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<document id="2D30BC814B3083E977804E80E191AD9D" ID-DOI="10.11646/phytotaxa.381.1.9" ID-ISSN="1179-3163" ID-Zenodo-Dep="13726112" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725675168103" checkinUser="felipe" docAuthor="Sang, Julia, Kiew, Ruth &amp; Yiing, Ling Chea" docDate="2018" docId="03CBE12E707B793F95B5A072C9FAE5BC" docLanguage="en" docName="phytotaxa.381.1.9.pdf" docOrigin="Phytotaxa 381 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.381.1.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia jokwaniana C. Y. Ling &amp; S. Julia. A. Habit 2018, spec. nov." docType="treatment" docVersion="2" lastPageNumber="67" masterDocId="FFF29956707C7936953DA56DCD78E70D" masterDocTitle="The Begonia flora of Gunung Mulu and Gunung Buda National Parks, Sarawak, Borneo, including five new species" masterLastPageNumber="79" masterPageNumber="58" pageNumber="65" updateTime="1725688374631" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="42A0A3C56181468AEBA8CEF7FE06F2D2">The Begonia flora of Gunung Mulu and Gunung Buda National Parks, Sarawak, Borneo, including five new species</mods:title>
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<mods:namePart id="380A88F1167011A9131F56C296C00F9A">Sang, Julia</mods:namePart>
<mods:nameIdentifier id="2C73AA2D46492860D8CADEB73DA03F18" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
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<mods:namePart id="CFF5FFFE1381C64684A8070C688CE6F1">Kiew, Ruth</mods:namePart>
<mods:nameIdentifier id="956F02FA21EE13089EA1DADC2FE3EB0E" type="email">ruth@frim.gov.my</mods:nameIdentifier>
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<mods:namePart id="353A97209D4D8C973FFAB426604CC5A9">Yiing, Ling Chea</mods:namePart>
<mods:nameIdentifier id="036E94775E7CC7DCBAE7774DF890A537" type="email">juliasang@sarawakforestry.com</mods:nameIdentifier>
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<mods:title id="FBCF22C3986A81622F3C7715CBF4865F">Phytotaxa</mods:title>
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<mods:date id="26A9D47458CED3F75CEB351350571371">2018</mods:date>
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<mods:number id="37C9E849D5295F208EE0E96C868750D7">2018-12-07</mods:number>
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<treatment id="03CBE12E707B793F95B5A072C9FAE5BC" LSID="urn:lsid:plazi:treatment:03CBE12E707B793F95B5A072C9FAE5BC" httpUri="http://treatment.plazi.org/id/03CBE12E707B793F95B5A072C9FAE5BC" lastPageId="9" lastPageNumber="67" pageId="7" pageNumber="65">
<subSubSection id="C37803B3707B793195B5A072CE0BE235" box="[136,883,1309,1336]" pageId="7" pageNumber="65" type="nomenclature">
<paragraph id="8BDD5038707B793195B5A072CE0BE235" blockId="7.[136,1452,1309,1587]" box="[136,883,1309,1336]" pageId="7" pageNumber="65">
<heading id="D095E754707B793195B5A072CE0BE235" box="[136,883,1309,1336]" fontSize="11" level="2" pageId="7" pageNumber="65" reason="4">
<emphasis id="B9168C2A707B793195B5A072CFFCE235" bold="true" box="[136,644,1309,1336]" pageId="7" pageNumber="65">
8.
<taxonomicName id="4C622BBB707B79319599A073CF06E235" ID-CoL="7SG5P" authority="C.Y.Ling &amp; S.Julia" authorityName="C. Y. Ling &amp; S. Julia. A. Habit" authorityYear="2018" box="[164,638,1309,1336]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="65" phylum="Tracheophyta" rank="species" species="jokwaniana" status="sp. nov.">
<emphasis id="B9168C2A707B79319599A073CCEBE235" bold="true" box="[164,403,1309,1336]" italics="true" pageId="7" pageNumber="65">Begonia jokwaniana</emphasis>
C.Y.Ling &amp; S.Julia
</taxonomicName>
,
</emphasis>
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<emphasis id="B9168C2A707B793197B6A073CF8DE235" bold="true" box="[651,757,1310,1336]" italics="true" pageId="7" pageNumber="65">spec. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="13594CBD707B79319637A072CE12E235" box="[778,874,1311,1336]" captionStart="FIGURE 5" captionStartId="8.[136,229,1929,1950]" captionTargetBox="[160,1426,200,1894]" captionTargetId="figure-14@8.[151,1436,190,1904]" captionTargetPageId="8" captionText="FIGURE 5. Begonia jokwaniana C.Y.Ling &amp; S.Julia. A. Habit; B. Stipule; C. Upper surface of leaf; D. Male inflorescence; E &amp; F. Male flower; G. Female flower; H. Fruit; I. Transverse section of fruit. All photos from SFC 6914 by C.Y. Ling." figureDoi="http://doi.org/10.5281/zenodo.13726124" httpUri="https://zenodo.org/record/13726124/files/figure.png" pageId="7" pageNumber="65">Figure 5</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37803B3707B793F95B5A02BC948E69C" lastPageId="9" lastPageNumber="67" pageId="7" pageNumber="65" type="description">
<paragraph id="8BDD5038707B793195B5A02BC95EE13E" blockId="7.[136,1452,1309,1587]" pageId="7" pageNumber="65">
In its habit and broadly ovate leaves, it is similar to
<taxonomicName id="4C622BBB707B79319741A02BCE8BE256" authority="Ridley (1906: 249)" authorityName="Ridley" authorityPageNumber="249" authorityYear="1906" box="[636,1011,1350,1371]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="65" phylum="Tracheophyta" rank="species" species="propinqua">
<emphasis id="B9168C2A707B79319741A02BCE4DE256" box="[636,821,1350,1371]" italics="true" pageId="7" pageNumber="65">Begonia propinqua</emphasis>
<bibRefCitation id="EFF32DC9707B79319601A02BCE8BE256" author="Ridley, H. N." box="[828,1011,1350,1371]" pageId="7" pageNumber="65" pagination="1 - 92" refId="ref16573" refString="Ridley, H. N. (1906) Begonias of Borneo. Journal of Straits Branch of the Royal Asiatic Society 43: 1 - 92." type="journal article" year="1906">Ridley (1906: 249)</bibRefCitation>
</taxonomicName>
but it is different in its smaller, ovate stipules 12–15 ×
<quantity id="4C9AFDDD707B79319428A007CC17E272" box="[277,367,1386,1407]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="7" pageNumber="65" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
(vs. broadly lanceolate stipule, 18–30 ×
<quantity id="4C9AFDDD707B793197C0A007CE29E272" box="[765,849,1386,1407]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="7" pageNumber="65" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
), more numerous veins, 5–8 veins on either side of the midrib and 2–3 veins in the basal lobes (vs. 3–5 veins on either side of the midrib and 4–5 veins in the basal lobes), longer peduncle,
<quantity id="4C9AFDDD707B79319048A0E3CDA2E2CA" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="7" pageNumber="65" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 cm</quantity>
long (vs. peduncle
<quantity id="4C9AFDDD707B793194A1A0DFCC8EE2CA" box="[412,502,1458,1479]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="7" pageNumber="65" unit="cm" value="5.75" valueMax="7.0" valueMin="4.5">4.5–7 cm</quantity>
long), wider bracts ca.
<quantity id="4C9AFDDD707B793197E0A0DFCE59E2CA" box="[733,801,1458,1479]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="65" unit="mm" value="12.0">12 mm</quantity>
wide (vs. ca.
<quantity id="4C9AFDDD707B79319696A0DFCE9BE2CA" box="[939,995,1458,1479]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="65" unit="mm" value="6.0">6 mm</quantity>
wide), male flower with longer pedicels
<quantity id="4C9AFDDD707B79319041A0DFCD9AE2E6" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="7" pageNumber="65" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long (vs.
<quantity id="4C9AFDDD707B79319478A0BBCCEDE2E6" box="[325,405,1494,1515]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="65" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
long), female flowers with larger ovate tepals 12–13 ×
<quantity id="4C9AFDDD707B7931968FA0BBC976E2E6" box="[946,1038,1494,1515]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="7" pageNumber="65" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
(vs. elliptic, 10–12 ×
<quantity id="4C9AFDDD707B793191D8A0BBC859E2E6" box="[1253,1313,1494,1515]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="65" unit="mm" value="5.0">5 mm</quantity>
), fruits single (vs. in pairs) and capsules shorter and broader 6–7 ×
<quantity id="4C9AFDDD707B7931978BA097CE63E102" box="[694,795,1530,1551]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="7" pageNumber="65" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
(vs. ca. 9 ×
<quantity id="4C9AFDDD707B793196ADA097CEB2E102" box="[912,970,1530,1551]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="65" unit="mm" value="6.0">6 mm</quantity>
).
<emphasis id="B9168C2A707B793196E7A097C96CE102" bold="true" box="[986,1044,1530,1551]" pageId="7" pageNumber="65">Type:</emphasis>
—
<collectingCountry id="F37510A8707B79319110A097C9D4E102" box="[1069,1196,1530,1551]" name="Malaysia" pageId="7" pageNumber="65">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="49A69EDA707B7931903BA097C824E102" box="[1286,1372,1530,1551]" country="Malaysia" name="Sarawak" pageId="7" pageNumber="65">Sarawak</collectingRegion>
. Marudi District: Gunung Mulu NP, Camp 4,
<date id="FFDC76F8707B79319722A373CFDEE13E" box="[543,678,1566,1587]" pageId="7" pageNumber="65" value="2016-04-18">18 April 2016</date>
,
<emphasis id="B9168C2A707B7931978DA373CE01E13E" box="[688,889,1566,1587]" italics="true" pageId="7" pageNumber="65">Ling et al. SFC 6914</emphasis>
(
<typeStatus id="54D9EE9A707B793196B8A373CEA3E13E" box="[901,987,1566,1587]" pageId="7" pageNumber="65" type="holotype">holotype</typeStatus>
SAR!).
</paragraph>
<paragraph id="8BDD5038707B793F95B5A30EC948E69C" blockId="7.[136,1452,1635,2092]" lastBlockId="9.[136,1452,160,689]" lastPageId="9" lastPageNumber="67" pageId="7" pageNumber="65">
Cane-like begonia to
<quantity id="4C9AFDDD707B793194BEA30ECCBEE171" box="[387,454,1635,1660]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="65" unit="m" value="1.5">1.5 m</quantity>
tall. Stem red brown, succulent,
<quantity id="4C9AFDDD707B79319671A30ECED4E171" box="[844,940,1635,1660]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="65" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
thick, pubescent, little-branched; internodes
<quantity id="4C9AFDDD707B793195B5A3EACC7DE1AD" box="[136,261,1671,1696]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.5" metricValueMin="3.5" pageId="7" pageNumber="65" unit="cm" value="5.5" valueMax="7.5" valueMin="3.5">3.5–7.5 cm</quantity>
long, slightly thicker at nodes. Stipule pale green, ovate, 12–15 ×
<quantity id="4C9AFDDD707B793196D5A3EAC92BE1AD" box="[1000,1107,1671,1696]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="7" pageNumber="65" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
, slightly keeled, margin entire, apex setose, seta to
<quantity id="4C9AFDDD707B79319459A3C6CCDCE1C9" box="[356,420,1707,1732]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="65" unit="mm" value="2.0">2 mm</quantity>
long, caducous. Leaves alternate, distant, held horizontally. Petiole red-brown, (2.5–)
<quantity id="4C9AFDDD707B79319063A3C6CDD2E1E5" metricMagnitude="-2" metricUnit="m" metricValue="9.25" metricValueMax="12.5" metricValueMin="6.0" pageId="7" pageNumber="65" unit="cm" value="9.25" valueMax="12.5" valueMin="6.0">6–12.5 cm</quantity>
long, grooved above. Lamina ovate, oblique, plain green above, pale green beneath, sparsely bristly above, hairs pale green on a raised base, glossy and fleshy in life, 11–16.5 ×
<quantity id="4C9AFDDD707B79319679A39ECEDFE001" box="[836,935,1779,1804]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="7" pageNumber="65" unit="cm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 cm</quantity>
, broad side
<quantity id="4C9AFDDD707B79319114A39EC9EBE001" box="[1065,1171,1779,1804]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="9.5" metricValueMin="6.0" pageId="7" pageNumber="65" unit="cm" value="7.75" valueMax="9.5" valueMin="6.0">6–9.5 cm</quantity>
, base cordate, sometimes overlapping, basal lobes
<quantity id="4C9AFDDD707B7931949AA27ACC84E03D" box="[423,508,1815,1840]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="7" pageNumber="65" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
wide, margin irregularly dentate; venation palmate-pinnate, reddish near the base above, otherwise pale green, 5–8 veins on either side of midrib, 2–3 veins in the basal lobes, prominent on both surfaces. Inflorescence protogynous, in the upper leaf axil, paniculate,
<quantity id="4C9AFDDD707B793196A2A232C975E075" box="[927,1037,1887,1912]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="3.3" metricValueMin="1.2" pageId="7" pageNumber="65" unit="cm" value="22.5" valueMax="33.0" valueMin="12.0">12–33 cm</quantity>
long, erect, peduncle
<quantity id="4C9AFDDD707B79319039A232C814E075" box="[1284,1388,1887,1912]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="7" pageNumber="65" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 cm</quantity>
long, reddish, pubescent, 1–
<specimenCount id="9D649BB1707B793194BCA2EECC9AE091" box="[385,482,1923,1948]" count="3" pageId="7" pageNumber="65" type="female">3 female</specimenCount>
flowers at the base, above many male flowers; bracts pale green, ca. 10 ×
<quantity id="4C9AFDDD707B79319026A2EEC814E091" box="[1307,1388,1923,1948]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="65" unit="mm" value="12.0">12 mm</quantity>
, apex acute, glabrous, margin entire, caducous; bracteoles up to 4 pairs, ovate, 8–10 ×
<quantity id="4C9AFDDD707B79319135A2CAC9FBE0CD" box="[1032,1155,1959,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="7" pageNumber="65" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
, pale green, margin entire, apex acute to rounded, caducous. Male flower: pedicel white,
<quantity id="4C9AFDDD707B7931967BA2A6CEC9E0E9" box="[838,945,1995,2020]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="7" pageNumber="65" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long, glabrous, tepals 4, outer 2 tepals white inside, pale green outside, broadly elliptic, 7–8 ×
<quantity id="4C9AFDDD707B79319790A282CE73EF05" box="[685,779,2031,2056]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="7" pageNumber="65" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
, margin entire, apex broadly acute to rounded; inner 2 tepals white or whitish green, lanceolate, 5–6 ×
<quantity id="4C9AFDDD707B7931975FAD7ECFA0EF21" box="[610,728,2067,2092]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="7" pageNumber="65" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
, margin entire, apex bluntly acute; stamens 16–20, pale yellow, cluster globose, subsessile; filaments pale yellow, ca.
<quantity id="4C9AFDDD7075793F97DFA5CDCE40E7B4" box="[738,824,160,185]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="67" unit="mm" value="0.5">0.5 mm</quantity>
long; anthers yellow, oblanceolate, ca. 1.5 ×
<quantity id="4C9AFDDD7075793F900EA5CDC8DFE7B4" box="[1331,1447,160,185]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="9" pageNumber="67" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
, apex emarginate. Female flower: pedicel pale green, ca.
<quantity id="4C9AFDDD7075793F97C6A5A9CE44E7D0" box="[763,828,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="67" unit="mm" value="9.0">9 mm</quantity>
long, glabrous; ovary broadly obovate, ca. 10 ×
<quantity id="4C9AFDDD7075793F906AA5A9C8DFE7D0" box="[1367,1447,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="67" unit="mm" value="15.0">15 mm</quantity>
, pale green-white, wings 3, equal, locules 3, placentas 2 per locule; tepals 5, whitish to white with a hint of pale green, ovate, 12–13 ×
<quantity id="4C9AFDDD7075793F9403A461CCD6E628" box="[318,430,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="9" pageNumber="67" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
, margin entire, apex rounded; styles 3, yellowish green, ca.
<quantity id="4C9AFDDD7075793F915AA461C9D2E628" box="[1127,1194,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="67" unit="mm" value="2.0">2 mm</quantity>
long, divided to base, shallowly Y-shaped above; stigma pale yellow, papillose forming a continuous twisted band. Fruit
<quantity id="4C9AFDDD7075793F91CDA45DC812E644" box="[1264,1386,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="9" pageNumber="67" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
long,
<quantity id="4C9AFDDD7075793F95B5A439CDA0E660" box="[136,216,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="67" unit="mm" value="17.0">17 mm</quantity>
wide across the wings, glabrous, locules 3, wings 3, dull green, equal,
<quantity id="4C9AFDDD7075793F96C3A439C924E660" box="[1022,1116,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="9" pageNumber="67" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
wide, thinly fibrous, rounded proximately, rounded to slightly truncate distally; pedicel pendent,
<quantity id="4C9AFDDD7075793F964AA415CE88E69C" box="[887,1008,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="9" pageNumber="67" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
long.
</paragraph>
</subSubSection>
<caption id="DF1D00B07074793E95B5A2E4C9F0E0CF" ID-DOI="http://doi.org/10.5281/zenodo.13726124" ID-Zenodo-Dep="13726124" httpUri="https://zenodo.org/record/13726124/files/figure.png" pageId="8" pageNumber="66" startId="8.[136,229,1929,1950]" targetBox="[160,1426,200,1894]" targetPageId="8" targetType="figure">
<paragraph id="8BDD50387074793E95B5A2E4C9F0E0CF" blockId="8.[136,1451,1928,1986]" pageId="8" pageNumber="66">
<emphasis id="B9168C2A7074793E95B5A2E4CD8FE093" bold="true" box="[136,247,1929,1950]" pageId="8" pageNumber="66">FIGURE 5</emphasis>
.
<taxonomicName id="4C622BBB7074793E943EA2E5CF93E093" authority="C. Y. Ling &amp; S. Julia. A. Habit" authorityName="C. Y. Ling &amp; S. Julia. A. Habit" authorityYear="2018" box="[259,747,1928,1950]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="66" phylum="Tracheophyta" rank="species" species="jokwaniana">
<emphasis id="B9168C2A7074793E943EA2E5CCB7E093" bold="true" box="[259,463,1928,1950]" italics="true" pageId="8" pageNumber="66">Begonia jokwaniana</emphasis>
C.Y.Ling &amp; S.Julia.
<emphasis id="B9168C2A7074793E97ABA2E4CFD6E093" bold="true" box="[662,686,1929,1950]" pageId="8" pageNumber="66">A.</emphasis>
Habit
</taxonomicName>
;
<emphasis id="B9168C2A7074793E97C9A2E4CE72E093" bold="true" box="[756,778,1929,1950]" pageId="8" pageNumber="66">B.</emphasis>
Stipule;
<emphasis id="B9168C2A7074793E965DA2E4CE00E093" bold="true" box="[864,888,1929,1950]" pageId="8" pageNumber="66">C.</emphasis>
Upper surface of leaf;
<emphasis id="B9168C2A7074793E916EA2E4C913E093" bold="true" box="[1107,1131,1929,1950]" pageId="8" pageNumber="66">D.</emphasis>
Male inflorescence;
<emphasis id="B9168C2A7074793E900EA2E4C80CE093" bold="true" box="[1331,1396,1929,1950]" pageId="8" pageNumber="66">E &amp; F.</emphasis>
Male flower;
<emphasis id="B9168C2A7074793E95EEA2C0CD94E0CF" bold="true" box="[211,236,1965,1986]" pageId="8" pageNumber="66">G.</emphasis>
Female flower;
<emphasis id="B9168C2A7074793E94B7A2C0CCDBE0CF" bold="true" box="[394,419,1965,1986]" pageId="8" pageNumber="66">H.</emphasis>
Fruit;
<emphasis id="B9168C2A7074793E94D9A2C0CC8BE0CF" bold="true" box="[484,499,1965,1986]" pageId="8" pageNumber="66">I.</emphasis>
Transverse section of fruit. All photos from SFC 6914 by C.Y. Ling.
</paragraph>
</caption>
<subSubSection id="C37803B37075793F9580A4F1CEE9E6D4" pageId="9" pageNumber="67" type="distribution">
<paragraph id="8BDD50387075793F9580A4F1C940E6B8" blockId="9.[136,1452,160,689]" box="[189,1080,412,437]" pageId="9" pageNumber="67">
<emphasis id="B9168C2A7075793F9580A4F1CC21E6B8" bold="true" box="[189,345,412,437]" pageId="9" pageNumber="67">Distribution:</emphasis>
—
<collectingCountry id="F37510A87075793F944BA4F1CF72E6B8" box="[374,522,412,437]" name="Malaysia" pageId="9" pageNumber="67">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="49A69EDA7075793F9748A4F1CFA1E6B8" box="[629,729,412,437]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="67">Sarawak</collectingRegion>
. Endemic in Gunung Mulu NP.
</paragraph>
<paragraph id="8BDD50387075793F9580A4ADCEE9E6D4" blockId="9.[136,1452,160,689]" box="[189,913,448,473]" pageId="9" pageNumber="67">
<emphasis id="B9168C2A7075793F9580A4ADCC5BE6D4" bold="true" box="[189,291,448,473]" pageId="9" pageNumber="67">Habitat:</emphasis>
—Mossy upper montane forest at ca.
<quantity id="4C9AFDDD7075793F97FAA4ADCE64E6D4" box="[711,796,448,473]" metricMagnitude="3" metricUnit="m" metricValue="1.5919999999999999" pageId="9" pageNumber="67" unit="m" value="1592.0">1592 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="C37803B37075793F9580A489C9FAE5BC" pageId="9" pageNumber="67" type="etymology">
<paragraph id="8BDD50387075793F9580A489CCBAE52C" blockId="9.[136,1452,160,689]" pageId="9" pageNumber="67">
<emphasis id="B9168C2A7075793F9580A489CC30E6F0" bold="true" box="[189,328,484,509]" pageId="9" pageNumber="67">Etymology:</emphasis>
—Named after Jok Wan, currently Park Ranger at Gunung Mulu NP, who facilitated the
<taxonomicName id="4C622BBB7075793F9073A489C8D3E6F0" box="[1358,1451,484,509]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B9168C2A7075793F9073A489C8D3E6F0" box="[1358,1451,484,509]" italics="true" pageId="9" pageNumber="67">Begonia</emphasis>
</taxonomicName>
survey at Gunung Mulu NP.
</paragraph>
<paragraph id="8BDD50387075793F9580A741C9EAE564" blockId="9.[136,1452,160,689]" pageId="9" pageNumber="67">
<emphasis id="B9168C2A7075793F9580A741CF3DE548" bold="true" box="[189,581,556,581]" pageId="9" pageNumber="67">Additional specimens examined:</emphasis>
—
<collectingCountry id="F37510A87075793F975FA741CF8EE548" box="[610,758,556,581]" name="Malaysia" pageId="9" pageNumber="67">MALAYSIA</collectingCountry>
. Borneo.
<collectingRegion id="49A69EDA7075793F9658A741CEB1E548" box="[869,969,556,581]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="67">Sarawak</collectingRegion>
. Marudi District: Gunung Mulu NP – Ulu Sg. Melinau Paku,
<emphasis id="B9168C2A7075793F9463A73DCF65E564" box="[350,541,592,617]" italics="true" pageId="9" pageNumber="67">Anderson S 4049</emphasis>
(SAR); Melinau Gorge,
<emphasis id="B9168C2A7075793F9609A73DC942E564" box="[820,1082,592,617]" italics="true" pageId="9" pageNumber="67">Abang Mohtar S 49525</emphasis>
(SAR).
</paragraph>
<paragraph id="8BDD50387075793F9580A719C9FAE5BC" blockId="9.[136,1452,160,689]" pageId="9" pageNumber="67">
<emphasis id="B9168C2A7075793F9580A719CC71E580" bold="true" box="[189,265,628,653]" pageId="9" pageNumber="67">Notes:</emphasis>
—The species is similar to
<taxonomicName id="4C622BBB7075793F9700A719CF97E580" box="[573,751,628,653]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="67" phylum="Tracheophyta" rank="species" species="melinauensis">
<emphasis id="B9168C2A7075793F9700A719CF97E580" box="[573,751,628,653]" italics="true" pageId="9" pageNumber="67">B. melinauensis</emphasis>
</taxonomicName>
in its broadly ovate leaves but is different in its glossy lamina (in life) that is sparsely bristly above, its male flowers with 2 tepals and its shorter capsule.
</paragraph>
</subSubSection>
</treatment>
</document>
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