier, M" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="37" pagination="999 - 1017" refId="B6" refString="Beier, M, 1974. Pseudoscorpione aus Suedindien des Naturhistorischen Museums in Genf. Revue Suisse de Zoologie 81 (4): 999 - 1017, DOI: https://doi.org/10.5962/bhl.part.76057" title="Pseudoscorpione aus Suedindien des Naturhistorischen Museums in Genf." url="https://doi.org/10.5962/bhl.part.76057" volume="81" year="1974">Beier 1974</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.82338" author="Mahnert, V" journalOrPublisher="Revue Suisse Zoologie" pageId="0" pageNumber="37" pagination="719 - 810" refId="B22" refString="Mahnert, V, 1979. Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse Zoologie 86 (3): 719 - 810, DOI: https://doi.org/10.5962/bhl.part.82338" title="Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien)." url="https://doi.org/10.5962/bhl.part.82338" volume="86" year="1979">Mahnert 1979</bibRefCitation>
;
<bibRefCitation author="Harvey, MS" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="0" pageNumber="37" pagination="21 - 29" refId="B14" refString="Harvey, MS, 1989. Two new cavernicolous chthoniids from Australia, with notes on the generic placement of the south-western Pacific species attributed to the genera Paraliochthonius Beier and Morikawa Chamberlin (Pseudoscorpionida: Chthoniidae). Bulletin of the British Arachnological Society 8 (1): 21 - 29" title="Two new cavernicolous chthoniids from Australia, with notes on the generic placement of the south-western Pacific species attributed to the genera Paraliochthonius Beier and Morikawa Chamberlin (Pseudoscorpionida: Chthoniidae)." volume="8" year="1989">Harvey 1989</bibRefCitation>
;
<bibRefCitation author="Muchmore, WB" journalOrPublisher="Proceedings of the Entomological Society of Hawaii" pageId="0" pageNumber="37" pagination="147 - 162" refId="B29" refString="Muchmore, WB, 2000. The Pseudoscorpionida of Hawaii. Part I. Introduction and Chthonioidea. Proceedings of the Entomological Society of Hawaii 34: 147 - 162" title="The Pseudoscorpionida of Hawaii. Part I. Introduction and Chthonioidea." volume="34" year="2000">Muchmore 2000</bibRefCitation>
).
<taxonomicName authorityName="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius eua" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">Tyrannochthonius eua</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName lsidName="T. convivus" pageId="0" pageNumber="37" rank="species" species="convivus">
<emphasis italics="true" pageId="0" pageNumber="37">T. convivus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. amazonicus" pageId="0" pageNumber="37" rank="species" species="amazonicus">
<emphasis italics="true" pageId="0" pageNumber="37">T. amazonicus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. rex" pageId="0" pageNumber="37" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="37">T. rex</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="T. swiftae" pageId="0" pageNumber="37" rank="species" species="swiftae">
<emphasis italics="true" pageId="0" pageNumber="37">T. swiftae</emphasis>
</taxonomicName>
by the significantly shorter palpal femur length (0.37 mm, versus 0.42-0.49 mm in
<taxonomicName lsidName="T. convivus" pageId="0" pageNumber="37" rank="species" species="convivus">
<emphasis italics="true" pageId="0" pageNumber="37">T. convivus</emphasis>
</taxonomicName>
, 0.46-0.56 mm in
<taxonomicName lsidName="T. amazonicus" pageId="0" pageNumber="37" rank="species" species="amazonicus">
<emphasis italics="true" pageId="0" pageNumber="37">T. amazonicus</emphasis>
</taxonomicName>
, 1.24-1.34 mm in
<taxonomicName lsidName="T. rex" pageId="0" pageNumber="37" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="37">T. rex</emphasis>
</taxonomicName>
, and 0.53 mm in
<taxonomicName lsidName="T. swiftae" pageId="0" pageNumber="37" rank="species" species="swiftae">
<emphasis italics="true" pageId="0" pageNumber="37">T. swiftae</emphasis>
</taxonomicName>
). It also differs from
<taxonomicName lsidName="T. amazonicus" pageId="0" pageNumber="37" rank="species" species="amazonicus">
<emphasis italics="true" pageId="0" pageNumber="37">T. amazonicus</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="T. rex" pageId="0" pageNumber="37" rank="species" species="rex">
<emphasis italics="true" pageId="0" pageNumber="37">T. rex</emphasis>
</taxonomicName>
, and
<taxonomicName lsidName="T. swiftae" pageId="0" pageNumber="37" rank="species" species="swiftae">
<emphasis italics="true" pageId="0" pageNumber="37">T. swiftae</emphasis>
</taxonomicName>
in having a lower number of teeth on fixed chelal fingers. In contrast,
<taxonomicName lsidName="T. brasiliensis" pageId="0" pageNumber="37" rank="species" species="brasiliensis">
<emphasis italics="true" pageId="0" pageNumber="37">T. brasiliensis</emphasis>
</taxonomicName>
has a shorter palpal femur (length 0.28 mm) than
<taxonomicName lsidName="T. eua" pageId="0" pageNumber="37" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">T. eua</emphasis>
</taxonomicName>
sp. nov., as well as lower number of coxal spines on coxae II (4-5, versus 8 in
<taxonomicName lsidName="T. eua" pageId="0" pageNumber="37" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">T. eua</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
<caption doi="10.3897/zookeys.927.49351.figure3" httpUri="https://binary.pensoft.net/fig/400618" pageId="0" pageNumber="37" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 3.</emphasis>
<taxonomicName authorityName="Krajčovičová &amp; Matyukhin &amp; Jana Christophoryová" authorityYear="2020" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tyrannochthonius eua" order="Pseudoscorpiones" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="eua">
<emphasis italics="true" pageId="0" pageNumber="37">Tyrannochthonius eua</emphasis>
</taxonomicName>
sp. nov., holotype male, dorsal
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
carapace
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
coxae
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
coxal spines
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
right chelicera
<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
right chela, showing trichobothrial pattern
<emphasis bold="true" pageId="0" pageNumber="37">F</emphasis>
chaetotaxy of genital area (sternites II-III). Abbreviations: trichobothria of moveable chelal finger:
<emphasis italics="true" pageId="0" pageNumber="37">t</emphasis>
-terminal,
<emphasis italics="true" pageId="0" pageNumber="37">b</emphasis>
-basal,
<emphasis italics="true" pageId="0" pageNumber="37">sb</emphasis>
-subbasal,
<emphasis italics="true" pageId="0" pageNumber="37">st</emphasis>
-subterminal; trichobothria of fixed chelal finger:
<emphasis italics="true" pageId="0" pageNumber="37">dx</emphasis>
-duplex trichobothria,
<emphasis italics="true" pageId="0" pageNumber="37">et</emphasis>
-exterior terminal,
<emphasis italics="true" pageId="0" pageNumber="37">it</emphasis>
-interior terminal,
<emphasis italics="true" pageId="0" pageNumber="37">est</emphasis>
-exterior subterminal,
<emphasis italics="true" pageId="0" pageNumber="37">ist</emphasis>
-interior subterminal,
<emphasis italics="true" pageId="0" pageNumber="37">esb</emphasis>
-exterior subbasal,
<emphasis italics="true" pageId="0" pageNumber="37">eb</emphasis>
-exterior basal,
<emphasis italics="true" pageId="0" pageNumber="37">isb</emphasis>
-interior subbasal,
<emphasis italics="true" pageId="0" pageNumber="37">ib</emphasis>
-interior basal. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>