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<mods:title id="40B5A9E163BE1AF58D9854247E016C94">Studies in the New World tribe Anisoscelini Laporte, 1832 (Heteroptera: Coreidae: Coreinae): updated distributions, nomenclatural changes, and keys to the genera of the tribe and the species of Anisoscelis Latreille, 1829 and Bitta Osuna, 1984</mods:title>
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<mods:namePart id="D359C8945634A53A4F459B3503AFAA8B">Leavengood, John M.</mods:namePart>
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<paragraph id="8BC836D22118582AA882F970FE3D0DC7" blockId="16.[151,505,1759,1823]" box="[151,505,1759,1786]" pageId="16" pageNumber="257">
<heading id="D08081BE2118582AA882F970FE3D0DC7" bold="true" box="[151,505,1759,1786]" fontSize="11" level="1" pageId="16" pageNumber="257" reason="1">
<taxonomicName id="4C774D512118582AA882F970FE3D0DC7" ID-CoL="3XRF4" authority="Stal, 1865" authorityName="Stal" authorityYear="1865" box="[151,505,1759,1786]" class="Insecta" family="Coreidae" genus="Malvana" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="serrulata">
<emphasis id="B903EAC02118582AA882F970FE3D0DC7" bold="true" box="[151,505,1759,1786]" pageId="16" pageNumber="257">
<emphasis id="B903EAC02118582AA882F970FEB30DC7" bold="true" box="[151,375,1759,1786]" italics="true" pageId="16" pageNumber="257">Malvana serrulata</emphasis>
<bibRefCitation id="EFE64B232118582AA96AF970FE3D0DC7" author="Stal, C." box="[383,505,1759,1786]" pageId="16" pageNumber="257" pagination="163 - 188" refId="ref21729" refString="Stal, C. (1865) Hemiptera nova vel minus cognita. Annales de la Societe Entomologique de France, 4 (5), 163 - 188." type="journal article" year="1865">Stål, 1865</bibRefCitation>
</emphasis>
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<paragraph id="8BC836D22118582AA882F8ABFE460C22" blockId="16.[151,505,1759,1823]" box="[151,386,1796,1823]" pageId="16" pageNumber="257">
(
<figureCitation id="134C2A572118582AA88AF8ABFEE30C22" box="[159,295,1796,1823]" captionStart="FIGURES 1925" captionStartId="12.[151,264,1978,2002]" captionTargetBox="[151,1437,725,1955]" captionTargetId="figure-210@12.[151,1437,725,1955]" captionTargetPageId="12" captionText="FIGURES 1925. 19. Leptoscelis divisa Stål; 20. Leptoscelis elongator (Fabricius); 21. Leptoscelis excellens Stål; 2225. Malvana serrulata Stål." figureDoi="http://doi.org/10.5281/zenodo.10692988" httpUri="https://zenodo.org/record/10692988/files/figure.png" pageId="16" pageNumber="257">Figs. 2225</figureCitation>
,
<figureCitation id="134C2A572118582AA926F8ABFEBF0C22" box="[307,379,1796,1823]" captionStart="FIGURES 5056" captionStartId="17.[151,264,1608,1632]" captionTargetBox="[151,1436,453,1585]" captionTargetId="figure-120@17.[151,1436,453,1585]" captionTargetPageId="17" captionText="FIGURES 5056. 50. Ugnius badius (Herrich-Schäffer); 51. Phthiacnemia picta (Drury); 52. Mamurius mopsus Stål; 5354. Malvana serrulata Stål; 55. Phthia lunata (Fabricius); 56. Phthiarella decorata (Stål)." figureDoi="http://doi.org/10.5281/zenodo.10693004" httpUri="https://zenodo.org/record/10693004/files/figure.png" pageId="16" pageNumber="257">5354</figureCitation>
)
</paragraph>
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<subSubSection id="C36D65592118582AA8D2F8E2FC9F0C5B" box="[199,859,1867,1895]" pageId="16" pageNumber="257" type="reference_group">
<paragraph id="8BC836D22118582AA8D2F8E2FC9F0C5B" blockId="16.[151,1437,1867,2039]" box="[199,859,1867,1895]" pageId="16" pageNumber="257">
=
<taxonomicName id="4C774D512118582AA8CBF8E4FD640C5B" ID-CoL="LBZL" authority="Uhler, 1869" authorityName="Uhler" authorityYear="1869" box="[222,672,1867,1894]" class="Insecta" family="Coreidae" genus="Belonomus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="annulaticornis" status="syn. nov.">
<emphasis id="B903EAC02118582AA8CBF8E4FDD60C5B" bold="true" box="[222,530,1867,1894]" italics="true" pageId="16" pageNumber="257">Belonomus annulaticornis</emphasis>
<bibRefCitation id="EFE64B232118582AAA0DF8E3FD640C5B" author="Uhler, P. R." box="[536,672,1868,1894]" pageId="16" pageNumber="257" pagination="321 - 327" refId="ref21807" refString="Uhler, P. R. (1869) Notices of the Hemiptera obtained by the expedition of Prof. James Orton in Ecuador and Brazil. Proceedings of the Boston Society of Natural History, 12, 321 - 327." type="journal article" year="1869">Uhler, 1869</bibRefCitation>
</taxonomicName>
,
<emphasis id="B903EAC02118582AAABEF8E3FC9F0C5B" bold="true" box="[683,859,1868,1894]" pageId="16" pageNumber="257">
<taxonomicNameLabel id="A23057BB2118582AAABEF8E3FC9F0C5B" box="[683,859,1868,1894]" pageId="16" pageNumber="257">new synonymy</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36D65592118582AA8D2F8DFFC390CCA" pageId="16" pageNumber="257" type="discussion">
<paragraph id="8BC836D22118582AA8D2F8DFFC390CCA" blockId="16.[151,1437,1867,2039]" pageId="16" pageNumber="257">
<emphasis id="B903EAC02118582AA8D2F8DFFEFF0CB7" bold="true" box="[199,315,1904,1930]" pageId="16" pageNumber="257">Remarks.</emphasis>
After careful examination of the
<typeStatus id="54CC88702118582AAABEF8DEFD220CB6" box="[683,742,1905,1931]" pageId="16" pageNumber="257">types</typeStatus>
of each species as well as many non-type specimens,
<taxonomicName id="4C774D512118582AAD2CF8DFFF3A0C93" class="Insecta" family="Coreidae" genus="Malvana" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="serrulata">
<emphasis id="B903EAC02118582AAD2CF8DFFF3A0C93" italics="true" pageId="16" pageNumber="257">Malvana serrulata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C774D512118582AA927F83BFD9C0C93" authorityName="Uhler" authorityYear="1869" box="[306,600,1940,1966]" class="Insecta" family="Coreidae" genus="Belonomus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="annulaticornis">
<emphasis id="B903EAC02118582AA927F83BFD9C0C93" box="[306,600,1940,1966]" italics="true" pageId="16" pageNumber="257">Belonomus annulaticornis</emphasis>
</taxonomicName>
are considered conspecific. Their known distributions have been combined herein and the ventral and posterior pyogphore are illustrated (
<figureCitation id="134C2A572118582AAB5EF817FC110CEE" box="[843,981,1976,2003]" captionStart="FIGURES 1925" captionStartId="12.[151,264,1978,2002]" captionTargetBox="[151,1437,725,1955]" captionTargetId="figure-210@12.[151,1437,725,1955]" captionTargetPageId="12" captionText="FIGURES 1925. 19. Leptoscelis divisa Stål; 20. Leptoscelis elongator (Fabricius); 21. Leptoscelis excellens Stål; 2225. Malvana serrulata Stål." figureDoi="http://doi.org/10.5281/zenodo.10692988" httpUri="https://zenodo.org/record/10692988/files/figure.png" pageId="16" pageNumber="257">Figs. 2425</figureCitation>
). As
<taxonomicName id="4C774D512118582AAC1FF816FB080CEF" authorityName="Uhler" authorityYear="1869" box="[1034,1228,1976,2002]" class="Insecta" family="Coreidae" genus="Belonomus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="species" species="annulaticornis">
<emphasis id="B903EAC02118582AAC1FF816FB080CEF" box="[1034,1228,1976,2002]" italics="true" pageId="16" pageNumber="257">B. annulaticornis</emphasis>
</taxonomicName>
is the only species of the genus, this likewise places
<taxonomicName id="4C774D512118582AAA1AF873FD4F0CCB" authorityName="Uhler" authorityYear="1869" box="[527,651,2012,2038]" class="Insecta" family="Coreidae" genus="Belonomus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAC02118582AAA1AF873FD4F0CCB" box="[527,651,2012,2038]" italics="true" pageId="16" pageNumber="257">Belonomus</emphasis>
</taxonomicName>
in junior synonymy of
<taxonomicName id="4C774D512118582AAB86F873FC330CCB" authorityName="Stal" authorityYear="1865" box="[915,1015,2012,2038]" class="Insecta" family="Coreidae" genus="Malvana" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="257" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAC02118582AAB86F873FC330CCB" box="[915,1015,2012,2038]" italics="true" pageId="16" pageNumber="257">Malvana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36D65592119582BA8D2FF39FCE70BC4" pageId="17" pageNumber="258" type="distribution">
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<emphasis id="B903EAC02119582BA8D2FF39FEA50B8D" bold="true" box="[199,353,150,176]" pageId="17" pageNumber="258">Distribution.</emphasis>
Known from
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,
<collectingCountry id="F36076422119582BAA4BFF38FD610B8C" box="[606,677,151,177]" name="Brazil" pageId="17" pageNumber="258">Brazil</collectingCountry>
,
<collectingCountry id="F36076422119582BAAA4FF38FCE90B8C" box="[689,813,151,177]" name="Costa Rica" pageId="17" pageNumber="258">Costa Rica</collectingCountry>
,
<collectingCountry id="F36076422119582BAB2DFF38FC510B8C" box="[824,917,151,177]" name="Ecuador" pageId="17" pageNumber="258">Ecuador</collectingCountry>
and
<collectingCountry id="F36076422119582BABD9FF38FC3B0B8C" box="[972,1023,151,177]" name="Peru" pageId="17" pageNumber="258">Peru</collectingCountry>
(
<bibRefCitation id="EFE64B232119582BAC18FF38FB100B8C" author="Packauskas, R." box="[1037,1236,151,177]" pageId="17" pageNumber="258" pagination="1 - 270" refId="ref21254" refString="Packauskas, R. (2010) Catalog of the Coreidae, or leaf-footed bugs of the New World. Fort Hays Studies, Series 4, 5, 1 - 270. https: // doi. org / 10.58809 / HCWH 1818" type="journal article" year="2010">Packauskas 2010</bibRefCitation>
).
<bibRefCitation id="EFE64B232119582BACF3FF38FF260BE8" author="Mattei, B. &amp; Mattei, B." pageId="17" pageNumber="258" refId="ref20946" refString="Mattei, B. &amp; Mattei, B. (2017) Hemiptera, Heteroptera, Coreidae: Catalogo de las especies hasta ahora encontradas en los alrededores de Puerto Ayacucho, Municipio Atures, Estado Amazonas, Venezuela. The Field Museum Field Guides. Guide 854, 10 pp." type="book" year="2017">Mattei &amp; Mattei (2017)</bibRefCitation>
reported this species from
<collectingRegion id="49B3F8302119582BAA00FF14FD4A0BE8" box="[533,654,187,213]" country="Venezuela" name="Amazonas" pageId="17" pageNumber="258">Amazonas</collectingRegion>
,
<collectingCountry id="F36076422119582BAA8CFF14FCCB0BE8" box="[665,783,187,213]" name="Venezuela" pageId="17" pageNumber="258">Venezuela</collectingCountry>
, which was formally published in
<bibRefCitation id="EFE64B232119582BAC83FF14FA860BE8" author="Cazorla, D." box="[1174,1346,187,213]" pageId="17" pageNumber="258" pagination="1 - 91" refId="ref19108" refString="Cazorla, D. (2021) Coreidae (Hemiptera: Heteroptera) de Venezuela. Revista Nicaraguense de Entomologia, 246, 1 - 91." type="journal article" year="2021">Cazorla (2021)</bibRefCitation>
. French
<collectingCountry id="F36076422119582BA882FF70FF350BC4" box="[151,241,223,249]" name="Guyana" pageId="17" pageNumber="258">Guyana</collectingCountry>
,
<collectingCountry id="F36076422119582BA8E9FF70FEA00BC4" box="[252,356,223,249]" name="Paraguay" pageId="17" pageNumber="258">Paraguay</collectingCountry>
and
<collectingCountry id="F36076422119582BA98EFF70FDC20BC4" box="[411,518,223,249]" name="Suriname" pageId="17" pageNumber="258">Suriname</collectingCountry>
are new country records.
</paragraph>
</subSubSection>
<subSubSection id="C36D65592119582BA8D2FEADFD700A90" pageId="17" pageNumber="258" type="materials_examined">
<paragraph id="8BC836D22119582BA8D2FEADFD700A90" blockId="17.[151,1437,150,429]" pageId="17" pageNumber="258">
<emphasis id="B903EAC02119582BA8D2FEADFD7F0A20" bold="true" box="[199,699,258,285]" pageId="17" pageNumber="258">Material Examined. New Records. French</emphasis>
<materialsCitation id="3B1F3C8F2119582BAAD4FEADFB470A7C" collectingDate="14 - 28 - XII- 2013, 8 - 28 - I- 2017, 9 - 15 - IV- 2017" collectionCode="MNHN, UNAM" collectorName="P. H. Dalens &amp; S. Brulle" country="Guyana" latitude="4.7200003" location="Montagne des Cheveaux" longLatPrecision="15" longitude="-52.416702" pageId="17" pageNumber="258" specimenCount="1" stateProvince="Concepcion">
<emphasis id="B903EAC02119582BAAD4FEADFCED0A21" bold="true" box="[705,809,258,284]" pageId="17" pageNumber="258">
<collectingCountry id="F36076422119582BAAD4FEADFCE00A21" box="[705,804,258,284]" name="Guyana" pageId="17" pageNumber="258">Guyana</collectingCountry>
:
</emphasis>
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,
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,
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,
<collectingDate id="EF8DE9FA2119582BA882FE88FD510A7C" box="[151,661,294,321]" isEnumeration="true" pageId="17" pageNumber="258">
<date id="FFC910122119582BA882FE88FE950A7C" box="[151,337,295,321]" pageId="17" pageNumber="258" value="2013-12-14" valueMax="2013-12-28" valueMin="2013-12-14">1428-XII-2013</date>
,
<date id="FFC910122119582BA94EFE88FE2E0A7C" box="[347,490,295,321]" pageId="17" pageNumber="258" value="2017-01-08" valueMax="2017-01-28" valueMin="2017-01-08">828-I-2017</date>
,
<date id="FFC910122119582BA9E1FE89FD510A7C" box="[500,661,294,321]" pageId="17" pageNumber="258" value="2017-04-09" valueMax="2017-04-15" valueMin="2017-04-09">915-IV-2017</date>
</collectingDate>
,
<collectorName id="268253042119582BAA8BFE88FCEE0A7C" box="[670,810,295,321]" pageId="17" pageNumber="258">P. H. Dalens</collectorName>
,
<collectorName id="268253042119582BAB26FE88FC510A7C" box="[819,917,295,321]" pageId="17" pageNumber="258">S. Brulle</collectorName>
(
<collectionCode id="ED66AE172119582BABB6FE88FC390A7C" box="[931,1021,295,321]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="258">MNHN</collectionCode>
,
<collectionCode id="ED66AE172119582BAC12FE88FB9B0A7C" box="[1031,1119,295,321]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="258">UNAM</collectionCode>
, 7)
</materialsCitation>
;
<materialsCitation id="3B1F3C8F2119582BAC9BFE88FDA90A58" collectorName="Kilm" country="Paraguay" location="Horqueta" pageId="17" pageNumber="258" specimenCount="1" stateProvince="Concepcion">
<emphasis id="B903EAC02119582BAC9BFE88FAC50A7C" bold="true" box="[1166,1281,295,321]" pageId="17" pageNumber="258">
<collectingCountry id="F36076422119582BAC9BFE88FAC50A7C" box="[1166,1281,295,321]" name="Paraguay" pageId="17" pageNumber="258">Paraguay</collectingCountry>
</emphasis>
:
<collectingRegion id="49B3F8302119582BAD1AFE88FA5D0A7C" box="[1295,1433,295,321]" country="Paraguay" name="Concepcion" pageId="17" pageNumber="258">Concepción</collectingRegion>
:
<location id="8EA860092119582BA882FEE4FEC60A58" LSID="urn:lsid:plazi:treatment:03DE87C42118582BA882F970FD700A90:8EA860092119582BA882FEE4FEC60A58" box="[151,258,331,357]" country="Paraguay" name="Horqueta" pageId="17" pageNumber="258" stateProvince="Concepcion">Horqueta</location>
, 57-10 W 23-24 N, 44
<collectorName id="268253042119582BAA14FEE4FDFF0A58" box="[513,571,331,357]" pageId="17" pageNumber="258">Kilm</collectorName>
east
</materialsCitation>
<materialsCitation id="3B1F3C8F2119582BAA66FEE4FAC30A58" box="[627,1287,331,357]" collectingDate="1986-01-13" collectorName="Alberto Schulze &amp; Drake" country="Paraguay" location="Riv." pageId="17" pageNumber="258" specimenCount="1" specimenCount-male="1" stateProvince="Asuncion">
<collectingRegion id="49B3F8302119582BAA66FEE4FD1F0A58" box="[627,731,331,357]" country="Paraguay" name="Asuncion" pageId="17" pageNumber="258">Paraguay</collectingRegion>
<location id="8EA860092119582BAAF7FEE4FCD60A58" LSID="urn:lsid:plazi:treatment:03DE87C42118582BA882F970FD700A90:8EA860092119582BAAF7FEE4FCD60A58" box="[738,786,331,357]" country="Paraguay" name="Riv." pageId="17" pageNumber="258" stateProvince="Asuncion">Riv.</location>
,
<date id="FFC910122119582BAB08FEE4FC4B0A58" box="[797,911,331,357]" pageId="17" pageNumber="258" value="1986-01-13">
<collectingDate id="EF8DE9FA2119582BAB08FEE4FC4B0A58" box="[797,911,331,357]" pageId="17" pageNumber="258" value="1986-01-13">13-I-1986</collectingDate>
</date>
,
<collectorName id="268253042119582BAB8CFEE4FB8A0A58" box="[921,1102,331,357]" pageId="17" pageNumber="258">Alberto Schulze</collectorName>
(
<collectorName id="268253042119582BAC49FEE4FB610A58" box="[1116,1189,331,357]" pageId="17" pageNumber="258">Drake</collectorName>
,
<specimenCount id="9D71FD5B2119582BACBAFEE4FB3B0A58" box="[1199,1279,331,357]" count="1" pageId="17" pageNumber="258" type="male">1 male</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B1F3C8F2119582BAD07FEE4FBA00AB4" collectingDate="1950-11-13" collectionCode="NMNH" collectorName="Col. Independencia &amp; Juan Foerster &amp; Lutz Collection" country="Paraguay" location="Guaira" pageId="17" pageNumber="258" specimenCount="1" specimenCount-male="1" stateProvince="Guaira">
<collectingRegion id="49B3F8302119582BAD07FEE4FAA70A58" box="[1298,1379,331,357]" country="Paraguay" name="Guaira" pageId="17" pageNumber="258">Guairá</collectingRegion>
:
<collectorName id="268253042119582BAD78FEE4FEF90AB4" pageId="17" pageNumber="258">Col. Independencia</collectorName>
,
<date id="FFC910122119582BA95DFEC1FE0B0AB4" box="[328,463,366,393]" pageId="17" pageNumber="258" value="1950-11-13">
<collectingDate id="EF8DE9FA2119582BA95DFEC1FE0B0AB4" box="[328,463,366,393]" pageId="17" pageNumber="258" value="1950-11-13">13-XI-1950</collectingDate>
</date>
,
<collectorName id="268253042119582BA9CEFEC0FDB70AB4" box="[475,627,367,393]" pageId="17" pageNumber="258">Juan Foerster</collectorName>
, J C
<collectorName id="268253042119582BAABCFEC0FC9D0AB4" box="[681,857,367,393]" pageId="17" pageNumber="258">Lutz Collection</collectorName>
1961 (
<collectionCode id="ED66AE172119582BABB3FEC0FBC40AB4" box="[934,1024,367,393]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="258" type="Museum">NMNH</collectionCode>
,
<specimenCount id="9D71FD5B2119582BAC19FEC0FB980AB4" box="[1036,1116,367,393]" count="1" pageId="17" pageNumber="258" type="male">1 male</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B1F3C8F2119582BAC7AFEC1FD750A90" collectingDate="20 - II- 1963, 8 - XI- 1963" collectionCode="UNAM" collectorName="Geijakes" country="Suriname" location="Paramaribo" pageId="17" pageNumber="258" specimenCount="1" stateProvince="Paramaribo">
<emphasis id="B903EAC02119582BAC7AFEC1FB250AB5" bold="true" box="[1135,1249,366,392]" pageId="17" pageNumber="258">
<collectingCountry id="F36076422119582BAC7AFEC1FB180AB5" box="[1135,1244,366,392]" name="Suriname" pageId="17" pageNumber="258">Surinam</collectingCountry>
:
</emphasis>
<collectingRegion id="49B3F8302119582BACFDFEC0FAA80AB4" box="[1256,1388,367,393]" country="Suriname" name="Paramaribo" pageId="17" pageNumber="258">Paramaribo</collectingRegion>
,
<collectingDate id="EF8DE9FA2119582BAD62FEC0FEB60A90" isEnumeration="true" pageId="17" pageNumber="258">
<date id="FFC910122119582BAD62FEC0FF290A90" pageId="17" pageNumber="258" value="1963-02-20">20- II-1963</date>
,
<date id="FFC910122119582BA8ECFE3CFEB60A90" box="[249,370,403,429]" pageId="17" pageNumber="258" value="1963-11-08">8-XI-1963</date>
</collectingDate>
, D. C.
<collectorName id="268253042119582BA9D5FE3CFDE60A90" box="[448,546,403,429]" pageId="17" pageNumber="258">Geijakes</collectorName>
(
<collectionCode id="ED66AE172119582BAA24FE3CFD4F0A90" box="[561,651,403,429]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="17" pageNumber="258">UNAM</collectionCode>
, 2)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>