613 lines
62 KiB
XML
613 lines
62 KiB
XML
<document id="103C16BDA9C9F6EB364ABFD73E955053" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. & Huijbregts, J." docDate="2017" docId="03E5BB4DE827FFD9AAE4F9BEFF0BFDC2" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus bisscrutator Krikken & Huijbregts, new species" docType="treatment" docVersion="7" lastPageNumber="312" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="304" updateTime="1698437609924" updateUser="ExternalLinkService">
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<mods:mods id="BB939B827B88EF8CB86718337B721445" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="230D2FE08894C69607A4C73C139F3F20">
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<mods:title id="59B12AE90F1D33465E87B24BB4EFFB2B">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F4AF5D798E1CC8D3347521B43CBA4B3B" type="personal">
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<mods:roleTerm id="85966714B5FE054EF85B25E79456808C">Author</mods:roleTerm>
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<mods:namePart id="82E1E3B4276C0D2922947F0CE2AE9015">Krikken, J.</mods:namePart>
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</mods:name>
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<mods:name id="3C76D864232834B150041FE6CCD6C6EA" type="personal">
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<mods:roleTerm id="4A0021C831C86BAD76CFCFE43D0D42E7">Author</mods:roleTerm>
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<mods:namePart id="54B8268F080467F5930DCDFE3FC84FCF">Huijbregts, J.</mods:namePart>
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<mods:typeOfResource id="A9992B8E17D9D9D44DAB52E9019DB3A6">text</mods:typeOfResource>
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<mods:titleInfo id="081F70DF0553394E8446DA78743D3AC6">
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<mods:title id="485526A07A7808C507A35D41334AABA3">Zootaxa</mods:title>
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<mods:part id="19A5F764F501CC5A9D76D82539D93B6B">
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<mods:date id="0ADB57C2A41850B53D810F32EF337746">2017</mods:date>
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<mods:detail id="2741B89D9EFE05A827EA822868A63D4D" type="volume">
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<mods:number id="2968A5273DCB6C102DC52E194AFA7E53">4238</mods:number>
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<mods:detail id="DDD751F13DD23888C41ED5D8870800E6" type="issue">
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<mods:number id="62DA4745AF30E725EFC9960B05050D96">3</mods:number>
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<mods:identifier id="285F4FA41E58B7CD07B0CAB15945479B" type="CLB-Dataset">36345</mods:identifier>
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<mods:identifier id="6E2510491AAC70529E96C84438C75005" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
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<mods:identifier id="782499A453333F975220359B8579BBC5" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03E5BB4DE827FFD9AAE4F9BEFF0BFDC2" ID-DOI="http://doi.org/10.5281/zenodo.6030177" ID-GBIF-Taxon="127719088" ID-Zenodo-Dep="6030177" LSID="urn:lsid:plazi:treatment:03E5BB4DE827FFD9AAE4F9BEFF0BFDC2" httpUri="http://treatment.plazi.org/id/03E5BB4DE827FFD9AAE4F9BEFF0BFDC2" lastPageId="11" lastPageNumber="312" pageId="3" pageNumber="304">
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<subSubSection id="C35659D0E827FFD1AAE4F9BEFEAEF9B8" pageId="3" pageNumber="304" type="nomenclature">
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<paragraph id="8BF30A5BE827FFD1AAE4F9BEFC55F99A" blockId="3.[151,911,1574,1634]" box="[151,911,1574,1600]" pageId="3" pageNumber="304">
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<heading id="D0BBBD37E827FFD1AAE4F9BEFC55F99A" bold="true" box="[151,911,1574,1600]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
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<emphasis id="B938D649E827FFD1AAE4F9BEFC55F99A" bold="true" box="[151,911,1574,1600]" pageId="3" pageNumber="304">
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<taxonomicName id="4C4C71D8E827FFD1AAE4F9BEFD28F99A" ID-CoL="6SMKD" authority="Krikken & Huijbregts" authorityName="Krikken & Huijbregts" box="[151,754,1574,1600]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="bisscrutator" status="sp. nov.">
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<emphasis id="B938D649E827FFD1AAE4F9BEFE08F99A" bold="true" box="[151,466,1574,1600]" italics="true" pageId="3" pageNumber="304">Onthophagus bisscrutator</emphasis>
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Krikken & Huijbregts
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20B6B32E827FFD1A88CF9BEFC55F99A" box="[767,911,1574,1600]" pageId="3" pageNumber="304" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F9D1FEAEF9B8" blockId="3.[151,911,1574,1634]" box="[151,372,1609,1634]" pageId="3" pageNumber="304">
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(
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<figureCitation id="137716DEE827FFD1AAECF9D1FF36F9B8" box="[159,236,1609,1634]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[207,1396,233,1891]" captionTargetId="figure@5.[177,1413,195,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Sulawesi Onthophagus species, oblique habitus. 1, O. bisscrutator, holotype male from Toraut. 2, O. scrutator, male from Moramo. 3, O. bongkudai, holotype male from Moajat. 4, O. sangirensis, male from Toraut. 5, O. holosericus, male from Marena, 6, female from Marena. 7, O. sopu, holotype male from Sopu. 8, O. manguliensis, male from Mangoli. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345757/files/figure.png" pageId="3" pageNumber="304">Figs. 1</figureCitation>
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,
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<figureCitation id="137716DEE827FFD1AA89F9D2FEC2F9B8" box="[250,280,1610,1634]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1935,1958]" captionTargetBox="[162,1436,205,1903]" captionTargetId="figure@7.[151,1436,193,1915]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Sulawesi Onthophagus species, dorsal forebody. 13, O. bisscrutator, holotype male from Toraut. 14, O. scrutator, male from Moramo. 15, O. bongkudai, holotype male from Moajat 16, paratype female. 17, O. sangirensis, male from Toraut. 18, O. sangirensis, female from Tangkoko. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345759/files/figure.png" pageId="3" pageNumber="304">13</figureCitation>
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,
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<figureCitation id="137716DEE827FFD1AB56F9D1FEB6F9B8" box="[293,364,1609,1634]" captionStart="FIGURES 31 – 38" captionStartId="10.[151,264,1674,1697]" captionTargetBox="[151,1366,302,1606]" captionTargetId="figure@10.[151,1436,193,1653]" captionTargetPageId="10" captionText="FIGURES 31 – 38. Sulawesi Onthophagus, body parts. 31 –, O. bisscrutator, head full-face, holotype male from Toraut. 32, O. bisscrutator, left elytron dorsal, holotype male from Toraut. 33, O. bisscrutator, protibial upper side, holotype male from Toraut. 34, O. bisscrutator, metatibial underside, paratype male from Mogogonipa. 35, O. bisscrutator, aedeagus lateral, paratype male from Toraut. 36, O. bisscrutator, head full-face, paratype female from Toraut. 37, O. scrutator, head full-face, female from Matanyo. 38, O. scrutator, aedeagus lateral, male from Morano. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345762/files/figure.png" pageId="3" pageNumber="304">31–36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35659D0E827FFD6AAE4F909FE47F93A" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304" type="materials_examined">
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<paragraph id="8BF30A5BE827FFD1AAE4F909FA47F970" blockId="3.[151,1437,1681,2030]" box="[151,1437,1681,1706]" pageId="3" pageNumber="304">
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<materialsCitation id="3B240006E827FFD1AAE4F909FA47F970" ID-GBIF-Occurrence="1438340097" box="[151,1437,1681,1706]" collectionCode="RMNH" country="Indonesia" location="Dumoga Bone National Park" pageId="3" pageNumber="304" specimenCount="1" stateProvince="North Sulawesi" typeStatus="holotype">
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<emphasis id="B938D649E827FFD1AAE4F909FE5FF970" bold="true" box="[151,389,1681,1706]" pageId="3" pageNumber="304">Material examined.</emphasis>
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<typeStatus id="54F7B4F9E827FFD1ABE2F909FE23F970" box="[401,505,1681,1706]" pageId="3" pageNumber="304" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="ED5D929EE827FFD1A83DF90AFD7CF970" box="[590,678,1682,1706]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="304">RMNH</collectionCode>
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) from
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<collectingCountry id="F35B4ACBE827FFD1A88FF909FCB7F970" box="[764,877,1681,1706]" name="Indonesia" pageId="3" pageNumber="304">Indonesia</collectingCountry>
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:
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<collectingRegion id="4988C4B9E827FFD1A90FF909FBE8F970" box="[892,1074,1681,1706]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="304">North Sulawesi</collectingRegion>
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:
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<location id="8E935C80E827FFD1AE31F90AFA43F970" LSID="urn:lsid:plazi:treatment:03E5BB4DE827FFD9AAE4F9BEFF0BFDC2:8E935C80E827FFD1AE31F90AFA43F970" box="[1090,1433,1681,1706]" country="Indonesia" name="Dumoga Bone National Park" pageId="3" pageNumber="304" stateProvince="North Sulawesi">Dumoga Bone National Park</location>
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:
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F92EFD78F928" blockId="3.[151,1437,1681,2030]" pageId="3" pageNumber="304">
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Toraut R3, alt.
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<quantity id="4CB4A7BEE827FFD1AB4CF92DFE52F917" box="[319,392,1717,1742]" metricMagnitude="2" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="304" unit="m" value="245.0">245 m</quantity>
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,
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<date id="FFF22C9BE827FFD1ABE7F92DFD92F914" box="[404,584,1717,1742]" pageId="3" pageNumber="304" value="1985-08-19" valueMax="1985-08-27" valueMin="1985-08-19">19–27.viii.1985</date>
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||
, J. Huijbregts HH433; multistral evergreen forest, 4 papaya traps. The
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<specimenCount id="9D4AC1D2E827FFD1AA8AF941FE4FF928" box="[249,405,1753,1778]" pageId="3" pageNumber="304" type="generic" typeStatus="paratypes">216 paratypes</specimenCount>
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are summarized below:
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F966FCD3F8CC" blockId="3.[151,1437,1681,2030]" box="[151,777,1789,1814]" pageId="3" pageNumber="304">
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||
<collectingRegion id="4988C4B9E827FFD1AAE4F966FE57F8CC" box="[151,397,1790,1814]" country="Indonesia" name="Sulawesi Utara" pageId="3" pageNumber="304">
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||
<collectionCode id="ED5D929EE827FFD1AAE4F966FE57F8CC" box="[151,397,1790,1814]" pageId="3" pageNumber="304">NORTH SULAWESI</collectionCode>
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||
</collectingRegion>
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:
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<collectionCode id="ED5D929EE827FFD1ABE4F966FDB2F8CC" box="[407,616,1790,1814]" pageId="3" pageNumber="304">DUMOGA BONE</collectionCode>
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National Park
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F8BAFA46F8E3" blockId="3.[151,1437,1681,2030]" box="[151,1436,1825,1850]" pageId="3" pageNumber="304">
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- Mount Mogogonipa,
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<quantity id="4CB4A7BEE827FFD1ABCAF8BAFDD9F8E3" box="[441,515,1826,1850]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="304" unit="m" value="400.0">400 m</quantity>
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||
,
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<date id="FFF22C9BE827FFD1A862F8B9FD1FF8E0" box="[529,709,1825,1850]" pageId="3" pageNumber="304" value="1985-08-22" valueMax="1985-08-26" valueMin="1985-08-22">22–26.viii.1985</date>
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, J. Huijbregts, hh424, multistratal evergreen forest, 2 fish traps,
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F8DEFE47F887" blockId="3.[151,1437,1681,2030]" box="[151,413,1861,1886]" pageId="3" pageNumber="304">29 paratypes in RMNH</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F8F2FA46F858" blockId="3.[151,1437,1681,2030]" box="[151,1436,1897,1922]" pageId="3" pageNumber="304">
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||
- Mount Mogogonipa,
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<quantity id="4CB4A7BEE827FFD1ABC6F8F1FE24F85B" box="[437,510,1897,1922]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="304" unit="m" value="600.0">600 m</quantity>
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||
,
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<date id="FFF22C9BE827FFD1A879F8F1FD65F858" box="[522,703,1897,1922]" pageId="3" pageNumber="304" value="1985-08-22" valueMax="1985-08-26" valueMin="1985-08-22">22–26.viii.1985</date>
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||
, J. Huijbregts, hh425, multistratal evergreen forest, 2 fish traps, 3
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</paragraph>
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<paragraph id="8BF30A5BE827FFD1AAE4F816FEA0F87F" blockId="3.[151,1437,1681,2030]" box="[151,378,1933,1958]" pageId="3" pageNumber="304">paratypes in RMNH</paragraph>
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||
<paragraph id="8BF30A5BE827FFD1AAE4F82AFA47F813" blockId="3.[151,1437,1681,2030]" box="[151,1437,1969,1994]" pageId="3" pageNumber="304">
|
||
- Page Subcamp,
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||
<quantity id="4CB4A7BEE827FFD1ABF7F82AFE0BF813" box="[388,465,1970,1994]" metricMagnitude="2" metricUnit="m" metricValue="3.02" pageId="3" pageNumber="304" unit="m" value="302.0">302 m</quantity>
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||
,
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||
<date id="FFF22C9BE827FFD1AB91F829FDB0F810" box="[482,618,1969,1994]" pageId="3" pageNumber="304" value="1985-09-03" valueMax="1985-09-08" valueMin="1985-09-03">3–8.ix.1985</date>
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||
, J. Huijbregts, hh439, multistratal evergreen forest, human excrement
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</paragraph>
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||
<paragraph id="8BF30A5BE827FFD1AAE4F84DFCE4F837" blockId="3.[151,1437,1681,2030]" box="[151,830,2005,2030]" pageId="3" pageNumber="304">
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||
baited,
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||
<collectingMethod id="520D724CE827FFD1AA9AF84DFDC8F837" box="[233,530,2005,2030]" pageId="3" pageNumber="304">baffled canopy funnel trap</collectingMethod>
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0 m, 1 paratype in RMNH
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||
</paragraph>
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||
<paragraph id="8BF30A5BE820FFD6AAE4FF00FEA0FF2D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Poniki Trail: Station 0 29,
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||
<quantity id="4CB4A7BEE820FFD6AB9AFF0FFD9AFF75" box="[489,576,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="305" unit="m" value="1600.0">1600 m</quantity>
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||
,
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<date id="FFF22C9BE820FFD6A83FFF0FFD41FF6A" box="[588,667,151,176]" pageId="4" pageNumber="305" value="1985-10">x.1985</date>
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||
, J. Van Stalle, 6 paratypes in IRSNB - Toraut R
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||
<quantity id="4CB4A7BEE820FFD6AB40FF24FE7BFF0E" box="[307,417,188,213]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
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||
<date id="FFF22C9BE820FFD6ABC0FF24FDBAFF0F" box="[435,608,188,213]" pageId="4" pageNumber="305" value="1985-05-31" valueMax="1985-06-08" valueMin="1985-05-31">31.v–8-vi.1985</date>
|
||
, J. Huijbregts, hh357, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AE9EFF24FAB4FF0E" box="[1261,1390,188,213]" pageId="4" pageNumber="305">fungi traps</collectingMethod>
|
||
, 36 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FE9DFEB5FEE5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB5FFE9CFE57FEC6" box="[300,397,260,285]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABEBFE9CFDFFFEC7" box="[408,549,260,285]" pageId="4" pageNumber="305" value="1985-05-08" valueMax="1985-05-10" valueMin="1985-05-08">8–10.v.1985</date>
|
||
, J. Huijbregts, hh311, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AE0FFE9CFA59FEC6" box="[1148,1411,260,285]" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 5 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FED5FE21FE5D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB5FFED4FE55FEBE" box="[300,399,332,357]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABE8FED4FDEFFEBF" box="[411,565,332,357]" pageId="4" pageNumber="305" value="1985-05-10" valueMax="1985-05-18" valueMin="1985-05-10">10–18.v.1985</date>
|
||
, J, Huijbregts, hh320, multistratal evergreen forest,
|
||
<collectingMethod id="520D724CE820FFD6AE0FFED4FF19FE5D" pageId="4" pageNumber="305">human excrement canopy trap</collectingMethod>
|
||
0 m, 3 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FE0DFF36FE15" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB5EFE0CFE55FE76" box="[301,399,404,429]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABE8FE0CFDD6FE77" box="[411,524,404,429]" pageId="4" pageNumber="305" value="1985-05-20">20.v.1985</date>
|
||
, J. Huijbregts, hh326, multistratal evergreen forest, decaying fungi, 1 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FE45FED2FDCD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB5FFE44FE57FE2E" box="[300,397,476,501]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABEAFE44FDE9FE2F" box="[409,563,476,501]" pageId="4" pageNumber="305" value="1985-05-20" valueMax="1985-05-23" valueMin="1985-05-20">20–23.v.1985</date>
|
||
, J. Huijbregts, hh332, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AEFEFE44FAC6FE2E" box="[1165,1308,476,501]" pageId="4" pageNumber="305">banana traps</collectingMethod>
|
||
, 1 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FDBDFEA0FD85" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB41FDBCFE44FDE6" box="[306,414,548,573]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABDCFDBCFDB9FDE7" box="[431,611,548,573]" pageId="4" pageNumber="305" value="1985-08-15" valueMax="1985-08-19" valueMin="1985-08-15">15–19.viii.1985</date>
|
||
, J. Huijbregts, hh415, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AE9AFDBDFAA7FDE6" box="[1257,1405,549,573]" pageId="4" pageNumber="305">papaya traps</collectingMethod>
|
||
, 5 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FDF5FC8DFD7D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB41FDF4FE44FD5E" box="[306,414,620,645]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABDCFDF4FDB9FD5F" box="[431,611,620,645]" pageId="4" pageNumber="305" value="1985-08-16" valueMax="1985-08-20" valueMin="1985-08-16">16–20.viii.1985</date>
|
||
, J. Huijbregts, hh422, multistratal evergreen forest, human excrement baited,
|
||
<collectingMethod id="520D724CE820FFD6AA9AFD17FDC8FD7D" box="[233,530,655,680]" pageId="4" pageNumber="305">baffled canopy funnel trap</collectingMethod>
|
||
0 m, 12 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FD2DFEA0FD35" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut R
|
||
<quantity id="4CB4A7BEE820FFD6AB42FD2CFE47FD16" box="[305,413,692,717]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="4" pageNumber="305" unit="m" value="3245.0">3, 245 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABDEFD2CFD83FD17" box="[429,601,692,717]" pageId="4" pageNumber="305" value="1985-07-19" valueMax="1985-07-27" valueMin="1985-07-19">19–27.vii.1985</date>
|
||
, J. Huijbregts, hh433, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AEAFFD2DFAAAFD16" box="[1244,1392,693,717]" pageId="4" pageNumber="305">papaya traps</collectingMethod>
|
||
, 21 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FD65FF36FCED" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut,
|
||
<quantity id="4CB4A7BEE820FFD6AB6EFD64FEBDFCCE" box="[285,359,764,788]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="4" pageNumber="305" unit="m" value="234.0">234 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6AB00FD64FDC4FCCF" box="[371,542,764,789]" pageId="4" pageNumber="305" value="1985-05-31" valueMax="1985-06-08" valueMin="1985-05-31">31.v–8.vi.1985</date>
|
||
, J. Huijbregts, hh356, multistratal evergreen forest,
|
||
<collectingMethod id="520D724CE820FFD6AE17FD64FB29FCCE" box="[1124,1267,764,789]" pageId="4" pageNumber="305">window trap</collectingMethod>
|
||
, 5 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FCDDFE55FCA5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut,
|
||
<quantity id="4CB4A7BEE820FFD6AB6EFCDCFEBCFC86" box="[285,358,836,860]" metricMagnitude="2" metricUnit="m" metricValue="2.17" pageId="4" pageNumber="305" unit="m" value="217.0">217 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6AB02FCDCFE26FC87" box="[369,508,836,861]" pageId="4" pageNumber="305" value="1985-05-09" valueMax="1985-05-11" valueMin="1985-05-09">9–11.v.1985</date>
|
||
, J. Huijbregts, hh313, multistratal evergreen forest edge, 4
|
||
<collectingMethod id="520D724CE820FFD6AEE2FCDCFA42FC86" box="[1169,1432,836,861]" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 2 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FC15FE09FC1D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut,
|
||
<quantity id="4CB4A7BEE820FFD6AB52FC14FEB4FC7E" box="[289,366,908,932]" metricMagnitude="2" metricUnit="m" metricValue="2.17" pageId="4" pageNumber="305" unit="m" value="217.0">217 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6AB0DFC14FDC2FC7F" box="[382,536,908,933]" pageId="4" pageNumber="305" value="1985-05-11" valueMax="1985-05-18" valueMin="1985-05-11">11–18.v.1985</date>
|
||
, J. Huijbregts, hh321, multistratal evergreen forest edge, 4
|
||
<collectingMethod id="520D724CE820FFD6AEA3FC14FF0AFC1D" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 3 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FC4DFE09FBD5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut,
|
||
<quantity id="4CB4A7BEE820FFD6AB52FC4CFEB4FC36" box="[289,366,980,1004]" metricMagnitude="2" metricUnit="m" metricValue="2.17" pageId="4" pageNumber="305" unit="m" value="217.0">217 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6AB0EFC4CFDC2FC37" box="[381,536,980,1005]" pageId="4" pageNumber="305" value="1985-05-18" valueMax="1985-05-22" valueMin="1985-05-18">18–22.v.1985</date>
|
||
, J. Huijbregts, hh329, multistratal evergreen forest edge, 4
|
||
<collectingMethod id="520D724CE820FFD6AEA2FC4CFF0AFBD5" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 6 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FB85FF36FB8D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut,
|
||
<quantity id="4CB4A7BEE820FFD6AB6DFB84FEBDFBEE" box="[286,359,1052,1076]" metricMagnitude="2" metricUnit="m" metricValue="2.34" pageId="4" pageNumber="305" unit="m" value="234.0">234 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6AB00FB84FDFDFBEF" box="[371,551,1052,1077]" pageId="4" pageNumber="305" value="1985-08-15" valueMax="1985-08-21" valueMin="1985-08-15">15–21.viii.1985</date>
|
||
, J. Huijbregts, hh416, multistratal evergreen forest,
|
||
<collectingMethod id="520D724CE820FFD6AE1DFB84FB24FBEE" box="[1134,1278,1052,1077]" pageId="4" pageNumber="305">window trap</collectingMethod>
|
||
, 1 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FBFDFE47FB7A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Toraut: Maze,
|
||
<quantity id="4CB4A7BEE820FFD6AB18FBFCFE6EFBA6" box="[363,436,1124,1148]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="305" unit="m" value="220.0">220 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABB3FBFCFD8CFBA7" box="[448,598,1124,1149]" pageId="4" pageNumber="305" value="1985-11-09" valueMax="1985-11-15" valueMin="1985-11-09">9–15.xi.1985</date>
|
||
, J. Krikken, pw23, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AEFCFBFCFA42FBA6" box="[1167,1432,1124,1149]" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 20 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FB34FCADFB1E" blockId="4.[151,1437,151,2013]" box="[151,887,1196,1220]" pageId="4" pageNumber="305">
|
||
<collectingRegion id="4988C4B9E820FFD6AAE4FB34FE57FB1E" box="[151,397,1196,1220]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="305">
|
||
<collectionCode id="ED5D929EE820FFD6AAE4FB34FE57FB1E" box="[151,397,1196,1220]" pageId="4" pageNumber="305">NORTH SULAWESI</collectionCode>
|
||
</collectingRegion>
|
||
:
|
||
<collectionCode id="ED5D929EE820FFD6ABE4FB34FCADFB1E" box="[407,887,1196,1220]" pageId="4" pageNumber="305">MOUNT AMBANG NATURE RESERVE</collectionCode>
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FB49FE13FAD6" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB1BFB48FE69FB32" box="[360,435,1231,1256]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB95FB48FDE5FB32" box="[486,575,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="305" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A83EFB57FD32FB32" box="[589,744,1231,1256]" pageId="4" pageNumber="305" value="1985-05-29" valueMax="1985-05-30" valueMin="1985-05-29">29–30.v.1985</date>
|
||
, J. Huijbregts, hh342, coffee plantation, 4
|
||
<collectingMethod id="520D724CE820FFD6AEA1FB57FF0AFAD6" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 1 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FA81FD9EFA8E" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB1BFA80FE6BFAEA" box="[360,433,1303,1328]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB90FA80FDE1FAEA" box="[483,571,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="4" pageNumber="305" unit="m" value="1120.0">1120 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A83BFA8FFD05FAEA" box="[584,735,1303,1328]" pageId="4" pageNumber="305" value="1985-09-09" valueMax="1985-09-13" valueMin="1985-09-09">9–13.ix.1985</date>
|
||
, J. Huijbregts, hh443a, multistratal evergreen forest, 2
|
||
<collectingMethod id="520D724CE820FFD6AF23FA8FFE91FA8E" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 1 paratype in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FAF9FE09FA46" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB19FAF8FE6CFAA2" box="[362,438,1375,1400]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB9EFAF8FD9CFAA2" box="[493,582,1376,1400]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="4" pageNumber="305" unit="m" value="1120.0">1120 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A825FAC7FD31FAA2" box="[598,747,1375,1400]" pageId="4" pageNumber="305" value="1985-09-09" valueMax="1985-09-13" valueMin="1985-09-09">9–13.ix.1985</date>
|
||
, J. Huijbregts, hh443b, multistratal evergreen forest, 2
|
||
<collectingMethod id="520D724CE820FFD6AF01FAC7FF0AFA46" pageId="4" pageNumber="305">fish traps</collectingMethod>
|
||
, 4 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FA31FE3BFA3E" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB19FA30FE6CFA1A" box="[362,438,1447,1472]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB9FFA30FD9CFA1A" box="[492,582,1448,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="4" pageNumber="305" unit="m" value="1080.0">1080 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A826FA3FFD36FA1A" box="[597,748,1447,1472]" pageId="4" pageNumber="305" value="1985-09-09" valueMax="1985-09-13" valueMin="1985-09-09">9–13.ix.1985</date>
|
||
, J. Huijbregts, hh443, multistratal evergreen forest, 4
|
||
<collectingMethod id="520D724CE820FFD6AF12FA3FFF0AFA3E" pageId="4" pageNumber="305">fungi traps</collectingMethod>
|
||
, 14 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4FA69FD78F9F6" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB1BFA68FE69F9D2" box="[360,435,1519,1544]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB95FA68FDE4F9D2" box="[486,574,1520,1544]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="305" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A83FFA77FD2BF9D2" box="[588,753,1519,1544]" pageId="4" pageNumber="305" value="1985-09-10" valueMax="1985-09-12" valueMin="1985-09-10">10–12.ix.1985</date>
|
||
, J. Huijbregts, hh446a, multistratal evergreen forest edge, 2
|
||
<collectingMethod id="520D724CE820FFD6AAE4F98CFE45F9F6" box="[151,415,1556,1581]" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 3 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4F9A1FDDEF9AE" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Kotamobagu (
|
||
<quantity id="4CB4A7BEE820FFD6AB1BF9A0FE68F98A" box="[360,434,1591,1616]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="305" unit="km" value="20.0">20 km</quantity>
|
||
E),
|
||
<quantity id="4CB4A7BEE820FFD6AB96F9A0FDE4F98A" box="[485,574,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="305" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6A83FF9AFFD2AF98A" box="[588,752,1591,1616]" pageId="4" pageNumber="305" value="1985-09-10" valueMax="1985-09-12" valueMin="1985-09-10">10–12.ix.1985</date>
|
||
, J. Huijbregts, hh446b, multistratal evergreen forest edge, 2
|
||
<collectingMethod id="520D724CE820FFD6AAE4F9C4FEDBF9AE" box="[151,257,1628,1653]" pageId="4" pageNumber="305">fish traps</collectingMethod>
|
||
, 5 paratypes in RMNH
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4F918FCF3F942" blockId="4.[151,1437,151,2013]" box="[151,809,1664,1688]" pageId="4" pageNumber="305">
|
||
<collectingRegion id="4988C4B9E820FFD6AAE4F918FE57F942" box="[151,397,1664,1688]" country="Indonesia" name="Sulawesi Utara" pageId="4" pageNumber="305">
|
||
<collectionCode id="ED5D929EE820FFD6AAE4F918FE57F942" box="[151,397,1664,1688]" pageId="4" pageNumber="305">NORTH SULAWESI</collectionCode>
|
||
</collectingRegion>
|
||
: TANGKOKO
|
||
<collectionCode id="ED5D929EE820FFD6A849F918FCF3F942" box="[570,809,1664,1688]" pageId="4" pageNumber="305">NATURE RESERVE</collectionCode>
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAE4F93DFE47F93A" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
- Pantai Trail,
|
||
<quantity id="4CB4A7BEE820FFD6AB25F93CFE45F966" box="[342,415,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="305" unit="m" value="100.0">100 m</quantity>
|
||
,
|
||
<date id="FFF22C9BE820FFD6ABD8F93CFD95F967" box="[427,591,1700,1725]" pageId="4" pageNumber="305" value="1985-11-27" valueMax="1985-11-29" valueMin="1985-11-27">27–29.xi.1985</date>
|
||
, J. Krikken, pw41a, multistratal evergreen forest, 2
|
||
<collectingMethod id="520D724CE820FFD6AEE3F93CFA42F966" box="[1168,1432,1700,1725]" pageId="4" pageNumber="305">human excrement traps</collectingMethod>
|
||
, 28 paratypes in RMNH
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35659D0E820FFD9AAB4F974FEC8FE0A" lastPageId="11" lastPageNumber="312" pageId="4" pageNumber="305" type="description">
|
||
<paragraph id="8BF30A5BE820FFD6AAB4F974FAD7F8F2" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">
|
||
<emphasis id="B938D649E820FFD6AAB4F974FE83F8DF" bold="true" box="[199,345,1772,1797]" pageId="4" pageNumber="305">Description.</emphasis>
|
||
<typeStatus id="54F7B4F9E820FFD6AB16F974FE0AF8DF" box="[357,464,1772,1797]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
|
||
, male. Body length approximately
|
||
<quantity id="4CB4A7BEE820FFD6A907F974FC09F8DE" box="[884,979,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="305" unit="mm" value="7.2">7.2 mm</quantity>
|
||
. Generally black-brown (not metallic), forebody shiny, elytra slightly less shiny. Pilosity pale white, dorsal side with numerous distinct setae.
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE820FFD6AAB4F8ACFEB9F806" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="305">Clypeus anteromedially with short, erect, broad lobe; top angles slightly round; clypeogenal transition virtually continuous; genal border widely round; clypeofrontal transition with median callosity; cephalic surface with abundant, partly seta-bearing, distinct punctures. Vertex indistinctly raised. Eyes dorsally very narrow, with approximately 8 facet rows across widest point; ratio interocular distance / single maximum eye width approximately 22.</paragraph>
|
||
<caption id="DF335AD3E821FFD7AAE4F8E7FE7DF828" httpUri="https://zenodo.org/record/345757/files/figure.png" pageId="5" pageNumber="306" targetBox="[207,1396,233,1891]" targetPageId="5">
|
||
<paragraph id="8BF30A5BE821FFD7AAE4F8E7FE7DF828" blockId="5.[151,1436,1919,2034]" pageId="5" pageNumber="306">
|
||
<emphasis id="B938D649E821FFD7AAE4F8E7FEE0F84F" bold="true" box="[151,314,1919,1942]" pageId="5" pageNumber="306">FIGURES 1–8.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E821FFD7ABD2F818FDF3F84C" box="[417,553,1920,1942]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E821FFD7ABD2F818FDF3F84C" box="[417,553,1920,1942]" italics="true" pageId="5" pageNumber="306">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
species, oblique habitus. 1,
|
||
<taxonomicName id="4C4C71D8E821FFD7A937F819FC06F84C" box="[836,988,1920,1942]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E821FFD7A937F819FC06F84C" box="[836,988,1920,1942]" italics="true" pageId="5" pageNumber="306">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Toraut. 2,
|
||
<taxonomicName id="4C4C71D8E821FFD7AF68F819FA4CF84C" box="[1307,1430,1921,1942]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="scrutator">
|
||
<emphasis id="B938D649E821FFD7AF68F819FA4CF84C" box="[1307,1430,1921,1942]" italics="true" pageId="5" pageNumber="306">O. scrutator</emphasis>
|
||
</taxonomicName>
|
||
, male from Moramo. 3,
|
||
<taxonomicName id="4C4C71D8E821FFD7ABF0F838FDD7F86F" box="[387,525,1951,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="bongkudai">
|
||
<emphasis id="B938D649E821FFD7ABF0F838FDD7F86F" box="[387,525,1951,1973]" italics="true" pageId="5" pageNumber="306">O. bongkudai</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Moajat. 4,
|
||
<emphasis id="B938D649E821FFD7A926F838FC37F86F" box="[853,1005,1952,1973]" italics="true" pageId="5" pageNumber="306">
|
||
<taxonomicName id="4C4C71D8E821FFD7A926F838FC33F86F" box="[853,1001,1952,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="sangirensis">O. sangirensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
male from Toraut. 5,
|
||
<taxonomicName id="4C4C71D8E821FFD7AEB9F838FA84F86F" box="[1226,1374,1951,1973]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="holosericus">
|
||
<emphasis id="B938D649E821FFD7AEB9F838FA84F86F" box="[1226,1374,1951,1973]" italics="true" pageId="5" pageNumber="306">O. holosericus</emphasis>
|
||
</taxonomicName>
|
||
, male from Marena, 6, female from Marena. 7,
|
||
<taxonomicName id="4C4C71D8E821FFD7A835F826FD4DF80E" box="[582,663,1982,2004]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="sopu">
|
||
<emphasis id="B938D649E821FFD7A835F826FD4DF80E" box="[582,663,1982,2004]" italics="true" pageId="5" pageNumber="306">O. sopu</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from
|
||
<taxonomicName id="4C4C71D8E821FFD7A909F825FC6BF809" box="[890,945,1981,2003]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="323" phylum="Arthropoda" rank="species" species="sopu">Sopu</taxonomicName>
|
||
. 8,
|
||
<taxonomicName id="4C4C71D8E821FFD7A9ABF826FBA4F809" box="[984,1150,1981,2003]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="manguliensis">
|
||
<emphasis id="B938D649E821FFD7A9ABF826FBA4F809" box="[984,1150,1981,2003]" italics="true" pageId="5" pageNumber="306">O. manguliensis</emphasis>
|
||
</taxonomicName>
|
||
, male from Mangoli. Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF335AD3E822FFD4AAE4FABFFCF4FAA1" httpUri="https://zenodo.org/record/345758/files/figure.png" pageId="6" pageNumber="307" targetBox="[172,1423,274,1297]" targetPageId="6">
|
||
<paragraph id="8BF30A5BE822FFD4AAE4FABFFCF4FAA1" blockId="6.[151,1436,1319,1403]" pageId="6" pageNumber="307">
|
||
<emphasis id="B938D649E822FFD4AAE4FABFFE93FAE7" bold="true" box="[151,329,1319,1342]" pageId="6" pageNumber="307">FIGURES 9–12.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E822FFD4ABC7FAB0FDE7FAE4" box="[436,573,1320,1342]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E822FFD4ABC7FAB0FDE7FAE4" box="[436,573,1320,1342]" italics="true" pageId="6" pageNumber="307">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
species, oblique habitus. 9,
|
||
<taxonomicName id="4C4C71D8E822FFD4A917FAB1FBCEFAE4" box="[868,1044,1320,1342]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="begoniophilus">
|
||
<emphasis id="B938D649E822FFD4A917FAB1FBCEFAE4" box="[868,1044,1320,1342]" italics="true" pageId="6" pageNumber="307">O. begoniophilus</emphasis>
|
||
</taxonomicName>
|
||
, paratype male from Ambang. 10,
|
||
<taxonomicName id="4C4C71D8E822FFD4AFF0FAB1FE9AFA86" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="annulopunctatus">
|
||
<emphasis id="B938D649E822FFD4AFF0FAB1FE9AFA86" italics="true" pageId="6" pageNumber="307">O. annulopunctatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype male from Mogogonipa. 11,
|
||
<taxonomicName id="4C4C71D8E822FFD4A8AAFADFFC84FA86" box="[729,862,1350,1372]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="hollowayi">
|
||
<emphasis id="B938D649E822FFD4A8AAFADFFC84FA86" box="[729,862,1350,1372]" italics="true" pageId="6" pageNumber="307">O. hollowayi</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Dumoga. 12,
|
||
<taxonomicName id="4C4C71D8E822FFD4AEA3FADFFAEFFA86" box="[1232,1333,1350,1372]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="seseba">
|
||
<emphasis id="B938D649E822FFD4AEA3FADFFAEFFA86" box="[1232,1333,1350,1372]" italics="true" pageId="6" pageNumber="307">O. seseba</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Batui (pygidium up). Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF30A5BE822FFD4AAB4FA30FD6FF9AF" blockId="6.[151,1437,1447,2013]" pageId="6" pageNumber="307">Pronotum strongly convex, midline hardly depressed, impunctate; anterior declivity broad, topped by extensive, anteriorly convergent protuberance; apex very shallowly bifid (dorsal view), with declivity laterally delimited by distinct cavity on either side; anterior section of lateral border weakly sinuate, anterolateral angle approximately 80°; posterior section of lateral border slightly sinuate; base immarginate; most of pronotal derm densely, distinctly punctate-setose but discal midline as well as anterolateral cavities virtually impunctate; discal punctures separated by 1–3 punctural diameters.</paragraph>
|
||
<paragraph id="8BF30A5BE822FFD4AAB4F9E7FF3CF8DF" blockId="6.[151,1437,1447,2013]" pageId="6" pageNumber="307">
|
||
Elytral striae discally well defined, with distinct punctures, mostly separated by 2–4 times their diameter; interstriae slightly crenulate, almost flat, microreticulate (
|
||
<date id="FFF22C9BE822FFD4A94AF93CFCBDF966" box="[825,871,1700,1724]" pageId="6" pageNumber="307" value="1980-10">x80</date>
|
||
), with poorly-defined, scattered, abundant, setabearing, finely asperate punctures; punctation on lateral interstriae coarse, more rugulate. Elytral base slightly rufous.
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE822FFD4AAB4F897FED1F8B5" blockId="6.[151,1437,1447,2013]" pageId="6" pageNumber="307">Antennal club grayish brown. Metasternum with shallow midline impression; disc along midline impunctate; much of disc with sparse, fine, seta-bearing punctures; lateral wings with abundant seta-bearing, subocellate punctures.</paragraph>
|
||
<paragraph id="8BF30A5BE822FFD4AAB4F8E4FE61F807" blockId="6.[151,1437,1447,2013]" pageId="6" pageNumber="307">Abdominal ventrites each with (somewhat irregular) transverse row of closely set seta-bearing punctures; juxta-elytral surface rufous. Pygidium almost matt; crowded with seta-bearing, subocellate punctures (somewhat rugulate); base marginate.</paragraph>
|
||
<caption id="DF335AD3E823FFD5AAE4F817FBC6F839" httpUri="https://zenodo.org/record/345759/files/figure.png" pageId="7" pageNumber="308" targetBox="[162,1436,205,1903]" targetPageId="7">
|
||
<paragraph id="8BF30A5BE823FFD5AAE4F817FBC6F839" blockId="7.[151,1436,1935,2019]" pageId="7" pageNumber="308">
|
||
<emphasis id="B938D649E823FFD5AAE4F817FE8CF87F" bold="true" box="[151,342,1935,1958]" pageId="7" pageNumber="308">FIGURES 13–18.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E823FFD5ABB7F808FD96F87C" box="[452,588,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E823FFD5ABB7F808FD96F87C" box="[452,588,1936,1958]" italics="true" pageId="7" pageNumber="308">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
species, dorsal forebody. 13,
|
||
<taxonomicName id="4C4C71D8E823FFD5A9FBF809FBF9F87C" box="[904,1059,1936,1958]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E823FFD5A9FBF809FBF9F87C" box="[904,1059,1936,1958]" italics="true" pageId="7" pageNumber="308">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Toraut. 14,
|
||
<taxonomicName id="4C4C71D8E823FFD5AFF0F809FF29F81F" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="scrutator">
|
||
<emphasis id="B938D649E823FFD5AFF0F809FF29F81F" italics="true" pageId="7" pageNumber="308">O. scrutator</emphasis>
|
||
</taxonomicName>
|
||
, male from Moramo. 15,
|
||
<taxonomicName id="4C4C71D8E823FFD5A877F828FD4AF81F" box="[516,656,1967,1989]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="bongkudai">
|
||
<emphasis id="B938D649E823FFD5A877F828FD4AF81F" box="[516,656,1967,1989]" italics="true" pageId="7" pageNumber="308">O. bongkudai</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Moajat 16, paratype female. 17,
|
||
<taxonomicName id="4C4C71D8E823FFD5AEBBF828FA81F81F" box="[1224,1371,1968,1989]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="sangirensis">
|
||
<emphasis id="B938D649E823FFD5AEBBF828FA81F81F" box="[1224,1371,1968,1989]" italics="true" pageId="7" pageNumber="308">O. sangirensis</emphasis>
|
||
</taxonomicName>
|
||
, male from Toraut.18,
|
||
<taxonomicName id="4C4C71D8E823FFD5AB4FF856FE17F839" box="[316,461,1998,2019]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="sangirensis">
|
||
<emphasis id="B938D649E823FFD5AB4FF856FE17F839" box="[316,461,1998,2019]" italics="true" pageId="7" pageNumber="308">O. sangirensis</emphasis>
|
||
</taxonomicName>
|
||
, female from Tangkoko. Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF335AD3E82CFFDAAAE4F8CFFD92F810" httpUri="https://zenodo.org/record/345760/files/figure.png" pageId="8" pageNumber="309" targetBox="[171,1436,193,1838]" targetPageId="8">
|
||
<paragraph id="8BF30A5BE82CFFDAAAE4F8CFFD92F810" blockId="8.[151,1437,1879,1994]" pageId="8" pageNumber="309">
|
||
<emphasis id="B938D649E82CFFDAAAE4F8CFFE8EF8B7" bold="true" box="[151,340,1879,1902]" pageId="8" pageNumber="309">FIGURES 19–24.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E82CFFDAABCEF8C0FD9FF8B4" box="[445,581,1880,1902]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E82CFFDAABCEF8C0FD9FF8B4" box="[445,581,1880,1902]" italics="true" pageId="8" pageNumber="309">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
species, dorsal, oblique (20 only) forebody. 19,
|
||
<taxonomicName id="4C4C71D8E82CFFDAAE40F8C1FB1FF8B4" box="[1075,1221,1881,1902]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="sangirensis">
|
||
<emphasis id="B938D649E82CFFDAAE40F8C1FB1FF8B4" box="[1075,1221,1881,1902]" italics="true" pageId="8" pageNumber="309">O. sangirensis</emphasis>
|
||
</taxonomicName>
|
||
, male from Manado. 20,
|
||
<taxonomicName id="4C4C71D8E82CFFDAAAB3F8EFFE8DF856" box="[192,343,1910,1932]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="holosericus">
|
||
<emphasis id="B938D649E82CFFDAAAB3F8EFFE8DF856" box="[192,343,1910,1932]" italics="true" pageId="8" pageNumber="309">O. holosericus</emphasis>
|
||
</taxonomicName>
|
||
, female from Kendari, oblique. 21,
|
||
<taxonomicName id="4C4C71D8E82CFFDAA8A6F8EFFCB6F856" box="[725,876,1910,1932]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="holosericus">
|
||
<emphasis id="B938D649E82CFFDAA8A6F8EFFCB6F856" box="[725,876,1910,1932]" italics="true" pageId="8" pageNumber="309">O. holosericus</emphasis>
|
||
</taxonomicName>
|
||
, male from Marena. 22,
|
||
<taxonomicName id="4C4C71D8E82CFFDAAE05F8EFFAD7F856" box="[1142,1293,1910,1932]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="holosericus">
|
||
<emphasis id="B938D649E82CFFDAAE05F8EFFAD7F856" box="[1142,1293,1910,1932]" italics="true" pageId="8" pageNumber="309">O. holosericus</emphasis>
|
||
</taxonomicName>
|
||
, female from Marena. 23,
|
||
<taxonomicName id="4C4C71D8E82CFFDAAB51F80EFE0CF871" box="[290,470,1941,1963]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="begoniophilus">
|
||
<emphasis id="B938D649E82CFFDAAB51F80EFE0CF871" box="[290,470,1941,1963]" italics="true" pageId="8" pageNumber="309">O. begoniophilus</emphasis>
|
||
</taxonomicName>
|
||
, holotype male major from Ambang. 24,
|
||
<taxonomicName id="4C4C71D8E82CFFDAA9D7F80EFB82F871" box="[932,1112,1941,1963]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="begoniophilus">
|
||
<emphasis id="B938D649E82CFFDAA9D7F80EFB82F871" box="[932,1112,1941,1963]" italics="true" pageId="8" pageNumber="309">O. begoniophilus</emphasis>
|
||
</taxonomicName>
|
||
, paratype male medium from Ambang. Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF335AD3E82DFFDBAAE4F81DFAA8F803" httpUri="https://zenodo.org/record/345761/files/figure.png" pageId="9" pageNumber="310" targetBox="[170,1436,198,1875]" targetPageId="9">
|
||
<paragraph id="8BF30A5BE82DFFDBAAE4F81DFAA8F803" blockId="9.[151,1437,1925,2009]" pageId="9" pageNumber="310">
|
||
<emphasis id="B938D649E82DFFDBAAE4F81DFE82F841" bold="true" box="[151,344,1925,1948]" pageId="9" pageNumber="310">FIGURES 25–30.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E82DFFDBABBBF81EFD8AF846" box="[456,592,1926,1948]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E82DFFDBABBBF81EFD8AF846" box="[456,592,1926,1948]" italics="true" pageId="9" pageNumber="310">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
species, dorsal forebody. 25,
|
||
<taxonomicName id="4C4C71D8E82DFFDBA9E7F81FFC3DF846" box="[916,999,1927,1948]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="sopu">
|
||
<emphasis id="B938D649E82DFFDBA9E7F81FFC3DF846" box="[916,999,1927,1948]" italics="true" pageId="9" pageNumber="310">O. sopu</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from
|
||
<taxonomicName id="4C4C71D8E82DFFDBAEA0F81EFAD0F846" box="[1235,1290,1926,1948]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="323" phylum="Arthropoda" rank="species" species="sopu">Sopu</taxonomicName>
|
||
. 26,
|
||
<taxonomicName id="4C4C71D8E82DFFDBAF31F81FFA4FF846" box="[1346,1429,1927,1948]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="sopu">
|
||
<emphasis id="B938D649E82DFFDBAF31F81FFA4FF846" box="[1346,1429,1927,1948]" italics="true" pageId="9" pageNumber="310">O. sopu</emphasis>
|
||
</taxonomicName>
|
||
, paratype male from
|
||
<taxonomicName id="4C4C71D8E82DFFDBAB10F83CFE40F860" box="[355,410,1956,1978]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="323" phylum="Arthropoda" rank="species" species="sopu">Sopu</taxonomicName>
|
||
. 27,
|
||
<taxonomicName id="4C4C71D8E82DFFDBABB9F83DFDB4F860" box="[458,622,1956,1978]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="manguliensis">
|
||
<emphasis id="B938D649E82DFFDBABB9F83DFDB4F860" box="[458,622,1956,1978]" italics="true" pageId="9" pageNumber="310">O. manguliensis</emphasis>
|
||
</taxonomicName>
|
||
, male from Mangoli. 28,
|
||
<taxonomicName id="4C4C71D8E82DFFDBA900F83DFBE1F860" box="[883,1083,1956,1978]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="annulopunctatus">
|
||
<emphasis id="B938D649E82DFFDBA900F83DFBE1F860" box="[883,1083,1956,1978]" italics="true" pageId="9" pageNumber="310">O. annulopunctatus</emphasis>
|
||
</taxonomicName>
|
||
, paratype male from Mogogonipa. 29,
|
||
<taxonomicName id="4C4C71D8E82DFFDBAACEF85CFE9AF803" box="[189,320,1987,2009]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="hollowayi">
|
||
<emphasis id="B938D649E82DFFDBAACEF85CFE9AF803" box="[189,320,1987,2009]" italics="true" pageId="9" pageNumber="310">O. hollowayi</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Dumoga. 30,
|
||
<taxonomicName id="4C4C71D8E82DFFDBA8D7F85CFCDDF803" box="[676,775,1987,2009]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="seseba">
|
||
<emphasis id="B938D649E82DFFDBA8D7F85CFCDDF803" box="[676,775,1987,2009]" italics="true" pageId="9" pageNumber="310">O. seseba</emphasis>
|
||
</taxonomicName>
|
||
, holotype male from Batui. Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF335AD3E82EFFD8AAE4F912FB2FF8C1" httpUri="https://zenodo.org/record/345762/files/figure.png" pageId="10" pageNumber="311" targetBox="[151,1366,302,1606]" targetPageId="10">
|
||
<paragraph id="8BF30A5BE82EFFD8AAE4F912FB2FF8C1" blockId="10.[151,1436,1674,1820]" pageId="10" pageNumber="311">
|
||
<emphasis id="B938D649E82EFFD8AAE4F912FE8EF97A" bold="true" box="[151,340,1674,1697]" pageId="10" pageNumber="311">FIGURES 31–38.</emphasis>
|
||
Sulawesi
|
||
<taxonomicName id="4C4C71D8E82EFFD8ABCEF913FD9FF97B" box="[445,581,1675,1697]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938D649E82EFFD8ABCEF913FD9FF97B" box="[445,581,1675,1697]" italics="true" pageId="10" pageNumber="311">Onthophagus</emphasis>
|
||
</taxonomicName>
|
||
, body parts. 31–,
|
||
<taxonomicName id="4C4C71D8E82EFFD8A88DF914FC4CF97B" box="[766,918,1675,1697]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8A88DF914FC4CF97B" box="[766,918,1675,1697]" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, head full-face, holotype male from Toraut. 32,
|
||
<taxonomicName id="4C4C71D8E82EFFD8AFF0F914FECAF91A" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8AFF0F914FECAF91A" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, left elytron dorsal, holotype male from Toraut. 33,
|
||
<taxonomicName id="4C4C71D8E82EFFD8A930F932FC04F91A" box="[835,990,1706,1728]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8A930F932FC04F91A" box="[835,990,1706,1728]" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, protibial upper side, holotype male from Toraut. 34,
|
||
<taxonomicName id="4C4C71D8E82EFFD8AB66F951FE6BF904" box="[277,433,1736,1758]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8AB66F951FE6BF904" box="[277,433,1736,1758]" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, metatibial underside, paratype male from Mogogonipa. 35,
|
||
<taxonomicName id="4C4C71D8E82EFFD8AE33F951FB06F904" box="[1088,1244,1736,1758]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8AE33F951FB06F904" box="[1088,1244,1736,1758]" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, aedeagus lateral, paratype male from Toraut. 36,
|
||
<taxonomicName id="4C4C71D8E82EFFD8ABACF970FDA3F927" box="[479,633,1767,1789]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="bisscrutator">
|
||
<emphasis id="B938D649E82EFFD8ABACF970FDA3F927" box="[479,633,1767,1789]" italics="true" pageId="10" pageNumber="311">O. bisscrutator</emphasis>
|
||
</taxonomicName>
|
||
, head full-face, paratype female from Toraut. 37,
|
||
<taxonomicName id="4C4C71D8E82EFFD8AE0DF970FB20F927" box="[1150,1274,1768,1789]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="scrutator">
|
||
<emphasis id="B938D649E82EFFD8AE0DF970FB20F927" box="[1150,1274,1768,1789]" italics="true" pageId="10" pageNumber="311">O. scrutator</emphasis>
|
||
</taxonomicName>
|
||
, head full-face, female from Matanyo. 38,
|
||
<taxonomicName id="4C4C71D8E82EFFD8ABD5F89EFDFBF8C6" box="[422,545,1798,1820]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="scrutator">
|
||
<emphasis id="B938D649E82EFFD8ABD5F89EFDFBF8C6" box="[422,545,1798,1820]" italics="true" pageId="10" pageNumber="311">O. scrutator</emphasis>
|
||
</taxonomicName>
|
||
, aedeagus lateral, male from Morano. Scale lines approximately 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF30A5BE82EFFD8AAB4F8D1FDB7F82B" blockId="10.[151,1436,1864,2033]" pageId="10" pageNumber="311">
|
||
Legs brown. Protibia slender, distally slightly curved inwards, apex internally slightly produced (angular), with 3+(1) larger denticles (tip of some worn off); spur short, acuminate. Mesotibiae and metatibiae and their spurs unmodified (some worn off). Profemoral, mesofemoral, and metafemoral underside sparsely, finely punctatesetose. Approximate proportions metatarsomeres 1–5:
|
||
<date id="FFF22C9BE82EFFD8A976F82DFC93F817" box="[773,841,1973,1997]" pageId="10" pageNumber="311" value="2011-03-04">11/4/3</date>
|
||
/2/4 (full spur in male
|
||
<typeStatus id="54F7B4F9E82EFFD8AE3CF82DFB6AF817" box="[1103,1200,1973,1997]" pageId="10" pageNumber="311" type="paratype">paratype</typeStatus>
|
||
slightly shorter than tarsomere 1: proportion approximately 9).
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE82FFFD9AAB4FF0FFDCEFF6A" blockId="11.[151,1437,151,536]" box="[199,532,151,176]" pageId="11" pageNumber="312">
|
||
Aedeagus,
|
||
<figureCitation id="137716DEE82FFFD9AB30FF0FFE48FF6A" box="[323,402,151,176]" captionStart="FIGURES 31 – 38" captionStartId="10.[151,264,1674,1697]" captionTargetBox="[151,1366,302,1606]" captionTargetId="figure@10.[151,1436,193,1653]" captionTargetPageId="10" captionText="FIGURES 31 – 38. Sulawesi Onthophagus, body parts. 31 –, O. bisscrutator, head full-face, holotype male from Toraut. 32, O. bisscrutator, left elytron dorsal, holotype male from Toraut. 33, O. bisscrutator, protibial upper side, holotype male from Toraut. 34, O. bisscrutator, metatibial underside, paratype male from Mogogonipa. 35, O. bisscrutator, aedeagus lateral, paratype male from Toraut. 36, O. bisscrutator, head full-face, paratype female from Toraut. 37, O. scrutator, head full-face, female from Matanyo. 38, O. scrutator, aedeagus lateral, male from Morano. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345762/files/figure.png" pageId="11" pageNumber="312">Fig. 35</figureCitation>
|
||
(
|
||
<typeStatus id="54F7B4F9E82FFFD9ABD2FF00FDD2FF6A" box="[417,520,152,176]" pageId="11" pageNumber="312" type="paratype">paratype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE82FFFD9AAB4FF25FC76FF22" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="312">Measurements in mm. Maximum width of head 2.1. Median length of pronotum (apart from protrusion) 2.3, maximum width 3.8. Sutural length of elytra 2.6, maximum width 4.0.</paragraph>
|
||
<paragraph id="8BF30A5BE82FFFD9AAB4FE9CFBE7FE9A" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="312">
|
||
<emphasis id="B938D649E82FFFD9AAB4FE9CFDB9FEC7" bold="true" box="[199,611,260,285]" pageId="11" pageNumber="312">Variation and sexual dimorphism.</emphasis>
|
||
Body length 5.0–
|
||
<quantity id="4CB4A7BEE82FFFD9A940FE9CFC57FEC6" box="[819,909,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="11" pageNumber="312" unit="mm" value="7.5">7.5 mm</quantity>
|
||
. Clypeal apex and pronotal gibbosity variably developed. Minor males may lack anteromedian clypeal lobe, looking like females.
|
||
</paragraph>
|
||
<paragraph id="8BF30A5BE82FFFD9AAB4FED4FEC8FE0A" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="312">
|
||
<emphasis id="B938D649E82FFFD9AAB4FED4FEC6FEBE" box="[199,284,332,356]" italics="true" pageId="11" pageNumber="312">Female</emphasis>
|
||
lacking anteromedian clypeal lobe. Clypeus rugulate-punctate. Clypeofrontal ridge present, weaklypronounced, curvilinear. Vertex with low, transverse costa. Pronotum more-or-less swollen anteromedially, otherwise unmodified. Legs different, protibia more robust, not curved, apex not produced; spur elongateacuminate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35659D0E82FFFD9AAB4FE44FF0BFDC2" pageId="11" pageNumber="312" type="etymology">
|
||
<paragraph id="8BF30A5BE82FFFD9AAB4FE44FF0BFDC2" blockId="11.[151,1437,151,536]" pageId="11" pageNumber="312">
|
||
<emphasis id="B938D649E82FFFD9AAB4FE44FE94FE2F" bold="true" box="[199,334,476,501]" pageId="11" pageNumber="312">Etymology.</emphasis>
|
||
The “bis-“ prefix has been added to the name of the related species,
|
||
<taxonomicName id="4C4C71D8E82FFFD9AE2EFE45FB19FE2E" box="[1117,1219,477,500]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="scrutator">
|
||
<emphasis id="B938D649E82FFFD9AE2EFE45FB19FE2E" box="[1117,1219,477,500]" italics="true" pageId="11" pageNumber="312">scrutator</emphasis>
|
||
</taxonomicName>
|
||
, meaning a second time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |