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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="9592D168D36A5E28BEF50A7A35B381F3" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Paraphaea philippinensis Shi &amp; Zhou &amp; Liang 2013, comb. n." docType="treatment" docVersion="3" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="20" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="19" updateTime="1668155586682" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
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<mods:start>1</mods:start>
<mods:end>129</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
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<treatment ID-GBIF-Taxon="182235607" LSID="urn:lsid:plazi:treatment:9592D168D36A5E28BEF50A7A35B381F3" httpUri="http://treatment.plazi.org/id/9592D168D36A5E28BEF50A7A35B381F3" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection pageId="18" pageNumber="19" type="nomenclature">
<paragraph pageId="18" pageNumber="19">
<taxonomicName LSID="9592D168-D36A-5E28-BEF5-0A7A35B381F3" authority="(Jedlicka, 1935 b)" authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paraphaea philippinensis" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="philippinensis" status="comb. n.">
<pageBreakToken pageId="18" pageNumber="19" start="start">Paraphaea</pageBreakToken>
philippinensis (
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, 1935b)
</taxonomicName>
<taxonomicNameLabel pageId="18" pageNumber="19">comb. n.</taxonomicNameLabel>
<figureCitation captionStart="Figures 3136" captionStartId="F6" captionText="Figures 31 - 36. Type materials, scale bars = 2.0 mm: 31 Lectotype of Plochionus binotatus Dejean [= Paraphaea binotata (Dejean)] 32 Lectotype of Calleida discophora Chaudoir [= Paraphaea binotata (Dejean)] 33 Lectotype of Paraphaea signifera Bates [= Paraphaea binotata (Dejean)] 34 Holotype of Parena formosana Jedlicka [= Paraphaea formosana (Jedlicka)] 35 Holotype of Paraphaea minor sp. n. 36 Holotype of Coptodera philippinensis Jedlicka [= Paraphaea philippinensis (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29679" pageId="18" pageNumber="19">Habitus: Fig. 36</figureCitation>
<figureCitation captionStart="Figures 111125" captionStartId="F41" captionText="Figures 111 - 125. Left ovipositors of females, ventral view, scale bars = 0.2 mm: 111 Paraphaea binotata (Dejean), a specimen from Guangxi 112 Paraphaea minor sp. n., a paratype from Hainan 113 Paraphaea philippinensis (Jedlicka), holotype 114 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 115 Physodera dejeani Eschscholtz, a specimen from Yunnan 116 Physodera eschscholtzii Parry, a specimen from Yunnan 117 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 118 Allocota aurata (Bates), a specimen from Shaanxi 119 Allocota bicolor sp. n., a paratype from Guangdong 120 Lachnoderma asperum Bates, a specimen from Zhejiang 121 Dasiosoma bellum (Habu), a specimen from Guangdong 122 Dasiosoma maindroni (Tian &amp; Deuve), a specimen from Tonkin 123 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 124 Orionella lewisii (Bates), a specimen from Zhejiang 125 Endynomena pradieri (Fairmaire), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29719" pageId="18" pageNumber="19">female genitalia: Figs 113</figureCitation>
<figureCitation captionStart="Figures 126131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="18" pageNumber="19">128</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="reference_group">
<paragraph pageId="18" pageNumber="19">
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="philippinensis" status="comb. n.">Paraphaea philippinensis</taxonomicName>
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Sbornik Entomologickeho Oddeleni pri Zoologichych Sbirkach Narodniho Musea v Praze" pageId="87" pageNumber="88" refId="B55" refString="Jedlicka, A, 1935b. Novy Druh Rodu Coptodera (Carab.). Eine Neue Coptodera-Art (Carab.). Casopis Ceskoslovenske spolecnosti entomologicke 32: 116." title="Novy Druh Rodu Coptodera (Carab.). Eine Neue Coptodera-Art (Carab.). Casopis Ceskoslovenske spolecnosti entomologicke 32: 116." year="1935 b">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1935b
</bibRefCitation>
: 116 (original:
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Coptodera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="18" pageNumber="19">Coptodera</emphasis>
</taxonomicName>
; type locality: Philippinen; holotype deposited in NHML);
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 306 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="18" pageNumber="19">Allocota</emphasis>
</taxonomicName>
);
<bibRefCitation author="Shpeley, D" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="88" pageNumber="89" pagination="1 - 182" refId="B82" refString="Shpeley, D, Ball, GE, 1993. Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini). Proceedings of the Entomological Society of Ontario 124: 1 - 182" title="Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini)." volume="124" year="1993">Shpeley and Ball 1993</bibRefCitation>
: 129 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="18" pageNumber="19">Anchista</emphasis>
</taxonomicName>
);
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="757 - 799" refId="B62" refString="Kirschenhofer, E, 1996. Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil. Linzer Biologische Beitraege 28 (2): 757 - 799" title="Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil." volume="28" year="1996">Kirschenhofer 1996</bibRefCitation>
: 763-764 (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus" status="comb. n.">
<emphasis italics="true" pageId="18" pageNumber="19">Allocota</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="type examined">
<paragraph pageId="18" pageNumber="19">Type examined.</paragraph>
<paragraph pageId="18" pageNumber="19">
Holotype of
<taxonomicName authorityName="Jedlicka" authorityYear="1935" class="Hexapoda" family="Carabidae" genus="Coptodera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis italics="true" pageId="18" pageNumber="19">Coptodera philippinensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
, by monotypy (NHML): female, body length = 7.6 mm, board mounted, with antennae missing except scapes,
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[round label with red ringed]; &quot;Philippine Is. / Coll. Bottcher. / B. M. 1929-201.&quot;; &quot;
<emphasis italics="true" pageId="18" pageNumber="19">217</emphasis>
&quot;; &quot;
<emphasis italics="true" pageId="18" pageNumber="19">Coptodera</emphasis>
/
<emphasis italics="true" pageId="18" pageNumber="19">philippinen- / type sis sp. n</emphasis>
. / Det. ING.
<normalizedToken originalValue="JEDLIČKA”">JEDLICKA&quot;</normalizedToken>
[pink label]; &quot;loan from / Brit. Mus.&quot;; &quot;
<emphasis italics="true" pageId="18" pageNumber="19">Coptodera / philippi- /</emphasis>
<emphasis italics="true" pageId="18" pageNumber="19">ensis</emphasis>
/ det.
<emphasis italics="true" pageId="18" pageNumber="19">Jedl/ 97</emphasis>
/ George E. Ball&quot;; &quot;
<emphasis italics="true" pageId="18" pageNumber="19">Anchista</emphasis>
/
<emphasis italics="true" pageId="18" pageNumber="19">philippinensis</emphasis>
/ det.
<emphasis italics="true" pageId="18" pageNumber="19">Jedlicka</emphasis>
/ D. Shpeley 19
<emphasis italics="true" pageId="18" pageNumber="19">93</emphasis>
&quot;[Figs 36, 113, 128].
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="notes on type">
<paragraph pageId="18" pageNumber="19">Notes on type.</paragraph>
<paragraph pageId="18" pageNumber="19">
<bibRefCitation author="Shpeley, D" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="88" pageNumber="89" pagination="1 - 182" refId="B82" refString="Shpeley, D, Ball, GE, 1993. Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini). Proceedings of the Entomological Society of Ontario 124: 1 - 182" title="Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini)." volume="124" year="1993">Shpeley and Ball (1993)</bibRefCitation>
mentioned the holotype is a male, but actually it is a female (
<figureCitation captionStart="Figures 3136" captionStartId="F6" captionText="Figures 31 - 36. Type materials, scale bars = 2.0 mm: 31 Lectotype of Plochionus binotatus Dejean [= Paraphaea binotata (Dejean)] 32 Lectotype of Calleida discophora Chaudoir [= Paraphaea binotata (Dejean)] 33 Lectotype of Paraphaea signifera Bates [= Paraphaea binotata (Dejean)] 34 Holotype of Parena formosana Jedlicka [= Paraphaea formosana (Jedlicka)] 35 Holotype of Paraphaea minor sp. n. 36 Holotype of Coptodera philippinensis Jedlicka [= Paraphaea philippinensis (Jedlicka)]." httpUri="https://binary.pensoft.net/fig/29679" pageId="18" pageNumber="19">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="19" type="diagnosis">
<paragraph pageId="18" pageNumber="19">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="19">Pronotum sparsely pubescent, widest at apical two-fifths, lateral margins slightly sinuate before hind angles; head and pronotum yellowish to reddish brown, elytra metallic blue; elytra with two or three primary setigerous pores on 3rd interval and one pore on base of 5th interval, intervals with a few very fine secondary setigerous pores.</paragraph>
<paragraph pageId="18" pageNumber="19">By the metallic blue elytra and sparse secondary setae on pronotum and elytra, this species can be readily distinguished from other allied species.</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph pageId="18" pageNumber="19">Description.</paragraph>
<paragraph lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Body length 7.6 mm; head reddish brown, pronotum, mouthparts and legs yellowish brown, apices of mandibles darker; elytra metallic blue with 1st and 2nd intervals slightly reddish, lateral margins, elytral suture and epipleurae reddish brown; ventral side brownish.
<emphasis bold="true" pageId="18" pageNumber="19">Head</emphasis>
nearly glabrous (only two very fine accessory setae visible), without punctures or microsculpture.
<emphasis bold="true" pageId="18" pageNumber="19">Pronotum</emphasis>
cordiform, widest at apical two-fifths; ratio PW/PL 1.52; pronotal base weakly lobed; disc sparsely and finely setose, lateral explanate areas and area along median line glabrous, microsculpture indistinct, without punctures; front angles with a few setae, distinctly longer than in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Dejean" baseAuthorityYear="1825" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="binotata">
<emphasis italics="true" pageId="18" pageNumber="19">Paraphaea binotata</emphasis>
</taxonomicName>
; lateral margins completely rounded in the widest part, slightly sinuate before hind angles; front angles widened and rounded, not projected; hind angles obtuse; disc moderately convex; basal foveae wide and even, with a few punctures.
<emphasis bold="true" pageId="18" pageNumber="19">Elytra</emphasis>
with striae very shallow, almost completely composed of rows of fine punctures; intervals flat, without punctures or microsculpture, very fine and sparse accessory setae present on all intervals; 3rd interval with two or three primary setigerous pores (in the holotype, two pores on left elytron, three pores on right one), first one placed at basal one-fourth, adjacent to 3rd stria, the last one placed at apical one-eighth, adjacent to 2nd stria, the middle one, if present, just behind the basal one, at basal one-third approximately; 5th interval with one pore near base, adjacent to 5th stria; 17-18 umbilical pores in 9th interval.
<emphasis bold="true" pageId="18" pageNumber="19">Male genitalia</emphasis>
unknown.
<emphasis bold="true" pageId="18" pageNumber="19">Female genitalia</emphasis>
. Spermatheca not distinctly bent in middle; spermathecal gland inserted near basal one-third of spermatheca, its
<pageBreakToken pageId="19" pageNumber="20" start="start">base</pageBreakToken>
slightly expanded, with a very short branch; apical part of spermatheca after the gland insertion with distinct ring-sculpture, slightly expanded to fusiform; basal part before the gland insertion without sculpture, slightly bent (
<figureCitation captionStart="Figures 126131" captionStartId="F42" captionText="Figures 126 - 131. Internal reproductive system of females, scale bars = 0.2 mm: 126 Paraphaea binotata (Dejean), a specimen from Guangxi 127 Paraphaea minor sp. n., a paratype from Hainan 128 Paraphaea philippinensis (Jedlicka), holotype 129 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 130 Physodera dejeani Eschscholtz, a specimen from Yunnan 131 Physodera eschscholtzii Parry, a specimen from Hainan." httpUri="https://binary.pensoft.net/fig/29720" pageId="19" pageNumber="20">Fig. 128</figureCitation>
). Apical segment of ovipositor scimitar-form, inner margin slightly angulate at apical fourth; length 4 times basal width; inner margin setose in apical fourth; apex narrow, with membranous extension long and slender (
<figureCitation captionStart="Figures 111125" captionStartId="F41" captionText="Figures 111 - 125. Left ovipositors of females, ventral view, scale bars = 0.2 mm: 111 Paraphaea binotata (Dejean), a specimen from Guangxi 112 Paraphaea minor sp. n., a paratype from Hainan 113 Paraphaea philippinensis (Jedlicka), holotype 114 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 115 Physodera dejeani Eschscholtz, a specimen from Yunnan 116 Physodera eschscholtzii Parry, a specimen from Yunnan 117 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 118 Allocota aurata (Bates), a specimen from Shaanxi 119 Allocota bicolor sp. n., a paratype from Guangdong 120 Lachnoderma asperum Bates, a specimen from Zhejiang 121 Dasiosoma bellum (Habu), a specimen from Guangdong 122 Dasiosoma maindroni (Tian &amp; Deuve), a specimen from Tonkin 123 Dasiosoma quadraticolle sp. n., a paratype from Yunnan 124 Orionella lewisii (Bates), a specimen from Zhejiang 125 Endynomena pradieri (Fairmaire), a specimen from Taiwan." httpUri="https://binary.pensoft.net/fig/29719" pageId="19" pageNumber="20">Fig. 113</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="distribution">
<paragraph pageId="19" pageNumber="20">Distribution</paragraph>
<paragraph pageId="19" pageNumber="20">
(
<figureCitation captionStart="Map 2" captionStartId="F46" captionText="Map 2. Known distribution of genus Paraphaea II: ▲ Paraphaea minor sp. n. ◆ Paraphaea philippinensis (Jedlicka) ★ Paraphaea formosana (Jedlicka)." httpUri="https://binary.pensoft.net/fig/29724" pageId="19" pageNumber="20">Map 2</figureCitation>
). Only known from the type locality, the Philippines.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="remarks">
<paragraph pageId="19" pageNumber="20">Remarks.</paragraph>
<paragraph pageId="19" pageNumber="20">
This species is only known from the holotype. It was originally described in genus
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Hexapoda" family="Carabidae" genus="Coptodera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Coptodera</emphasis>
</taxonomicName>
and later moved to
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Allocota</emphasis>
</taxonomicName>
by
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Sbornik Entomologickeho Oddeleni pri Zoologichych Sbirkach Narodniho Musea v Praze" pageId="87" pageNumber="88" refId="B55" refString="Jedlicka, A, 1935b. Novy Druh Rodu Coptodera (Carab.). Eine Neue Coptodera-Art (Carab.). Casopis Ceskoslovenske spolecnosti entomologicke 32: 116." title="Novy Druh Rodu Coptodera (Carab.). Eine Neue Coptodera-Art (Carab.). Casopis Ceskoslovenske spolecnosti entomologicke 32: 116." year="1935 b">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
(1935b
</bibRefCitation>
,
<bibRefCitation author="Jedlicka, A" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="87" pageNumber="88" pagination="269 - 579" refId="B58" refString="Jedlicka, A, 1963. Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae). Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 269 - 579" title="Monographie der Truncatipennen aus Ostasien, Lebiinae- Odacanthinae- Braehyninae (Coleptera, Carabidae)." volume="28" year="1963">1963</bibRefCitation>
).
<bibRefCitation author="Shpeley, D" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="88" pageNumber="89" pagination="1 - 182" refId="B82" refString="Shpeley, D, Ball, GE, 1993. Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini). Proceedings of the Entomological Society of Ontario 124: 1 - 182" title="Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini)." volume="124" year="1993">Shpeley and Ball (1993)</bibRefCitation>
indicated that it is an
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Anchista</emphasis>
</taxonomicName>
species.
<bibRefCitation author="Kirschenhofer, E" journalOrPublisher="Linzer Biologische Beitraege" pageId="87" pageNumber="88" pagination="757 - 799" refId="B62" refString="Kirschenhofer, E, 1996. Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil. Linzer Biologische Beitraege 28 (2): 757 - 799" title="Weitere neue und wenig bekannte Carabidae aus Sued- und Suedostasien (Coleoptera: Carabidae: Lebiinae: Colliurinae, Callistinae, Hexagoninae, Brachininae, Panagaeinae). II. Teil." volume="28" year="1996">Kirschenhofer (1996)</bibRefCitation>
overlooked
<bibRefCitation author="Shpeley, D" journalOrPublisher="Proceedings of the Entomological Society of Ontario" pageId="88" pageNumber="89" pagination="1 - 182" refId="B82" refString="Shpeley, D, Ball, GE, 1993. Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini). Proceedings of the Entomological Society of Ontario 124: 1 - 182" title="Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini)." volume="124" year="1993">
Shpeley and
<normalizedToken originalValue="Balls">Ball's</normalizedToken>
(1993)
</bibRefCitation>
comments, and kept it in
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" baseAuthorityName="Shi &amp; Zhou &amp; Liang" baseAuthorityYear="2013" class="Insecta" family="Carabidae" genus="Allocota" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Allocota</emphasis>
</taxonomicName>
. Based on the pronotal shape and female genitalia characters, this species is most similar to
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="minor">
<emphasis italics="true" pageId="19" pageNumber="20">Paraphaea minor</emphasis>
</taxonomicName>
sp. n., so we herein move it into
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="19" pageNumber="20">Paraphaea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>