treatments-xml/data/3C/90/DF/3C90DF0693B359C6AA3E2A1E72DCAFEA.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.99.99030" ID-Pensoft-Pub="1860-0743-1-195" ID-Pensoft-UUID="C59B2291F71D52B89499FD9433B4B57E" ID-ZooBank="E28DD35A30EA4C53B5A6F0A80CA0CFC0" ModsDocID="1860-0743-99-1-195" checkinTime="1678295867757" checkinUser="pensoft" docAuthor="Boonmachai, Tuangthong, Bergey, Elizabeth A. &amp; Nantarat, Nattawadee" docDate="2023" docId="3C90DF0693B359C6AA3E2A1E72DCAFEA" docLanguage="en" docName="ZoosystEvol 99(1): 195-207" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-03-07" docSource="http://dx.doi.org/10.3897/zse.99.99030" docTitle="Diplommatina bulonensis Boonmachai &amp; Nantarat 2023, sp. nov." docType="treatment" docUuid="94CA70AE-54F1-4B9D-8604-29766AE5FB80" docUuidSource="ZooBank" docVersion="2" id="C59B2291F71D52B89499FD9433B4B57E" lastPageNumber="195" masterDocId="C59B2291F71D52B89499FD9433B4B57E" masterDocTitle="First record and description of three new species in the land snail genus Diplommatina Benson, 1849 (Caenogastropoda, Diplommatinidae) from Satun Province, Thailand" masterLastPageNumber="207" masterPageNumber="195" pageNumber="195" updateTime="1678296169138" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>First record and description of three new species in the land snail genus Diplommatina Benson, 1849 (Caenogastropoda, Diplommatinidae) from Satun Province, Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Boonmachai, Tuangthong</mods:namePart>
<mods:affiliation>Ph. D. ' s Degree Program in Biology (International Program), Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bergey, Elizabeth A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7160-6590</mods:nameIdentifier>
<mods:affiliation>Department of Biology, Faculty of Science, Chiang Mai University 50200, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nantarat, Nattawadee</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3317-0041</mods:nameIdentifier>
<mods:affiliation>Ph. D. ' s Degree Program in Biology (International Program), Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">n_nantarat@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>99</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:extent unit="page">
<mods:start>195</mods:start>
<mods:end>207</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.99.99030</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.99.99030</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-195</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:94CA70AE-54F1-4B9D-8604-29766AE5FB80" httpUri="http://treatment.plazi.org/id/3C90DF0693B359C6AA3E2A1E72DCAFEA" lastPageNumber="195" pageId="0" pageNumber="195">
<subSubSection pageId="0" pageNumber="195" type="nomenclature">
<paragraph pageId="0" pageNumber="195">
<taxonomicName LSID="https://zoobank.org/94CA70AE-54F1-4B9D-8604-29766AE5FB80" authority="Boonmachai &amp; Nantarat" authorityName="Boonmachai &amp; Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Diplommatina bulonensis" order="Mesogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="bulonensis" status="sp. nov.">Diplommatina bulonensis Boonmachai &amp; Nantarat</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="195">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="description">
<paragraph pageId="0" pageNumber="195">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">Figs 2A</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">, 3A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="type material">
<paragraph pageId="0" pageNumber="195">Type material.</paragraph>
<paragraph pageId="0" pageNumber="195">
<emphasis bold="true" italics="true" pageId="0" pageNumber="195">Holotype</emphasis>
CMUZ 9050001 (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">2A</figureCitation>
); Shell measurements: SH = 2.72 mm, SW = 1.27 mm, AH = 0.94 mm, AW = 0.91 mm, W = 6 1/2. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="195">Paratypes</emphasis>
CMUZ 9050002-9050006 (5 shells); Shell measurements: SH = 2.58-2.82 mm, SW = 1.26-1.34 mm, AH = 0.97-1.04 mm, AW = 0.92-1.03 mm, W = 6 1/2-7 1/2.
</paragraph>
<caption doi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="195">
<emphasis bold="true" pageId="0" pageNumber="195">Figure 2.</emphasis>
Shell morphology of 
<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
</taxonomicName>
from Satun Province. 
<emphasis bold="true" pageId="0" pageNumber="195">A.</emphasis>
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); 
<emphasis bold="true" pageId="0" pageNumber="195">B.</emphasis>
<taxonomicName lsidName="D. laemsonensis" pageId="0" pageNumber="195" rank="species" species="laemsonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. laemsonensis</emphasis>
</taxonomicName>
sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); 
<emphasis bold="true" pageId="0" pageNumber="195">C.</emphasis>
<taxonomicName lsidName="D. prakaiphetensis" pageId="0" pageNumber="195" rank="species" species="prakaiphetensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. prakaiphetensis</emphasis>
</taxonomicName>
sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); 
<emphasis bold="true" pageId="0" pageNumber="195">D.</emphasis>
<taxonomicName lsidName="D. naiyanetri" pageId="0" pageNumber="195" rank="species" species="naiyanetri">
<emphasis italics="true" pageId="0" pageNumber="195">D. naiyanetri</emphasis>
</taxonomicName>
(CMUZ 905090; Tarutao Island); 
<emphasis bold="true" pageId="0" pageNumber="195">E.</emphasis>
<taxonomicName lsidName="D. canaliculata" pageId="0" pageNumber="195" rank="species" species="canaliculata">
<emphasis italics="true" pageId="0" pageNumber="195">D. canaliculata</emphasis>
</taxonomicName>
(CMUZ 9050158; Tharn Pliew Waterfall).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="type locality">
<paragraph pageId="0" pageNumber="195">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="195">
Thailand, Satun Province, La-ngu District, Bulon Pai Island on limestone rocks and under leaves, 
<geoCoordinate degrees="6" direction="north" minutes="49" orientation="latitude" precision="1" seconds="53.3" value="6.8314724">6°49'53.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="35" orientation="longitude" precision="1" seconds="19.5" value="99.58875">99°35'19.5&quot;E</geoCoordinate>
, 22 December 2020, coll. T. Booonmachai.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="materials_examined">
<paragraph pageId="0" pageNumber="195">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="195">
<materialsCitation collectingDate="2020-12-22" collectingDateMax="2022-02-15" collectingDateMin="2020-12-22" country="Thailand" county="La-ngu District" latitude="6.8314724" location="La-ngu District" longLatPrecision="1" longitude="99.58875" municipality="Bulon Pai Island" specimenCount="1" stateProvince="Satun Province">
<collectingCountry name="Thailand">Thailand</collectingCountry>
, 
<collectingRegion country="Thailand" name="Satun">Satun Province</collectingRegion>
; 
<collectingCounty>La-ngu District</collectingCounty>
, 
<collectingMunicipality>Bulon Pai Island</collectingMunicipality>
, 
<geoCoordinate degrees="6" direction="north" minutes="49" orientation="latitude" precision="1" seconds="53.3" value="6.8314724">6°49'53.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="35" orientation="longitude" precision="1" seconds="19.5" value="99.58875">99°35'19.5&quot;E</geoCoordinate>
, 
<collectingDate value="2020-12-22">22 December 2020</collectingDate>
: CMUZ 9050007-9050016 (10 shells); 
<location LSID="urn:lsid:plazi:treatment:3C90DF0693B359C6AA3E2A1E72DCAFEA:E11671797ECC718C2B981683A027B5BB" country="Thailand" county="La-ngu District" latitude="6.8314724" longLatPrecision="1" longitude="99.58875" municipality="Bulon Pai Island" name="La-ngu District" stateProvince="Satun Province">La-ngu District</location>
, 
<location LSID="urn:lsid:plazi:treatment:3C90DF0693B359C6AA3E2A1E72DCAFEA:591EEF8DA377CD75809CE7566FC80594" country="Thailand" county="La-ngu District" latitude="6.8314724" longLatPrecision="1" longitude="99.58875" municipality="Bulon Pai Island" name="Bulon Pai Island" stateProvince="Satun Province">Bulon Pai Island</location>
, 
<geoCoordinate degrees="6" direction="north" minutes="49" orientation="latitude" precision="1" seconds="53.3" value="6.8314724">6°49'53.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="99" direction="east" minutes="35" orientation="longitude" precision="1" seconds="19.5" value="99.58875">99°35'19.5&quot;E</geoCoordinate>
, 
<collectingDate value="2022-02-15">15 February 2022</collectingDate>
: CMUZ 9050017-9050018 (2 shells)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="etymology">
<paragraph pageId="0" pageNumber="195">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="195">
The specific epithet 
<emphasis italics="true" pageId="0" pageNumber="195">bulonensis</emphasis>
is an adjective referring to the type locality (Bulon Pai Island in Satun Province, Thailand).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="differential diagnosis">
<paragraph pageId="0" pageNumber="195">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="195">
Among approximately 400 species of 
<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
</taxonomicName>
, the new species is most similar to 
<taxonomicName lsidName="D. conditioria" pageId="0" pageNumber="195" rank="species" species="conditioria">
<emphasis italics="true" pageId="0" pageNumber="195">D. conditioria</emphasis>
</taxonomicName>
Maassen, 2007, 
<taxonomicName lsidName="D. baliana" pageId="0" pageNumber="195" rank="species" species="baliana">
<emphasis italics="true" pageId="0" pageNumber="195">D. baliana</emphasis>
</taxonomicName>
Fulton, 1899, 
<taxonomicName lsidName="D. maduana" pageId="0" pageNumber="195" rank="species" species="maduana">
<emphasis italics="true" pageId="0" pageNumber="195">D. maduana</emphasis>
</taxonomicName>
Laidlaw, 1949, 
<taxonomicName lsidName="D. gomantongensis" pageId="0" pageNumber="195" rank="species" species="gomantongensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. gomantongensis</emphasis>
</taxonomicName>
Smith, 1894 and 
<taxonomicName lsidName="D. antheae" pageId="0" pageNumber="195" rank="species" species="antheae">
<emphasis italics="true" pageId="0" pageNumber="195">D. antheae</emphasis>
</taxonomicName>
Vermeulen, 1993 (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P &lt;0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
). Typical characteristics shared by 
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
with all these species are shell dextral with a slender fusiform shape with rather evenly rounded sides, radial ribs not sinuous, constriction situated halfway on the parietal side of the peristome and aperture round. However, 
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov. differs from 
<taxonomicName lsidName="D. gomantongensis" pageId="0" pageNumber="195" rank="species" species="gomantongensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. gomantongensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. antheae" pageId="0" pageNumber="195" rank="species" species="antheae">
<emphasis italics="true" pageId="0" pageNumber="195">D. antheae</emphasis>
</taxonomicName>
by the presence of a transverse palatalis and an inconspicuous columellar denticle in the aperture (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell internal terminology of new species genus Diplommatina from Satun Province under the stereo microscope and the drawing. A, B. D. bulonensis sp. nov. (Bulon Pai Island); C, D. D. laemsonensis sp. nov. (Laem Son Hill); E, F. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure4" httpUri="https://binary.pensoft.net/fig/820987" pageId="0" pageNumber="195">4A</figureCitation>
). 
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov differs in the combination of shell height, shell width, aperture height and aperture width from 
<taxonomicName lsidName="D. maduana" pageId="0" pageNumber="195" rank="species" species="maduana">
<emphasis italics="true" pageId="0" pageNumber="195">D. maduana</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. antheae" pageId="0" pageNumber="195" rank="species" species="antheae">
<emphasis italics="true" pageId="0" pageNumber="195">D. antheae</emphasis>
</taxonomicName>
(Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P &lt;0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
). 
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov differs from 
<taxonomicName lsidName="D. conditioria" pageId="0" pageNumber="195" rank="species" species="conditioria">
<emphasis italics="true" pageId="0" pageNumber="195">D. conditioria</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. maduana" pageId="0" pageNumber="195" rank="species" species="maduana">
<emphasis italics="true" pageId="0" pageNumber="195">D. maduana</emphasis>
</taxonomicName>
in the number of whorls (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P &lt;0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
) and differs from 
<taxonomicName lsidName="D. maduana" pageId="0" pageNumber="195" rank="species" species="maduana">
<emphasis italics="true" pageId="0" pageNumber="195">D. maduana</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="D. gomantongensis" pageId="0" pageNumber="195" rank="species" species="gomantongensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. gomantongensis</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="D. antheae" pageId="0" pageNumber="195" rank="species" species="antheae">
<emphasis italics="true" pageId="0" pageNumber="195">D. antheae</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="D. baliana" pageId="0" pageNumber="195" rank="species" species="baliana">
<emphasis italics="true" pageId="0" pageNumber="195">D. baliana</emphasis>
</taxonomicName>
in the number of radial ribs/0.5 mm on the penultimate whorl (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P &lt;0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
). 
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov has fewer radial ribs on body whorls above the aperture than 
<taxonomicName lsidName="D. conditioria" pageId="0" pageNumber="195" rank="species" species="conditioria">
<emphasis italics="true" pageId="0" pageNumber="195">D. conditioria</emphasis>
</taxonomicName>
(12-13 vs. 16 ribs/1 mm) (
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="195" pagination="189 - 208" refId="B25" refString="Maassen, WJM, 2007. Notes on terrestrial molluscs of the island of Sulawesi. The genus Diplommatina (Gastropoda, Caenogastropoda, Diplommatinidae). Basteria 71: 189 - 208" title="Notes on terrestrial molluscs of the island of Sulawesi. The genus Diplommatina (Gastropoda, Caenogastropoda, Diplommatinidae)." volume="71" year="2007">Maassen 2007</bibRefCitation>
). Moreover, it differs from 
<taxonomicName lsidName="D. baliana" pageId="0" pageNumber="195" rank="species" species="baliana">
<emphasis italics="true" pageId="0" pageNumber="195">D. baliana</emphasis>
</taxonomicName>
in having a columella fold.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="195" type="description">
<paragraph pageId="0" pageNumber="195">Description.</paragraph>
<paragraph pageId="0" pageNumber="195">
Shell dextral, fusiform, convex and red rushes or red-orange in color (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">2A</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3A</figureCitation>
). Penultimate whorl slightly wider than body whorl, side flat. Spire conical with slightly convex sides. Suture impressed. Constriction level with the middle parietal side of the peristome, with three lamellae: one parietalis very distinct, one distinct and long transversal palatalis and one distinct columellaris. (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell internal terminology of new species genus Diplommatina from Satun Province under the stereo microscope and the drawing. A, B. D. bulonensis sp. nov. (Bulon Pai Island); C, D. D. laemsonensis sp. nov. (Laem Son Hill); E, F. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure4" httpUri="https://binary.pensoft.net/fig/820987" pageId="0" pageNumber="195">4A, B</figureCitation>
). Protoconch smooth with dimples and about 2 1/2 whorls (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3B</figureCitation>
). Radial ribs straight, inconspicuous but distinct, moderately spaced about 6 1/2-7 ribs/0.5 mm on the penultimate whorl (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P &lt;0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
). The aperture slightly tilted against the coiling axis, columellar denticle slightly weak in the aperture. Peristome double, rather expanding, palatal side not sinuous, basal side and columella side weakly sinuous. Outer peristome expanded less than inner peristome, inner peristome expanded at parietalis side, with a palatal lip. Umbilicus closed. Operculum multispiral, flat, corneous, transparent, slightly concave, outer surface smooth, inner surface smooth with raised peripheral circular margin (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3C</figureCitation>
). The radula of the taenioglossate type (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Radular morphology of Diplommatina. A. Diplommatina laemsonensis sp. nov. (Laem Son Hill); B. Diplommatina bulonensis sp. nov. (Bulon Pai Island); C. Diplommatina prakaiphetensis sp. nov. (Prakaiphet Hill). Colours show the position of the teeth of the radula; blue: central tooth, yellow: the basal plate of the central tooth, green: lateral teeth, pink: inner marginal teeth, and red: outer marginal teeth." figureDoi="10.3897/zse.99.99030.figure5" httpUri="https://binary.pensoft.net/fig/820988" pageId="0" pageNumber="195">5B</figureCitation>
). The central tooth round and flat with seven cusps of similar size. The basal plate of the central tooth, cuspate and small. The lateral teeth with five cusps. The marginal teeth with inner and outer teeth, the inner teeth with seven small cusps and the outer teeth with five cusps and larger than the inner teeth (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Radular morphology of Diplommatina. A. Diplommatina laemsonensis sp. nov. (Laem Son Hill); B. Diplommatina bulonensis sp. nov. (Bulon Pai Island); C. Diplommatina prakaiphetensis sp. nov. (Prakaiphet Hill). Colours show the position of the teeth of the radula; blue: central tooth, yellow: the basal plate of the central tooth, green: lateral teeth, pink: inner marginal teeth, and red: outer marginal teeth." figureDoi="10.3897/zse.99.99030.figure5" httpUri="https://binary.pensoft.net/fig/820988" pageId="0" pageNumber="195">5B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="195">
<emphasis bold="true" pageId="0" pageNumber="195">Figure 3.</emphasis>
Scanning electron microscope of shell morphology, protoconch and operculum of 
<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
</taxonomicName>
new species. 
<emphasis bold="true" pageId="0" pageNumber="195">A-C.</emphasis>
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov. (Bulon Pai Island); 
<emphasis bold="true" pageId="0" pageNumber="195">D-F.</emphasis>
<taxonomicName lsidName="D. laemsonensis" pageId="0" pageNumber="195" rank="species" species="laemsonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. laemsonensis</emphasis>
</taxonomicName>
sp. nov. (Laem Son Hill); 
<emphasis bold="true" pageId="0" pageNumber="195">G-I.</emphasis>
<taxonomicName lsidName="D. prakaiphetensis" pageId="0" pageNumber="195" rank="species" species="prakaiphetensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. prakaiphetensis</emphasis>
</taxonomicName>
sp. nov. (Prakaiphet Hill).
</paragraph>
</caption>
<caption doi="10.3897/zse.99.99030.figure4" httpUri="https://binary.pensoft.net/fig/820987" pageId="0" pageNumber="195" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="195">
<emphasis bold="true" pageId="0" pageNumber="195">Figure 4.</emphasis>
Shell internal terminology of new species genus 
<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
</taxonomicName>
from Satun Province under the stereo microscope and the drawing. 
<emphasis bold="true" pageId="0" pageNumber="195">A, B.</emphasis>
<taxonomicName lsidName="D. bulonensis" pageId="0" pageNumber="195" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. bulonensis</emphasis>
</taxonomicName>
sp. nov. (Bulon Pai Island); 
<emphasis bold="true" pageId="0" pageNumber="195">C, D.</emphasis>
<taxonomicName lsidName="D. laemsonensis" pageId="0" pageNumber="195" rank="species" species="laemsonensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. laemsonensis</emphasis>
</taxonomicName>
sp. nov. (Laem Son Hill); 
<emphasis bold="true" pageId="0" pageNumber="195">E, F.</emphasis>
<taxonomicName lsidName="D. prakaiphetensis" pageId="0" pageNumber="195" rank="species" species="prakaiphetensis">
<emphasis italics="true" pageId="0" pageNumber="195">D. prakaiphetensis</emphasis>
</taxonomicName>
sp. nov. (Prakaiphet Hill).
</paragraph>
</caption>
<caption doi="10.3897/zse.99.99030.figure5" httpUri="https://binary.pensoft.net/fig/820988" pageId="0" pageNumber="195" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="195">
<emphasis bold="true" pageId="0" pageNumber="195">Figure 5.</emphasis>
Radular morphology of 
<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
</taxonomicName>
. 
<emphasis bold="true" pageId="0" pageNumber="195">A.</emphasis>
<taxonomicName authorityName="Boonmachai &amp; Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina laemsonensis" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="laemsonensis">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina laemsonensis</emphasis>
</taxonomicName>
sp. nov. (Laem Son Hill); 
<emphasis bold="true" pageId="0" pageNumber="195">B.</emphasis>
<taxonomicName authorityName="Boonmachai &amp; Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina bulonensis" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="bulonensis">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina bulonensis</emphasis>
</taxonomicName>
sp. nov. (Bulon Pai Island); 
<emphasis bold="true" pageId="0" pageNumber="195">C.</emphasis>
<taxonomicName authorityName="Boonmachai &amp; Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina prakaiphetensis" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="prakaiphetensis">
<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina prakaiphetensis</emphasis>
</taxonomicName>
sp. nov. (Prakaiphet Hill). Colours show the position of the teeth of the radula; blue: central tooth, yellow: the basal plate of the central tooth, green: lateral teeth, pink: inner marginal teeth, and red: outer marginal teeth.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>