New species and records of the genus Paraphloeostiba Steel, 1960 (Coleoptera: Staphylinidae: Omaliinae) from Papua New Guinea, and notes on some species from India, Indonesia and Philippines Author Shavrin, Alexey V. text Zootaxa 2023 2023-06-13 5301 4 486 500 http://dx.doi.org/10.11646/zootaxa.5301.4.6 journal article 10.11646/zootaxa.5301.4.6 1175-5326 8036117 01F6333F-807B-448A-80C8-EC1F8CE030EA Paraphloeostiba flaveola sp.n. ( Figs. 3 , 15–21 ) Type material. Holotype ♁ ( Fig. 3 ; dissected): ‘12 VIII 79 [handwritten in black] | PNG /EHProv. | Umg. Kainantu | Onerunka <printed>, Papua | Nlle Guinée | W. G. Ullrich` <printed>, ‘ HOLOTYPE | Paraphloeostiba | flaveola sp.n. | Shavrin A.V. des. 2023`<red, printed> ( MHNG ). Paratypes : 3 ♀♀ ( one specimen dissected): ‘PAPUA N. GUINEA | Onerunka VIII. 79 [handwritten in black] | nr Kainantu | W. G. Ullrich` <printed >, ‘ PARATYPE | Paraphloeostiba | flaveola sp.n. | Shavrin A.V. des. 2023`<red, printed> ( 1 ♀ : cSh, 2 ♀♀ : MHNG ) . Description. Measurements (n=4): HW: 0.37–0.41; HL: 0.20–0.23; OL: 0.11–0.12; TL: 0.01–0.02; AL ( holotype ): 0.57; PL: 0.28–0.33; PWmax: 0.55–0.57; PWmin: 0.47–0.52; ESL: 0.50–0.56; EW: 0.63–0.68; MTbL ( holotype ): 0.30; MTrL ( holotype ): 0.16 (MTrL 1–4: 0.06; MTrL 5: 0.10); AW: 0.62–0.66; AedL: 0.45; BL: 2.00– 2.35 ( holotype : 2.18). Habitus as in Fig. 3 . Body yellow-brown, with slightly darker apical portion of head, latero-apical and apical portions of elytra, and sometimes apical margins of abdominal tergites; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow. Pronotum with indistinct and very sparse fine punctation, larger in lateral and indistinct or missing in middle portions; punctation of latero-apical and lateral portions of elytra slightly larger and deeper than that in pronotum; abdominal tergites without or with very fine and sparse punctation more distinct in lateral portion of each tergite. Head with coarse and dense microsculpture; neck with dense and fine transverse meshes; pronotum with very dense isodiametric sculpture, slightly finer than that in middle portion of head; microsculpture of elytra isodiametric, slightly coarser and larger than that on pronotum; abdomen with dense and transverse microreticulation. Head 1.7–1.8 times as broad as long, with short (about as long as diameter of ocellus) and deep anteocellar foveae; postocular carina present, indistinct and subacute, distance between posterior margin of eye and carina about as long as one nearest ommatidium. Apical maxillary palpomere narrow, slightly more than twice as long as preapical segment. Antennomere 7 slightly and antennomeres 8–10 strongly transverse; antennomere 2 slightly narrower than basal antennomere, 3 distinctly narrower than 2, 4 small, about twice shorter than 3, 5 slightly broader and shorter than 4, 6 about twice broader and slightly longer than 5, 7–8 slightly broader than 6, 9–10 slightly longer than 8, apical antennomere twice longer than 10. Pronotum distinctly transverse, 1.7–1.8 times as broad as long, 1.3–1.4 times as broad as head, widest in about middle; laterobasal strongly concaved in front of obtuse or subacute posterior angles; anterior margin widely concave in middle; middle portion without visible paramedian impressions except of two paratypes with very indistinct long and wide impressions. FIGURES 15–25. Aedeagus and abdominal segments of Paraphloeostiba flaveola (Figs, 15–21: holotype) and P. papuana (Figs. 22–25): 15—aedeagus, parameral view, 16—aedeagus, (lateral view), 17—male tergite VIII, dorsal view, 18—male sternite VIII, ventral view, 19, 22—female tergite VIII, dorsal view, 20, 23—female sternite VIII, ventral view, 21, 24—female genital segment, ventral view, 25—spermatheca, lateral view. Scale bar: 0.1 mm. Elytra 1.2 times as broad as long, 1.6–1.7 times as long as pronotum; posterior margin straight. Abdomen slightly narrower than elytra,. Male. Posterior margin of abdominal tergite VIII truncate ( Fig. 17 ). Posterior margin of abdominal sternite VIII widely and deeply concave ( Fig. 18 ). Aedeagus with wide basal bulb, gradually narrowed toward moderately narrow and rounded apical portion; parameres significantly narrowed apically toward widely rounded apical lobes, distinctly exceeding apex of median lobe, with three moderately long apical setae, with a row of 4–6 of moderately short setae on inner margins and 2–3 small setae on outer margins of eache paramere; internal sac moderately long and wide, with several sclerotized and very long structures in basal portion ( Fig. 15 ). Lateral aspect of the aedeagus as in Fig. 16 . Female. Posterior margin of abdominal tergite VIII straight ( Fig. 19 ). Posterior margin of abdominal sternite VIII widely rounded ( Fig. 20 ). Female accessory sclerite as in Fig. 21 . Spermatheca not recognized. Comparative notes. Based on the shape of the lateral portions of the pronotum widely and deeply concave in front of posterior margin, P. flaveola sp.n. is similar to P. conjuncta sp.n. , from which it can be distinguished by the paler coloration of the body, the missing of paramedian impressions on the pronotum, and broader apical portions of the median lobe and the parameres. Distribution. Paraphloeostiba flaveola sp.n. is known only from the type locality in north-eastern part of Papua New Guinea . Etymology. The specific epithet is the Latin adjective flaveolus , - a , - um (yellowish). It refers to the general coloration of the body.