240 lines
33 KiB
XML
240 lines
33 KiB
XML
<document id="BF12EE14497E661CA50F2EC022A2F8BD" ID-DOI="10.11646/zootaxa.5301.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8036117" ID-ZooBank="01F6333F-807B-448A-80C8-EC1F8CE030EA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686723262927" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2023" docId="002087EAFFA1947C64A8FCB7FE77FD9E" docLanguage="en" docName="zootaxa.5301.4.6.pdf" docOrigin="Zootaxa 5301 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5301.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba flaveola Shavrin 2023, sp.n." docType="treatment" docVersion="2" lastPageNumber="493" masterDocId="FC19FF92FFA4947B643FFFA9FFE2FFA3" masterDocTitle="New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines" masterLastPageNumber="500" masterPageNumber="486" pageNumber="491" updateTime="1686723785076" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DF015EBC1CE465F7605ACFC24CC0A99A">
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<mods:title id="C4EF7DF340779C979DDDDC3ECABE2146">New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines</mods:title>
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<mods:namePart id="113CF8BC7E81E7F752E4424331E19EB5">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="DE7D39F414B86DE3D3C4FDA338D050A5">text</mods:typeOfResource>
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<mods:title id="DD3820D2E0493190C5FA9A56F5D5323D">Zootaxa</mods:title>
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<mods:part id="9C7710B259044A01F5DED9FFB9C4EECA">
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<mods:date id="CB37A593988FB573C74121518252A3D4">2023</mods:date>
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<mods:number id="A628BE87365B4D193C7A677799D5E54E">2023-06-13</mods:number>
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<mods:detail id="C85BE613B64BB006C4E358B3049552A5" type="volume">
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<mods:number id="A66F6008BD356CC54582BC0ECAEBACC2">5301</mods:number>
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<mods:start id="BE67024877DB3B73F5FA5B593A05540B">486</mods:start>
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<mods:end id="045661F5643E9D918341C7C50A81A9F6">500</mods:end>
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<mods:url id="7F5199510E23E279B5B3BA3147C7B4E5">http://dx.doi.org/10.11646/zootaxa.5301.4.6</mods:url>
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<mods:classification id="8DD21E57AC6651143E2DA762C5589C9D">journal article</mods:classification>
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<mods:identifier id="D1A29F763F6E34289740C642CC2174ED" type="DOI">10.11646/zootaxa.5301.4.6</mods:identifier>
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<mods:identifier id="DD544E41C3D28FA108ED36845FA8EA6F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="002087EAFFA1947C64A8FCB7FE77FD9E" ID-DOI="http://doi.org/10.5281/zenodo.8036109" ID-Zenodo-Dep="8036109" LSID="urn:lsid:plazi:treatment:002087EAFFA1947C64A8FCB7FE77FD9E" httpUri="http://treatment.plazi.org/id/002087EAFFA1947C64A8FCB7FE77FD9E" lastPageId="7" lastPageNumber="493" pageId="5" pageNumber="491">
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<subSubSection id="C0936577FFA1947E64A8FCB7FDE4FC9A" box="[151,518,798,825]" pageId="5" pageNumber="491" type="nomenclature">
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<paragraph id="883636FCFFA1947E64A8FCB7FDE4FC9A" blockId="5.[151,518,798,861]" box="[151,518,798,825]" pageId="5" pageNumber="491">
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<heading id="D37E8190FFA1947E64A8FCB7FDE4FC9A" bold="true" box="[151,518,798,825]" fontSize="11" level="1" pageId="5" pageNumber="491" reason="1">
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<emphasis id="BAFDEAEEFFA1947E64A8FCB7FDE4FC9A" bold="true" box="[151,518,798,825]" pageId="5" pageNumber="491">
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||
<taxonomicName id="4F894D7FFFA1947E64A8FCB7FE26FC9A" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[151,452,798,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="flaveola" status="sp.n.">
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<emphasis id="BAFDEAEEFFA1947E64A8FCB7FE26FC9A" bold="true" box="[151,452,798,825]" italics="true" pageId="5" pageNumber="491">Paraphloeostiba flaveola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A1CE5795FFA1947E65F4FCB6FDE4FC9A" box="[459,518,799,825]" pageId="5" pageNumber="491" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C0936577FFA1947E64A8FCEAFEA8FCFE" box="[151,330,834,861]" pageId="5" pageNumber="491" type="description">
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||
<paragraph id="883636FCFFA1947E64A8FCEAFEA8FCFE" blockId="5.[151,518,798,861]" box="[151,330,834,861]" pageId="5" pageNumber="491">
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||
(
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||
<figureCitation id="10B22A79FFA1947E64A0FCEAFF0CFCFE" box="[159,238,835,861]" captionStart="FIGURES 1–6" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="5" pageNumber="491">Figs. 3</figureCitation>
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,
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<figureCitation id="10B22A79FFA1947E64C4FCEBFEA0FCFE" box="[251,322,834,861]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="5" pageNumber="491">15–21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C0936577FFA1947E64A8FC22FD8FFBFA" pageId="5" pageNumber="491" type="materials_examined">
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<paragraph id="883636FCFFA1947E64A8FC22FCEEFC4E" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">
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<emphasis id="BAFDEAEEFFA1947E64A8FC22FEA6FC07" bold="true" box="[151,324,906,933]" pageId="5" pageNumber="491">Type material.</emphasis>
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<typeStatus id="5732885EFFA1947E6574FC22FE51FC06" box="[331,435,907,933]" pageId="5" pageNumber="491">Holotype</typeStatus>
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♁ (
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<figureCitation id="10B22A79FFA1947E65E2FC22FDC2FC06" box="[477,544,907,933]" captionStart="FIGURES 1–6" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="5" pageNumber="491">Fig. 3</figureCitation>
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; dissected): ‘12 VIII 79 [handwritten in black] |
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<collectingCountry id="F09E766CFFA1947E6007FC22FB92FC06" box="[1080,1136,907,933]" name="Papua New Guinea" pageId="5" pageNumber="491">PNG</collectingCountry>
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/EHProv. | Umg. Kainantu | Onerunka <printed>, Papua | Nlle Guinée | W. G. Ullrich` <printed>, ‘
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<typeStatus id="5732885EFFA1947E67FDFC06FBB9FC6A" box="[962,1115,943,969]" pageId="5" pageNumber="491">HOLOTYPE</typeStatus>
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<taxonomicName id="4F894D7FFFA1947E604EFC06FACEFC6A" authorityName="Steel" authorityYear="1960" box="[1137,1324,943,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="genus">
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<emphasis id="BAFDEAEEFFA1947E604EFC06FACEFC6A" box="[1137,1324,943,969]" italics="true" pageId="5" pageNumber="491">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="4F894D7FFFA1947E617DFC06FA7EFC6A" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[1346,1436,943,969]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="flaveola" status="sp.n.">
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<emphasis id="BAFDEAEEFFA1947E617DFC06FA7EFC6A" box="[1346,1436,943,969]" italics="true" pageId="5" pageNumber="491">flaveola</emphasis>
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</taxonomicName>
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<emphasis id="BAFDEAEEFFA1947E64A8FC7AFF2DFC4E" bold="true" box="[151,207,979,1005]" pageId="5" pageNumber="491">
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<taxonomicNameLabel id="A1CE5795FFA1947E64A8FC7AFF2DFC4E" box="[151,207,979,1005]" pageId="5" pageNumber="491" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2023`<red, printed> (
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<collectionCode id="EE98AE39FFA1947E669BFC7AFCE2FC4E" box="[676,768,979,1005]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="491" type="Museum">MHNG</collectionCode>
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).
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</paragraph>
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<paragraph id="883636FCFFA1947E64F8FC5EFD8FFBFA" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">
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||
<materialsCitation id="38E13CA1FFA1947E64F8FC5EFDAEFB96" country="Guinea-Bissau" location="Onerunka" municipality="Kainantu" pageId="5" pageNumber="491" specimenCount="4" specimenCount-female="3" typeStatus="paratype">
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<typeStatus id="5732885EFFA1947E64F8FC5EFEDAFBB2" box="[199,312,1015,1041]" pageId="5" pageNumber="491">Paratypes</typeStatus>
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:
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<specimenCount id="9E8FFD75FFA1947E657BFC5EFE67FBB2" box="[324,389,1015,1041]" pageId="5" pageNumber="491" type="female">3 ♀♀</specimenCount>
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(
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<specimenCount id="9E8FFD75FFA1947E65AAFC5EFDD3FBB2" box="[405,561,1015,1041]" pageId="5" pageNumber="491" type="generic">one specimen</specimenCount>
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dissected): ‘PAPUA N.
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<collectingCountry id="F09E766CFFA1947E6779FC5EFC52FBB2" box="[838,944,1015,1041]" name="Guinea-Bissau" pageId="5" pageNumber="491">GUINEA</collectingCountry>
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<location id="8D566027FFA1947E67F9FC5EFBD6FBB2" LSID="urn:lsid:plazi:treatment:002087EAFFA1947C64A8FCB7FE77FD9E:8D566027FFA1947E67F9FC5EFBD6FBB2" box="[966,1076,1015,1041]" country="Guinea-Bissau" municipality="Kainantu" name="Onerunka" pageId="5" pageNumber="491">Onerunka</location>
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VIII. 79 [handwritten in black] | nr
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<collectingMunicipality id="6852AC86FFA1947E64FBFBB2FEC9FB96" box="[196,299,1051,1077]" pageId="5" pageNumber="491">Kainantu</collectingMunicipality>
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| W. G. Ullrich` <printed
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</materialsCitation>
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>, ‘
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<materialsCitation id="38E13CA1FFA1947E6655FBB2FD88FBFA" collectionCode="MHNG" collectorName="Shavrin A. V." country="Guinea-Bissau" pageId="5" pageNumber="491" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
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<typeStatus id="5732885EFFA1947E6655FBB2FD18FB96" box="[618,762,1051,1077]" pageId="5" pageNumber="491">PARATYPE</typeStatus>
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<taxonomicName id="4F894D7FFFA1947E672FFBB2FC29FB96" authorityName="Steel" authorityYear="1960" box="[784,971,1051,1077]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="genus">
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<emphasis id="BAFDEAEEFFA1947E672FFBB2FC29FB96" box="[784,971,1051,1077]" italics="true" pageId="5" pageNumber="491">Paraphloeostiba</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="4F894D7FFFA1947E67DFFBB2FBD8FB96" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[992,1082,1051,1077]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="flaveola" status="sp.n.">
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<emphasis id="BAFDEAEEFFA1947E67DFFBB2FBD8FB96" box="[992,1082,1051,1077]" italics="true" pageId="5" pageNumber="491">flaveola</emphasis>
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</taxonomicName>
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<emphasis id="BAFDEAEEFFA1947E607DFBB2FB9BFB96" bold="true" box="[1090,1145,1051,1077]" pageId="5" pageNumber="491">
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<taxonomicNameLabel id="A1CE5795FFA1947E607DFBB2FB9BFB96" box="[1090,1145,1051,1077]" pageId="5" pageNumber="491" rank="species">sp.n.</taxonomicNameLabel>
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||
</emphasis>
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||
|
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||
<collectorName id="257C532AFFA1947E60B0FBB2FAFDFB96" box="[1167,1311,1051,1077]" pageId="5" pageNumber="491">Shavrin A.V.</collectorName>
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des. 2023`<red, printed> (
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||
<specimenCount id="9E8FFD75FFA1947E6575FB96FE96FBFA" box="[330,372,1087,1113]" pageId="5" pageNumber="491" type="female">1 ♀</specimenCount>
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: cSh,
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<specimenCount id="9E8FFD75FFA1947E6586FB96FE18FBFA" box="[441,506,1087,1113]" pageId="5" pageNumber="491" type="female">2 ♀♀</specimenCount>
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||
:
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<collectionCode id="EE98AE39FFA1947E6638FB96FD80FBFA" box="[519,610,1087,1113]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="491" type="Museum">MHNG</collectionCode>
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||
)
|
||
</materialsCitation>
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||
.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C0936577FFA1947C64F8FBCBFDA1FE0E" lastPageId="7" lastPageNumber="493" pageId="5" pageNumber="491" type="description">
|
||
<paragraph id="883636FCFFA1947E64F8FBCBFE69FB4A" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">
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||
<emphasis id="BAFDEAEEFFA1947E64F8FBCBFEBBFBDF" bold="true" box="[199,345,1122,1148]" pageId="5" pageNumber="491">Description.</emphasis>
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Measurements (n=4): HW: 0.37–0.41; HL: 0.20–0.23; OL: 0.11–0.12; TL: 0.01–0.02; AL (
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<typeStatus id="5732885EFFA1947E64A0FB2EFEEAFB02" box="[159,264,1159,1185]" pageId="5" pageNumber="491">holotype</typeStatus>
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): 0.57; PL: 0.28–0.33; PWmax: 0.55–0.57; PWmin: 0.47–0.52; ESL: 0.50–0.56; EW: 0.63–0.68; MTbL (
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||
<typeStatus id="5732885EFFA1947E64A0FB02FEEAFB66" box="[159,264,1195,1221]" pageId="5" pageNumber="491">holotype</typeStatus>
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||
): 0.30; MTrL (
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<typeStatus id="5732885EFFA1947E6591FB02FDF5FB66" box="[430,535,1195,1221]" pageId="5" pageNumber="491">holotype</typeStatus>
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||
): 0.16 (MTrL 1–4: 0.06; MTrL 5: 0.10); AW: 0.62–0.66; AedL: 0.45; BL: 2.00– 2.35 (
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<typeStatus id="5732885EFFA1947E64E8FB66FEDDFB4A" box="[215,319,1231,1257]" pageId="5" pageNumber="491">holotype</typeStatus>
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: 2.18).
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</paragraph>
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<paragraph id="883636FCFFA1947E64F8FB5AFE3AF98E" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">
|
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Habitus as in
|
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<figureCitation id="10B22A79FFA1947E655BFB5AFE4BFAAE" box="[356,425,1267,1293]" captionStart="FIGURES 1–6" captionStartId="2.[151,275,1541,1567]" captionTargetBox="[197,1385,250,1508]" captionTargetId="figure-45@2.[187,1400,237,1520]" captionTargetPageId="2" captionText="FIGURES 1–6. Habitus of Paraphloeostiba: 1—P. conjuncta (holotype), 2—P. coriacea, 3—P. flaveola (holotype), 4—P. gayndahensis (Papua New Guinea), 5—P. papuana (holotype), 6—P. ullrichi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8036119" httpUri="https://zenodo.org/record/8036119/files/figure.png" pageId="5" pageNumber="491">Fig. 3</figureCitation>
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. Body yellow-brown, with slightly darker apical portion of head, latero-apical and apical portions of elytra, and sometimes apical margins of abdominal tergites; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Pronotum with indistinct and very sparse fine punctation, larger in lateral and indistinct or missing in middle portions; punctation of latero-apical and lateral portions of elytra slightly larger and deeper than that in pronotum; abdominal tergites without or with very fine and sparse punctation more distinct in lateral portion of each tergite. Head with coarse and dense microsculpture; neck with dense and fine transverse meshes; pronotum with very dense isodiametric sculpture, slightly finer than that in middle portion of head; microsculpture of elytra isodiametric, slightly coarser and larger than that on pronotum; abdomen with dense and transverse microreticulation.
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</paragraph>
|
||
<paragraph id="883636FCFFA1947E64F8F99EFD5AF88A" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">Head 1.7–1.8 times as broad as long, with short (about as long as diameter of ocellus) and deep anteocellar foveae; postocular carina present, indistinct and subacute, distance between posterior margin of eye and carina about as long as one nearest ommatidium. Apical maxillary palpomere narrow, slightly more than twice as long as preapical segment. Antennomere 7 slightly and antennomeres 8–10 strongly transverse; antennomere 2 slightly narrower than basal antennomere, 3 distinctly narrower than 2, 4 small, about twice shorter than 3, 5 slightly broader and shorter than 4, 6 about twice broader and slightly longer than 5, 7–8 slightly broader than 6, 9–10 slightly longer than 8, apical antennomere twice longer than 10.</paragraph>
|
||
<paragraph id="883636FCFFA1947E64F8F89AFE70F81A" blockId="5.[151,1437,906,1977]" pageId="5" pageNumber="491">
|
||
Pronotum distinctly transverse, 1.7–1.8 times as broad as long, 1.3–1.4 times as broad as head, widest in about middle; laterobasal strongly concaved in front of obtuse or subacute posterior angles; anterior margin widely concave in middle; middle portion without visible paramedian impressions except of
|
||
<specimenCount id="9E8FFD75FFA1947E67D5F8D2FB67F836" box="[1002,1157,1915,1941]" pageId="5" pageNumber="491" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
with very indistinct long and wide impressions.
|
||
</paragraph>
|
||
<caption id="DCF66674FFA2947D64A8F982FC2EF90C" ID-DOI="http://doi.org/10.5281/zenodo.8036123" ID-Zenodo-Dep="8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="6" pageNumber="492" startId="6.[151,264,1579,1603]" targetBox="[168,1422,200,1548]" targetPageId="6" targetType="figure">
|
||
<paragraph id="883636FCFFA2947D64A8F982FC2EF90C" blockId="6.[151,1437,1579,1712]" pageId="6" pageNumber="492">
|
||
<emphasis id="BAFDEAEEFFA2947D64A8F982FEB5F9E0" bold="true" box="[151,343,1579,1603]" pageId="6" pageNumber="492">FIGURES 15–25.</emphasis>
|
||
Aedeagus and abdominal segments of
|
||
<taxonomicName id="4F894D7FFFA2947D66CDF982FC14F9E0" authorityName="Shavrin" authorityYear="2023" box="[754,1014,1579,1603]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="flaveola">
|
||
<emphasis id="BAFDEAEEFFA2947D66CDF982FC14F9E0" box="[754,1014,1579,1603]" italics="true" pageId="6" pageNumber="492">Paraphloeostiba flaveola</emphasis>
|
||
</taxonomicName>
|
||
(Figs, 15–21: holotype) and
|
||
<taxonomicName id="4F894D7FFFA2947D6117F985FA7FF9E0" baseAuthorityName="Cameron" baseAuthorityYear="1931" box="[1320,1437,1580,1603]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="BAFDEAEEFFA2947D6117F985FA7FF9E0" box="[1320,1437,1580,1603]" italics="true" pageId="6" pageNumber="492">P. papuana</emphasis>
|
||
</taxonomicName>
|
||
(Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="883636FCFFA2947D64F8F8A8FB0DF8B8" blockId="6.[151,1436,1793,2036]" box="[199,1263,1793,1820]" pageId="6" pageNumber="492">Elytra 1.2 times as broad as long, 1.6–1.7 times as long as pronotum; posterior margin straight.</paragraph>
|
||
<paragraph id="883636FCFFA2947D64F8F88CFD6BF89C" blockId="6.[151,1436,1793,2036]" box="[199,649,1829,1856]" pageId="6" pageNumber="492">Abdomen slightly narrower than elytra,.</paragraph>
|
||
<paragraph id="883636FCFFA2947C64F8F8E0FECBFF76" blockId="6.[151,1436,1793,2036]" lastBlockId="7.[151,1437,150,573]" lastPageId="7" lastPageNumber="493" pageId="6" pageNumber="492">
|
||
Male. Posterior margin of abdominal tergite VIII truncate (
|
||
<figureCitation id="10B22A79FFA2947D674CF8E0FC2BF8C7" box="[883,969,1865,1892]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="6" pageNumber="492">Fig. 17</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII widely and deeply concave (
|
||
<figureCitation id="10B22A79FFA2947D662FF8C4FD86F82B" box="[528,612,1901,1928]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="6" pageNumber="492">Fig. 18</figureCitation>
|
||
). Aedeagus with wide basal bulb, gradually narrowed toward moderately narrow and rounded apical portion; parameres significantly narrowed apically toward widely rounded apical lobes, distinctly exceeding apex of median lobe, with three moderately long apical setae, with a row of 4–6 of moderately short setae on inner margins and 2–3 small setae on outer margins of eache paramere; internal sac moderately long and wide, with several sclerotized and very long structures in basal portion (
|
||
<figureCitation id="10B22A79FFA3947C67D9FF3EFBDBFF12" box="[998,1081,150,177]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">Fig. 15</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="10B22A79FFA3947C64EDFF12FEC7FF76" box="[210,293,187,213]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">Fig. 16</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="883636FCFFA3947C64F8FF76FB15FEBE" blockId="7.[151,1437,150,573]" pageId="7" pageNumber="493">
|
||
Female. Posterior margin of abdominal tergite VIII straight (
|
||
<figureCitation id="10B22A79FFA3947C6744FF76FC32FF5A" box="[891,976,223,249]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">Fig. 19</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII widely rounded (
|
||
<figureCitation id="10B22A79FFA3947C65B2FEAAFE03FEBE" box="[397,481,259,285]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">Fig. 20</figureCitation>
|
||
). Female accessory sclerite as in
|
||
<figureCitation id="10B22A79FFA3947C676AFEAAFC45FEBE" box="[853,935,259,285]" captionStart="FIGURES 15–25" captionStartId="6.[151,264,1579,1603]" captionTargetBox="[168,1422,200,1548]" captionTargetId="figure-15@6.[151,1436,181,1555]" captionTargetPageId="6" captionText="FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8036123" httpUri="https://zenodo.org/record/8036123/files/figure.png" pageId="7" pageNumber="493">Fig. 21</figureCitation>
|
||
. Spermatheca not recognized.
|
||
</paragraph>
|
||
<paragraph id="883636FCFFA3947C64F8FE8FFDA1FE0E" blockId="7.[151,1437,150,573]" pageId="7" pageNumber="493">
|
||
<emphasis id="BAFDEAEEFFA3947C64F8FE8FFE53FEE2" bold="true" box="[199,433,294,321]" pageId="7" pageNumber="493">Comparative notes.</emphasis>
|
||
Based on the shape of the lateral portions of the pronotum widely and deeply concave in front of posterior margin,
|
||
<taxonomicName id="4F894D7FFFA3947C6587FEE2FDCFFEC6" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[440,557,331,357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="flaveola" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFA3947C6587FEE2FDCFFEC6" box="[440,557,331,357]" italics="true" pageId="7" pageNumber="493">P. flaveola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFA3947C660CFEE2FD89FEC6" bold="true" box="[563,619,331,357]" pageId="7" pageNumber="493">
|
||
<taxonomicNameLabel id="A1CE5795FFA3947C660CFEE2FD89FEC6" box="[563,619,331,357]" pageId="7" pageNumber="493" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4F894D7FFFA3947C66C4FEE2FC66FEC7" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[763,900,331,356]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="conjuncta" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFA3947C66C4FEE2FC66FEC7" box="[763,900,331,356]" italics="true" pageId="7" pageNumber="493">P. conjuncta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFA3947C67B4FEE2FC20FEC6" bold="true" box="[907,962,331,357]" pageId="7" pageNumber="493">
|
||
<taxonomicNameLabel id="A1CE5795FFA3947C67B4FEE2FC20FEC6" box="[907,962,331,357]" pageId="7" pageNumber="493" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, from which it can be distinguished by the paler coloration of the body, the missing of paramedian impressions on the pronotum, and broader apical portions of the median lobe and the parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFA3947C64F8FE1FFE95FE56" pageId="7" pageNumber="493" type="distribution">
|
||
<paragraph id="883636FCFFA3947C64F8FE1FFE95FE56" blockId="7.[151,1437,150,573]" pageId="7" pageNumber="493">
|
||
<emphasis id="BAFDEAEEFFA3947C64F8FE1FFE83FE73" bold="true" box="[199,353,438,464]" pageId="7" pageNumber="493">Distribution.</emphasis>
|
||
<taxonomicName id="4F894D7FFFA3947C6554FE1EFD68FE72" authority="Shavrin, 2023" authorityName="Shavrin" authorityYear="2023" box="[363,650,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="flaveola" status="sp.n.">
|
||
<emphasis id="BAFDEAEEFFA3947C6554FE1EFD68FE72" box="[363,650,439,465]" italics="true" pageId="7" pageNumber="493">Paraphloeostiba flaveola</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BAFDEAEEFFA3947C66AAFE1EFD2FFE72" bold="true" box="[661,717,439,465]" pageId="7" pageNumber="493">
|
||
<taxonomicNameLabel id="A1CE5795FFA3947C66AAFE1EFD2FFE72" box="[661,717,439,465]" pageId="7" pageNumber="493" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="5732885EFFA3947C67C9FE1EFBC4FE72" box="[1014,1062,439,465]" pageId="7" pageNumber="493">type</typeStatus>
|
||
locality in north-eastern part of
|
||
<collectingCountry id="F09E766CFFA3947C64A8FE72FE91FE56" box="[151,371,475,501]" name="Papua New Guinea" pageId="7" pageNumber="493">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0936577FFA3947C64F8FE56FE77FD9E" pageId="7" pageNumber="493" type="etymology">
|
||
<paragraph id="883636FCFFA3947C64F8FE56FE77FD9E" blockId="7.[151,1437,150,573]" pageId="7" pageNumber="493">
|
||
<emphasis id="BAFDEAEEFFA3947C64F8FE56FEACFDBA" bold="true" box="[199,334,511,537]" pageId="7" pageNumber="493">Etymology.</emphasis>
|
||
The specific epithet is the Latin adjective
|
||
<emphasis id="BAFDEAEEFFA3947C6704FE56FC42FDBA" box="[827,928,511,537]" italics="true" pageId="7" pageNumber="493">flaveolus</emphasis>
|
||
, -
|
||
<emphasis id="BAFDEAEEFFA3947C6785FE56FC2AFDBB" box="[954,968,511,536]" italics="true" pageId="7" pageNumber="493">a</emphasis>
|
||
, -
|
||
<emphasis id="BAFDEAEEFFA3947C67DEFE56FBE1FDBB" box="[993,1027,511,536]" italics="true" pageId="7" pageNumber="493">um</emphasis>
|
||
(yellowish). It refers to the general coloration of the body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |