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<document id="29ABCAA0751AE7CADCC026B5B3F47287" ID-DOI="10.5852/ejt.2016.190" ID-GBIF-Dataset="3c438ae3-9ea0-4eda-b893-acf6d5756c90" ID-ISSN="2118-9773" ID-Zenodo-Dep="831004" ID-ZooBank="urn:lsid:zoobank.org:pub:BE92596B-62D9-46CD-8486-CF6B36C640B1 1" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1500288028508" checkinUser="jeremy" docAuthor="Berhard A. Huber, Booppa Petchard, Charles Leh Moi Ung, Joseph K. H. Koh &amp; Amir R. M. Ghazali" docDate="2016" docId="7F7187D54E457021FD91C310C8D0FAB7" docLanguage="en" docName="Huber, B. A., Petcharad, B., Ung, C. L. M., Koh, J. K., &amp; Ghazali, A. R. (2016). The Southeast Asian Pholcus halabala species group (Done).imf" docOrigin="European Journal of Taxonomy 190 (190)" docSource="http://www.europeanjournaloftaxonomy.eu/index.php/ejt/article/view/313" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pholcus ubin Huber, sp. nov." docType="treatment" docUuid="5EC32BD5-FE22-45DE-A5EB-7659D61FFC03" docUuidSource="ZooBank" docVersion="14" lastPageNumber="12" masterDocId="8348FFAD4E4C702AFFFCC07ECC17FFF8" masterDocTitle="The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae): new data from field observations and ultrastructure" masterLastPageNumber="55" masterPageNumber="1" pageNumber="10" updateTime="1733942487093" updateUser="ExternalLinkService">
<mods:mods id="327CB8F156D57B983EDC724F27BA79C6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="862F62280B9ED7A15C8D26005D0C48AD">The Southeast Asian Pholcus halabala species group (Araneae, Pholcidae): new data from field observations and ultrastructure</mods:title>
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<mods:namePart id="DDFFBCCEB38C2F7E6F19AE5A9B065792">Berhard A. Huber</mods:namePart>
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<mods:namePart id="FC78CE77CA340D8B58E705764A1B74EB">Booppa Petchard</mods:namePart>
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<mods:namePart id="30E6136157B014DDEEC76E943B1626AA">Charles Leh Moi Ung</mods:namePart>
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<mods:namePart id="28A7C68F05F5506779C7D155D280B708">Joseph K. H. Koh</mods:namePart>
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<mods:namePart id="3650EAA8F50BABA606FE2EC1A6794650">Amir R. M. Ghazali</mods:namePart>
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<mods:title id="75D76BDE344CAC576453BF4411902DB6">European Journal of Taxonomy</mods:title>
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<mods:date id="046EFB57F2567EE064A7754852039F8B">2016</mods:date>
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<mods:number id="4DC2DAC9609AED2A4A4839B43A9F3326">2016-04-14</mods:number>
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<mods:number id="D1DBE05AB8FAD73B3C4E3C21C4C9EAAD">190</mods:number>
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<treatment id="7F7187D54E457021FD91C310C8D0FAB7" ID-DOI="http://doi.org/10.5281/zenodo.6076772" ID-GBIF-Taxon="132549846" ID-Zenodo-Dep="6076772" LSID="urn:lsid:zoobank.org:act:5EC32BD5-FE22-45DE-A5EB-7659D61FFC03" httpUri="http://treatment.plazi.org/id/7F7187D54E457021FD91C310C8D0FAB7" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" scope_family="Pholcidae" scope_order="Araneae">
<subSubSection id="BFC265484E457023FD91C310CFD0FC71" box="[621,967,878,905]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="F76736C34E457023FD91C310CFD0FC71" blockId="9.[361,1227,878,976]" box="[621,967,878,905]" pageId="9" pageNumber="10">
<heading id="AC2F81AF4E457023FD91C310CFD0FC71" box="[621,967,878,905]" centered="true" fontSize="11" level="3" pageId="9" pageNumber="10" reason="4">
<taxonomicName id="30D84D404E457023FD91C310CF77FC71" ID-CoL="9473P" authority="Huber" authorityName="Huber" box="[621,864,878,905]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ubin" status="sp. nov.">
<emphasis id="C5ACEAD14E457023FD91C310CF1AFC71" bold="true" box="[621,781,878,905]" italics="true" pageId="9" pageNumber="10">Pholcus ubin</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="DE9F57AA4E457023FC90C311CFD0FC71" box="[876,967,879,905]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E457023FE95C3ECCFB2FC28" pageId="9" pageNumber="10" type="description">
<paragraph id="F76736C34E457023FE95C3ECC8DCFC55" blockId="9.[361,1227,878,976]" box="[361,1227,914,941]" pageId="9" pageNumber="10">urn:lsid:zoobank.org:act:5EC32BD5-FE22-45DE-A5EB-7659D61FFC03</paragraph>
<paragraph id="F76736C34E457023FD73C3C8CFB2FC28" blockId="9.[361,1227,878,976]" box="[655,933,950,976]" pageId="9" pageNumber="10">
<figureCitation id="6FE32A464E457023FD73C3C8CEE0FC28" box="[655,759,950,976]" captionStart="Figs 1 – 8" captionStartId="3.[189,241,1895,1921]" captionTargetBox="[198,1389,620,1852]" captionTargetId="figure@3.[198,1389,620,1853]" captionTargetPageId="3" captionText="Figs 1 – 8. Live specimens, Pholcus halabala Huber, 2011 (1 – 6) and Ph. ubin Huber, sp. nov. (7 – 8). 1. ♂ from MacRitchie, Singapore. 2. ♂ from Gunung Jerai, Malaysia. 3. ♀ with egg-sac from Gunung Liang, Malaysia. 4. Juvenile in web with silk tufts, Upper Selatar, Singapore. 5. ♀ with egg-sac from Hala Bala, Thailand. 6. ♀ with egg-sac from Ulu Dong, Malaysia. 7 – 8. ♂ and ♀ from Pulau Ubin, Singapore." figureDoi="http://doi.org/10.5281/zenodo.1183673" httpUri="https://zenodo.org/record/1183673/files/figure.png" pageId="9" pageNumber="10">Figs 7–8</figureCitation>
,
<figureCitation id="6FE32A464E457023FCFFC3C8CF58FC28" box="[771,847,950,976]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="9" pageNumber="10">29–33</figureCitation>
,
<figureCitation id="6FE32A464E457023FCA7C3C8CFB2FC28" box="[859,933,950,976]" captionStart="Figs 90 – 101" captionStartId="25.[189,241,1930,1956]" captionTargetBox="[189,1399,430,1890]" captionTargetId="figure@25.[189,1399,430,1890]" captionTargetPageId="25" captionText="Figs 90 – 101. Female genitalia, untreated in ventral view, cleared in ventral and dorsal views. 90 – 92. Pholcus ubin Huber, sp. nov. 93 – 95. Ph. lintang Huber, sp. nov. 96 – 98. Ph. kuhapimuk Huber, sp. nov. 99 – 101. Ph. khaolek Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.1183699" httpUri="https://zenodo.org/record/1183699/files/figure.png" pageId="9" pageNumber="10">90–92</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E457023FF41C389C959FB71" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="F76736C34E457023FF41C389CD21FBE9" blockId="9.[189,1398,1015,1161]" box="[189,310,1015,1041]" pageId="9" pageNumber="10">
<heading id="AC2F81AF4E457023FF41C389CD21FBE9" bold="true" box="[189,310,1015,1041]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="6">
<emphasis id="C5ACEAD14E457023FF41C389CD21FBE9" bold="true" box="[189,310,1015,1041]" pageId="9" pageNumber="10">Diagnosis</emphasis>
</heading>
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<paragraph id="F76736C34E457023FF41C458C959FB71" blockId="9.[189,1398,1015,1161]" pageId="9" pageNumber="10">
Easily distinguished from putatively closest known relatives (other species in the
<taxonomicName id="30D84D404E457023FB82C459C8F1FBB9" box="[1150,1254,1063,1089]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="halabala">
<emphasis id="C5ACEAD14E457023FB82C459C8F1FBB9" box="[1150,1254,1063,1089]" italics="true" pageId="9" pageNumber="10">halabala</emphasis>
</taxonomicName>
core group) by morphology of male palps (
<figureCitation id="6FE32A464E457023FDD3C434CEAFFB9C" box="[559,696,1098,1124]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="9" pageNumber="10">Figs 29–30</figureCitation>
; procursus with heavily sclerotized dorsal process distally; shapes of uncus and appendix) and by female internal genitalia (
<figureCitation id="6FE32A464E457023FC53C410C820FB70" box="[943,1079,1134,1160]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="9" pageNumber="10">Figs 32–33</figureCitation>
; triangular pore plates).
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E457023FF41C4CECFE9FB00" box="[189,1022,1200,1273]" pageId="9" pageNumber="10" type="etymology">
<paragraph id="F76736C34E457023FF41C4CECD53FB32" blockId="9.[189,1022,1200,1273]" box="[189,324,1200,1226]" pageId="9" pageNumber="10">
<heading id="AC2F81AF4E457023FF41C4CECD53FB32" bold="true" box="[189,324,1200,1226]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="6">
<emphasis id="C5ACEAD14E457023FF41C4CECD53FB32" bold="true" box="[189,324,1200,1226]" pageId="9" pageNumber="10">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E457023FF41C4A0CFE9FB00" blockId="9.[189,1022,1200,1273]" box="[189,1022,1246,1273]" pageId="9" pageNumber="10">
The species name is derived from the
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locality; noun in apposition.
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<typeStatus id="286388614E457023FF41C55ECCEDFAC2" box="[189,250,1312,1338]" pageId="9" pageNumber="10">Type</typeStatus>
material
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</heading>
</paragraph>
<paragraph id="F76736C34E457023FF41C531C917FA75" blockId="9.[189,1399,1359,1421]" pageId="9" pageNumber="10">
<materialsCitation id="47B03C9E4E457023FF41C531C8EBFA75" ID-GBIF-Occurrence="1575903807" collectedFrom="on leaves" collectingDate="2015-02-16" collectionCode="ZFMK" collectorName="B. A. Huber &amp; J. Koh &amp; D. Court" country="Singapore" elevation="20" latitude="1.4033333" location="degraded forest near park headquarters" longLatPrecision="130" longitude="103.97" pageId="9" pageNumber="10" specimenCode="Ar 15010" specimenCount="1" specimenCount-male="1" stateProvince="Pulau Ubin" typeStatus="holotype">
<collectingCountry id="8FCF76534E457023FF41C531CD71FA91" box="[189,358,1359,1385]" name="Singapore" pageId="9" pageNumber="10" value="Singapore">SINGAPORE</collectingCountry>
:
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,
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,
<collectionCode id="91C9AE064E457023FDE8C52ECE72FA92" box="[532,613,1360,1386]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" pageId="9" pageNumber="10">ZFMK</collectionCode>
(
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),
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Pulau
<taxonomicName id="30D84D404E457023FCAAC531CF83FA91" box="[854,916,1359,1385]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="26" phylum="Arthropoda" rank="species" species="ubin">Ubin</taxonomicName>
</collectingRegion>
,
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(
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,
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),
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</quantity>
a.s.l.,
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,
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<collectingDate id="9322E9EB4E457023FD10C50DCF91FA75" box="[748,902,1394,1421]" pageId="9" pageNumber="10" value="2015-02-16">16 Feb. 2015</collectingDate>
</date>
(
<collectorName id="5A2D53154E457023FC6BC50DC836FA75" box="[919,1057,1395,1421]" pageId="9" pageNumber="10">B.A. Huber</collectorName>
,
<collectorName id="5A2D53154E457023FBD0C50DC86CFA75" box="[1068,1147,1395,1421]" pageId="9" pageNumber="10">J. Koh</collectorName>
,
<collectorName id="5A2D53154E457023FB74C50DC8E4FA75" box="[1160,1267,1395,1421]" pageId="9" pageNumber="10">D. Court</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F76736C34E457023FF41C5CACDEEFA36" blockId="9.[189,1399,1460,1569]" box="[189,505,1460,1486]" pageId="9" pageNumber="10">
<heading id="AC2F81AF4E457023FF41C5CACDEEFA36" bold="true" box="[189,505,1460,1486]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="6">
<emphasis id="C5ACEAD14E457023FF41C5CACDEEFA36" bold="true" box="[189,505,1460,1486]" pageId="9" pageNumber="10">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E457023FF41C59DCE6BF9D9" blockId="9.[189,1399,1460,1569]" pageId="9" pageNumber="10">
<materialsCitation id="47B03C9E4E457023FF41C59DC83AFA05" ID-GBIF-Occurrence="1575903825" box="[189,1069,1507,1534]" collectedFrom="on leaves" collectingDate="2015-02-16" collectionCode="ZFMK" collectorName="B. A. Huber &amp; J. Koh &amp; D. Court" country="Singapore" elevation="20" latitude="1.4033333" location="SINGAPORE" longLatPrecision="130" longitude="103.97" pageId="9" pageNumber="10" specimenCode="Ar 15011" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Pulau Ubin">
<collectingCountry id="8FCF76534E457023FF41C59DCD71FA05" box="[189,358,1507,1533]" name="Singapore" pageId="9" pageNumber="10" value="Singapore">SINGAPORE</collectingCountry>
:
<specimenCount id="E1DEFD4A4E457023FE89C59DCDAAFA06" box="[373,445,1507,1534]" count="2" pageId="9" pageNumber="10" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="E1DEFD4A4E457023FE31C59DCE03FA06" box="[461,532,1507,1534]" count="3" pageId="9" pageNumber="10" type="female">3 ♀♀</specimenCount>
,
<collectionCode id="91C9AE064E457023FDDAC59ACE60FA06" box="[550,631,1508,1534]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" pageId="9" pageNumber="10">ZFMK</collectionCode>
(
<specimenCode id="A77E9EB84E457023FD77C59DCF14FA06" box="[651,771,1507,1534]" pageId="9" pageNumber="10">Ar 15011</specimenCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="47B03C9E4E457023FBC1C59DCE6FF9D9" ID-GBIF-Occurrence="1575903808" collectedFrom="on leaves" collectingDate="2015-02-16" collectionCode="ZFMK" collectorName="B. A. Huber &amp; J. Koh &amp; D. Court" country="Singapore" elevation="20" latitude="1.4033333" location="SINGAPORE" longLatPrecision="130" longitude="103.97" pageId="9" pageNumber="10" specimenCode="3 juvs, Mal 232" specimenCount="2" specimenCount-female="2" specimenCount-juvenile="3" stateProvince="Pulau Ubin">
<specimenCount id="E1DEFD4A4E457023FBC1C59DC893FA06" box="[1085,1156,1507,1534]" count="2" pageId="9" pageNumber="10" type="female">2 ♀♀</specimenCount>
,
<specimenCode id="A77E9EB84E457023FB69C59DC8F4FA05" box="[1173,1251,1507,1533]" pageId="9" pageNumber="10">3 juvs</specimenCode>
, in absolute ethanol,
<collectionCode id="91C9AE064E457023FEDEC679CD64F9D9" box="[290,371,1543,1569]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" pageId="9" pageNumber="10">ZFMK</collectionCode>
(
<specimenCode id="A77E9EB84E457023FE7FC679CDFCF9D9" box="[387,491,1543,1569]" pageId="9" pageNumber="10">Mal 232</specimenCode>
), same data
</materialsCitation>
.
</paragraph>
<paragraph id="F76736C34E457023FF41C637CD5EF97F" blockId="9.[189,1399,1609,1671]" pageId="9" pageNumber="10">
<materialsCitation id="47B03C9E4E457023FF41C637CD52F97F" ID-GBIF-Occurrence="1575903810" collectingDate="2009-08-22" collectionCode="RMNH" collectorName="A. Floren" country="MALAYSIA-BORNEO" latitude="6.015" location="Gaya Island" longLatPrecision="13" longitude="116.0195" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Sabah">
<collectingCountry id="8FCF76534E457023FF41C637CDCBF99B" box="[189,476,1609,1635]" pageId="9" pageNumber="10" value="Malaysia-borneo">MALAYSIA-BORNEO</collectingCountry>
:
<specimenCount id="E1DEFD4A4E457023FE14C637CE00F99B" box="[488,535,1609,1635]" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
,
<collectionCode id="91C9AE064E457023FDD9C637CE97F99B" box="[549,640,1609,1635]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="9" pageNumber="10">RMNH</collectionCode>
,
<taxonomicName id="30D84D404E457023FD70C637CEC0F99B" box="[652,727,1609,1635]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="12" phylum="Arthropoda" rank="species" species="sabah">
<collectingRegion id="351CF8214E457023FD70C637CEC0F99B" box="[652,727,1609,1635]" pageId="9" pageNumber="10">Sabah</collectingRegion>
</taxonomicName>
,
<location id="F20760184E457023FD18C637CF65F99B" LSID="urn:lsid:plazi:treatment:7F7187D54E457021FD91C310C8D0FAB7:F20760184E457023FD18C637CF65F99B" box="[740,882,1609,1635]" country="MALAYSIA-BORNEO" latitude="6.015" longLatPrecision="13" longitude="116.0195" name="Gaya Island" pageId="9" pageNumber="10" stateProvince="Sabah">Gaya Island</location>
(
<geoCoordinate id="92EC50044E457023FC78C637C819F99B" box="[900,1038,1609,1635]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="9" value="6.015">6° 00.90' N</geoCoordinate>
,
<geoCoordinate id="92EC50044E457023FBE7C637C8A8F99B" box="[1051,1215,1609,1635]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="9" value="116.0195">116° 01.17' E</geoCoordinate>
),
<date id="836610034E457023FB29C637C960F99B" box="[1237,1399,1609,1635]" pageId="9" pageNumber="10" value="2009-08-22">
<collectingDate id="9322E9EB4E457023FB29C637C960F99B" box="[1237,1399,1609,1635]" pageId="9" pageNumber="10" value="2009-08-22">22 Aug. 2009</collectingDate>
</date>
(
<collectorName id="5A2D53154E457023FF3AC612CD2AF97F" box="[198,317,1644,1671]" pageId="9" pageNumber="10">A. Floren</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E457021FF41C6D0CEF7FBE3" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
<paragraph id="F76736C34E457023FF41C6D0CD59F930" blockId="9.[189,1399,1710,1891]" box="[189,334,1710,1736]" pageId="9" pageNumber="10">
<heading id="AC2F81AF4E457023FF41C6D0CD59F930" bold="true" box="[189,334,1710,1736]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="6">
<emphasis id="C5ACEAD14E457023FF41C6D0CD59F930" bold="true" box="[189,334,1710,1736]" pageId="9" pageNumber="10">Description</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E457023FF16C6A3CDBCF900" blockId="9.[189,1399,1710,1891]" box="[234,427,1757,1784]" pageId="9" pageNumber="10">
<emphasis id="C5ACEAD14E457023FF16C6A3CD3DF90F" bold="true" box="[234,298,1757,1783]" pageId="9" pageNumber="10">Male</emphasis>
(
<typeStatus id="286388614E457023FEC5C6A3CDB4F900" box="[313,419,1757,1784]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="F76736C34E457023FF41C77FC863F89A" blockId="9.[189,1399,1710,1891]" pageId="9" pageNumber="10">MEASUREMENTS. Total body length 4.1, carapace width 1.0. Leg 1: 30.4 (7.3 + 0.4 + 7.2 + 14.2 + 1.3), tibia 2: 4.7, tibia 3: 2.7, tibia 4: 3.9; tibia 1 L/d: 86. Distance PME-PME 220 µm, diameter PME 95 µm, distance PME-ALE ~35 µm; distance AME-AME 35 µm; diameter AME 60 µm.</paragraph>
<paragraph id="F76736C34E457020FF41C7F4CDFEFEB0" blockId="9.[189,1399,1929,2027]" lastBlockId="10.[189,1398,267,328]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
COLOR. Carapace ochre-yellow with brown pattern of radiating marks posteriorly, in live specimens with reddish color in median area between posterior marks (
<figureCitation id="6FE32A464E457023FCACC7D3CF8DF830" box="[848,922,1965,1992]" captionStart="Figs 1 – 8" captionStartId="3.[189,241,1895,1921]" captionTargetBox="[198,1389,620,1852]" captionTargetId="figure@3.[198,1389,620,1853]" captionTargetPageId="3" captionText="Figs 1 – 8. Live specimens, Pholcus halabala Huber, 2011 (1 – 6) and Ph. ubin Huber, sp. nov. (7 – 8). 1. ♂ from MacRitchie, Singapore. 2. ♂ from Gunung Jerai, Malaysia. 3. ♀ with egg-sac from Gunung Liang, Malaysia. 4. Juvenile in web with silk tufts, Upper Selatar, Singapore. 5. ♀ with egg-sac from Hala Bala, Thailand. 6. ♀ with egg-sac from Ulu Dong, Malaysia. 7 – 8. ♂ and ♀ from Pulau Ubin, Singapore." figureDoi="http://doi.org/10.5281/zenodo.1183673" httpUri="https://zenodo.org/record/1183673/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
); ocular area and clypeus not darkened; sternum with some dark marks posteriorly; legs ochre-yellow with dark brown patellae and tibiametatarsus joints; abdomen pale ochre-gray with small black and white marks dorsally and laterally, monochromous ventrally.
</paragraph>
<paragraph id="F76736C34E467020FF41C10EC962FE55" blockId="10.[189,1398,368,430]" pageId="10" pageNumber="11">
BODY. Habitus as in
<figureCitation id="6FE32A464E467020FE5CC10ECDF3FE72" box="[416,484,368,394]" captionStart="Figs 1 – 8" captionStartId="3.[189,241,1895,1921]" captionTargetBox="[198,1389,620,1852]" captionTargetId="figure@3.[198,1389,620,1853]" captionTargetPageId="3" captionText="Figs 1 – 8. Live specimens, Pholcus halabala Huber, 2011 (1 – 6) and Ph. ubin Huber, sp. nov. (7 – 8). 1. ♂ from MacRitchie, Singapore. 2. ♂ from Gunung Jerai, Malaysia. 3. ♀ with egg-sac from Gunung Liang, Malaysia. 4. Juvenile in web with silk tufts, Upper Selatar, Singapore. 5. ♀ with egg-sac from Hala Bala, Thailand. 6. ♀ with egg-sac from Ulu Dong, Malaysia. 7 – 8. ♂ and ♀ from Pulau Ubin, Singapore." figureDoi="http://doi.org/10.5281/zenodo.1183673" httpUri="https://zenodo.org/record/1183673/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
; ocular area slightly raised, with brushes of ~10 stronger hairs behind each PME; carapace without median furrow; clypeus unmodified; sternum wider than long (0.62/0.40), unmodified.
</paragraph>
<caption id="A3A7664B4E467020FF41C73ACE6BF813" ID-DOI="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="10" pageNumber="11" targetBox="[203,1386,477,1813]" targetPageId="10">
<paragraph id="F76736C34E467020FF41C73ACE6BF813" blockId="10.[189,1399,1860,2027]" pageId="10" pageNumber="11">
<emphasis id="C5ACEAD14E467020FF41C73ACD5AF8A6" bold="true" box="[189,333,1860,1886]" pageId="10" pageNumber="11">Figs 29–33.</emphasis>
<taxonomicName id="30D84D404E467020FEA4C73ACE5AF8A6" authority="Huber" authorityName="Huber" box="[344,589,1860,1886]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ubin" status="sp. nov.">
<emphasis id="C5ACEAD14E467020FEA4C73ACDE1F8A6" box="[344,502,1860,1886]" italics="true" pageId="10" pageNumber="11">Pholcus ubin</emphasis>
Huber
</taxonomicName>
,
<taxonomicNameLabel id="DE9F57AA4E467020FDA1C73ACEA9F8A6" box="[605,702,1860,1886]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, ZFMK Ar 15011.
<emphasis id="C5ACEAD14E467020FC52C73ACFEFF8A6" bold="true" box="[942,1016,1860,1886]" pageId="10" pageNumber="11">29–30</emphasis>
. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus).
<emphasis id="C5ACEAD14E467020FE93C7F4CD9BF85C" bold="true" box="[367,396,1930,1956]" pageId="10" pageNumber="11">31</emphasis>
. Male chelicerae, frontal view.
<emphasis id="C5ACEAD14E467020FCF1C7F4CF40F85C" bold="true" box="[781,855,1930,1956]" pageId="10" pageNumber="11">32–33</emphasis>
. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29–30, 32–33).
</paragraph>
</caption>
<paragraph id="F76736C34E477021FF41C175CDFAFEB3" blockId="11.[189,1398,267,331]" pageId="11" pageNumber="12">
CHELICERAE. As in
<figureCitation id="6FE32A464E477021FE60C175CDE3FEDD" box="[412,500,267,293]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Fig. 31</figureCitation>
, with pair of frontal apophyses provided with two modified hairs each and rounded lateral processes.
</paragraph>
<paragraph id="F76736C34E477021FF41C10BCFDAFDDC" blockId="11.[189,1398,372,548]" pageId="11" pageNumber="12">
PALPS. As in
<figureCitation id="6FE32A464E477021FEA7C10ACDF3FE77" box="[347,484,372,399]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Figs 29–30</figureCitation>
; coxa unmodified; trochanter with retrolatero-ventral apophysis; femur with small retrolateral apophysis proximally; tibia very large; tarsus with dorsal elongation carrying subdistal tarsal organ; procursus with prominent ventral ‘knee’, with distinctive heavily sclerotized dorsal process distally (arrow in
<figureCitation id="6FE32A464E477021FE73C19BCDF1FE07" box="[399,486,485,511]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Fig. 30</figureCitation>
); bulb oval, with distinctive uncus and appendix; weakly sclerotized embolus with subdistal fringed side branch (hidden by appendix in
<figureCitation id="6FE32A464E477021FC95C274CFA8FDDC" box="[873,959,522,548]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="F76736C34E477021FF41C230CE54FD4C" blockId="11.[189,1398,590,692]" pageId="11" pageNumber="12">LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 5%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 with ~20 pseudosegments (only distally ~15 fairly distinct).</paragraph>
<paragraph id="F76736C34E477021FF16C2A3CDBAFD0F" blockId="11.[189,578,732,796]" box="[234,429,732,759]" pageId="11" pageNumber="12">
<emphasis id="C5ACEAD14E477021FF16C2A3CD3DFD0F" bold="true" box="[234,298,733,759]" pageId="11" pageNumber="12">Male</emphasis>
(variation)
</paragraph>
<paragraph id="F76736C34E477021FF41C37CCE55FCE4" blockId="11.[189,578,732,796]" box="[189,578,770,796]" pageId="11" pageNumber="12">
Tibia
<quantity id="30209B264E477021FEFDC37CCD39FCE4" box="[257,302,770,796]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
2 other males: 6.1, 7.3.
</paragraph>
<paragraph id="F76736C34E477021FF16C338CD52FC98" blockId="11.[189,1399,838,1051]" box="[234,325,838,864]" pageId="11" pageNumber="12">
<heading id="AC2F81AF4E477021FF16C338CD52FC98" bold="true" box="[234,325,838,864]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="6">
<emphasis id="C5ACEAD14E477021FF16C338CD52FC98" bold="true" box="[234,325,838,864]" pageId="11" pageNumber="12">Female</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E477021FF41C312CEF7FBE3" blockId="11.[189,1399,838,1051]" pageId="11" pageNumber="12">
In general similar to male (
<figureCitation id="6FE32A464E477021FDFEC315CE5CFC7D" box="[514,587,875,901]" captionStart="Figs 1 – 8" captionStartId="3.[189,241,1895,1921]" captionTargetBox="[198,1389,620,1852]" captionTargetId="figure@3.[198,1389,620,1853]" captionTargetPageId="3" captionText="Figs 1 – 8. Live specimens, Pholcus halabala Huber, 2011 (1 – 6) and Ph. ubin Huber, sp. nov. (7 – 8). 1. ♂ from MacRitchie, Singapore. 2. ♂ from Gunung Jerai, Malaysia. 3. ♀ with egg-sac from Gunung Liang, Malaysia. 4. Juvenile in web with silk tufts, Upper Selatar, Singapore. 5. ♀ with egg-sac from Hala Bala, Thailand. 6. ♀ with egg-sac from Ulu Dong, Malaysia. 7 – 8. ♂ and ♀ from Pulau Ubin, Singapore." figureDoi="http://doi.org/10.5281/zenodo.1183673" httpUri="https://zenodo.org/record/1183673/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
), but without stronger hairs behind PME; sternum mostly dark with some small light marks; eye triads closer together than in male (PME-PME distance: 185 µm). Tibia
<quantity id="30209B264E477021FA94C3EFCCC3FC28" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="12" unit="in" value="1.0">1 in</quantity>
<specimenCount id="E1DEFD4A4E477021FF20C3C8CD43FC28" box="[220,340,950,976]" pageId="11" pageNumber="12" type="female">3 females</specimenCount>
: 5.8, 5.9, 6.1. Epigynum weakly sclerotized whitish plate (
<figureCitation id="6FE32A464E477021FBF3C3C8C87FFC28" box="[1039,1128,950,976]" captionStart="Figs 90 – 101" captionStartId="25.[189,241,1930,1956]" captionTargetBox="[189,1399,430,1890]" captionTargetId="figure@25.[189,1399,430,1890]" captionTargetPageId="25" captionText="Figs 90 – 101. Female genitalia, untreated in ventral view, cleared in ventral and dorsal views. 90 – 92. Pholcus ubin Huber, sp. nov. 93 – 95. Ph. lintang Huber, sp. nov. 96 – 98. Ph. kuhapimuk Huber, sp. nov. 99 – 101. Ph. khaolek Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.1183699" httpUri="https://zenodo.org/record/1183699/files/figure.png" pageId="11" pageNumber="12">Fig. 90</figureCitation>
), anterior internal arch visible through cuticle, posterior margin laterally slightly more sclerotized; with median ‘knob’ (
<figureCitation id="6FE32A464E477021FAE4C3A5C965FC0E" box="[1304,1394,987,1014]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Figs 32</figureCitation>
,
<figureCitation id="6FE32A464E477021FF41C47FCD1CFBE3" box="[189,267,1025,1051]" captionStart="Figs 90 – 101" captionStartId="25.[189,241,1930,1956]" captionTargetBox="[189,1399,430,1890]" captionTargetId="figure@25.[189,1399,430,1890]" captionTargetPageId="25" captionText="Figs 90 – 101. Female genitalia, untreated in ventral view, cleared in ventral and dorsal views. 90 – 92. Pholcus ubin Huber, sp. nov. 93 – 95. Ph. lintang Huber, sp. nov. 96 – 98. Ph. kuhapimuk Huber, sp. nov. 99 – 101. Ph. khaolek Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.1183699" httpUri="https://zenodo.org/record/1183699/files/figure.png" pageId="11" pageNumber="12">90–91</figureCitation>
); internal genitalia as in
<figureCitation id="6FE32A464E477021FDD7C47FCE95FBE3" box="[555,642,1025,1051]" captionStart="Figs 29 – 33" captionStartId="10.[189,241,1860,1886]" captionTargetBox="[203,1386,477,1813]" captionTargetId="figure@10.[197,1390,477,1818]" captionTargetPageId="10" captionText="Figs 29 – 33. Pholcus ubin Huber, sp. nov., ZFMK Ar 15011. 29 – 30. Left male palp, prolateral and retrolateral views (arrow points at distinctive dorsal process; asterisk marks dorsal elongation of male palpal tarsus). 31. Male chelicerae, frontal view. 32 – 33. Cleared female genitalia, ventral and dorsal views. Abbreviations: a = appendix; b = genital bulb; e = embolus; p = procursus; u = uncus. Scale lines: 0.3 mm (31); 0.5 mm (29 – 30, 32 – 33)." figureDoi="http://doi.org/10.5281/zenodo.1183681" httpUri="https://zenodo.org/record/1183681/files/figure.png" pageId="11" pageNumber="12">Figs 33</figureCitation>
and
<figureCitation id="6FE32A464E477021FD47C47FCECBFBE3" box="[699,732,1025,1051]" captionStart="Figs 90 – 101" captionStartId="25.[189,241,1930,1956]" captionTargetBox="[189,1399,430,1890]" captionTargetId="figure@25.[189,1399,430,1890]" captionTargetPageId="25" captionText="Figs 90 – 101. Female genitalia, untreated in ventral view, cleared in ventral and dorsal views. 90 – 92. Pholcus ubin Huber, sp. nov. 93 – 95. Ph. lintang Huber, sp. nov. 96 – 98. Ph. kuhapimuk Huber, sp. nov. 99 – 101. Ph. khaolek Huber, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.1183699" httpUri="https://zenodo.org/record/1183699/files/figure.png" pageId="11" pageNumber="12">92</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E477021FF41C43BC8F7FB22" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="F76736C34E477021FF41C43BCD69FBA6" blockId="11.[189,382,1092,1119]" box="[189,382,1092,1119]" pageId="11" pageNumber="12">
<heading id="AC2F81AF4E477021FF41C43BCD69FBA6" bold="true" box="[189,382,1092,1119]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="C5ACEAD14E477021FF41C43BCD69FBA6" bold="true" box="[189,382,1092,1119]" pageId="11" pageNumber="12">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E477021FF41C40BC8F7FB22" blockId="11.[189,1399,1141,1243]" pageId="11" pageNumber="12">
All Pulau
<taxonomicName id="30D84D404E477021FECEC40BCD7AFB77" box="[306,365,1141,1167]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="26" phylum="Arthropoda" rank="species" species="ubin">Ubin</taxonomicName>
specimens were found in a highly degraded patch of forest near park headquarters, while no specimen was found in a well preserved forest about
<quantity id="30209B264E477021FCA8C4E5CFB1FB4D" box="[852,934,1179,1205]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="12" unit="km" value="3.3">3.3 km</quantity>
WNW. Most specimens were collected by beating of branches, but some were observed in their flat resting position on the leaves.
</paragraph>
</subSubSection>
<subSubSection id="BFC265484E477021FF41C57AC8D0FAB7" box="[189,1223,1284,1360]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="F76736C34E477021FF41C57ACD41FAE6" blockId="11.[189,342,1284,1310]" box="[189,342,1284,1310]" pageId="11" pageNumber="12">
<heading id="AC2F81AF4E477021FF41C57ACD41FAE6" bold="true" box="[189,342,1284,1310]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="C5ACEAD14E477021FF41C57ACD41FAE6" bold="true" box="[189,342,1284,1310]" pageId="11" pageNumber="12">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="F76736C34E477021FF41C54BC8D0FAB7" blockId="11.[189,1223,1333,1360]" box="[189,1223,1333,1360]" pageId="11" pageNumber="12">
Known from two localities in Singapore and Gaya Island (
<taxonomicName id="30D84D404E477021FC96C54BCFA2FAA8" box="[874,949,1333,1360]" class="Arachnida" family="Pholcidae" genus="Pholcus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
) respectively (
<figureCitation id="6FE32A464E477021FB9EC54BC8AEFAB7" box="[1122,1209,1333,1359]" captionStart="Fig. 17" captionStartId="5.[189,232,2000,2026]" captionTargetBox="[189,1398,895,1959]" captionTargetId="figure@5.[189,1399,895,1959]" captionTargetPageId="5" captionText="Fig. 17. Known distribution of the core group of the Pholcus halabala species group." figureDoi="http://doi.org/10.5281/zenodo.1183677" httpUri="https://zenodo.org/record/1183677/files/figure.png" pageId="11" pageNumber="12">Fig. 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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