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<mods:title id="50D869BA6B274B5813F53C06321E2AD0">Snailfishes of the genus Careproctus (Perciformes: Liparidae) with a reduced pelvic disk: three new species and new records from the western North Pacific with comments on their phenotypic diversity</mods:title>
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<mods:namePart id="59B1072793988CA1D1FDF0960B5932FE">Kai, Yoshiaki</mods:namePart>
<mods:affiliation id="9C4297BC14B40A5F77929B5FECC46ADA">Maizuru Fisheries Research Station, Field Science Education and Research Center, Kyoto University, Nagahama, Maizuru, Kyoto 625 - 0086, Japan</mods:affiliation>
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<mods:namePart id="A9DA80CAAAAA462806B501E0C19EE886">Matsuzaki, Koji</mods:namePart>
<mods:affiliation id="D4AB0F17E6353E9C6AD15D88534DAD97">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:namePart id="9A5334AA631A12A8320E6E7E6EC63177">Mori, Toshiaki</mods:namePart>
<mods:affiliation id="77525B4CB8CAECEF40AA024E5FF6F7B7">Marine Science Museum, Fukushima (Aquamarine Fukushima), Onahama, Iwaki, Fukushima 971 - 8101, Japan</mods:affiliation>
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<mods:affiliation id="AD5D84275AE4EC406C5A00F866D8B551">A. V. Zhirmunsky National Scientific Center of Marine Biology, Far Eastern Branch of the Russian Academy of Sciences, ul. Palchevskogo 17, Vladivostok 690041, Russia</mods:affiliation>
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<mods:namePart id="46C70AB325DBCD46CE356E97196B56A5">Misawa, Ryo</mods:namePart>
<mods:affiliation id="31983C4EAC0448DF4EFEA20715FA50EE">Demersal Fish Resources Division, Fisheries Stock Assessment Center, Fisheries Resources Institute, Japan Fisheries Research and Education Agency, National Research and Development Agency, 25 - 259 Shimomekurakubo, Same, Hachinohe, Aomori 031 - 0841, Japan</mods:affiliation>
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<mods:affiliation id="798127355AF1400205CE1E7A94BD1B43">Fisheries Science Center, The Hokkaido University Museum, 3 - 1 - 1 Minato-cho, Hakodate, Hokkaido 041 - 8611, Japan</mods:affiliation>
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<taxonomicName id="4C2D4D61FF86386B8BCDAB6CD182FDFF" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[151,437,549,576]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF86386B8BCDAB6CD182FDFF" bold="true" box="[151,437,549,576]" italics="true" pageId="3" pageNumber="194">Careproctus barbatulus</emphasis>
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Kai, Matsuzaki &amp; Mori
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<paragraph id="8B9236E2FF86386B8BCDAB03D218FDDC" blockId="3.[151,975,549,683]" box="[151,559,585,612]" pageId="3" pageNumber="194">New Japanese name: Isaribi-bikunin</paragraph>
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,
<figureCitation id="13162A67FF86386B8BABAB24D172FD38" box="[241,325,621,647]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="3" pageNumber="194">2A, 2B</figureCitation>
;
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SL, female, off
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depth
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, F/
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, coll. by Koji Matsuzaki, Shigeki Fujimoto, and Shigetada Fujimoto.
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<typeStatus id="54968840FF86386B8B9DAA68D176FC84" box="[199,321,801,827]" pageId="3" pageNumber="194" type="paratype">Paratypes</typeStatus>
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(ex.
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),
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SL, male,
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depth
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, F/
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,
<date id="FF931022FF86386B885EAA0CD349FCE0" box="[772,894,837,863]" pageId="3" pageNumber="194" value="2018-08">
<collectingDate id="EFD7E9CAFF86386B885EAA0CD349FCE0" box="[772,894,837,863]" pageId="3" pageNumber="194" value="2018-08">Aug. 2018</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B88BFAA0CD4A1FCE0" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B88BFAA0CD4A1FCE0" box="[997,1174,837,863]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8FF8AA0CD55EFCE0" box="[1186,1385,837,863]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8BCDAA20D141FC3C" box="[151,374,873,899]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8AD8AA20D551FC3C" box="[386,1382,873,900]" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8AD8AA20D1FBFC3C" box="[386,460,873,899]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8A89AA20D291FC3B" box="[467,678,873,900]" metricMagnitude="5" metricUnit="m" metricValue="1.466331903" pageId="3" pageNumber="194" unit="mm" value="1.466331903E8">146633, 190.3 mm</quantity>
SL, female, off
<collectorName id="26D85334FF86386B8806AA23D392FC3B" box="[860,933,874,900]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B88F5AA20D417FC3C" box="[943,1056,873,899]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8F76AA23D458FC3B" box="[1068,1135,874,900]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B8FFAAA20D551FC3C" box="[1184,1382,873,900]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B8FFAAA20D528FC3B" box="[1184,1311,873,900]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8ED3AA20D4E8FC18" collectingDate="2014-09-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8ED3AA20D5ABFC3C" box="[1417,1436,873,899]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8BCDAAC7D111FC18" box="[151,294,910,935]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8BCDAAC7D111FC18" box="[151,294,910,935]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8A6DAAC4D1F2FC18" box="[311,453,909,935]" pageId="3" pageNumber="194" value="2014-09-08">
<collectingDate id="EFD7E9CAFF86386B8A6DAAC4D1F2FC18" box="[311,453,909,935]" pageId="3" pageNumber="194" value="2014-09-08">8 Sep. 2014</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8969AAC4D2D1FC18" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8969AAC4D2D1FC18" box="[563,742,909,935]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B89AEAAC4D38AFC18" box="[756,957,909,935]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B88A7AAC4D4E8FC18" box="[1021,1247,909,935]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8FB7AAC4D3F3FC74" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8FB7AAC4D500FC18" box="[1261,1335,909,935]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8E1BAAC4D13DFC73" metricMagnitude="5" metricUnit="m" metricValue="1.466342208" pageId="3" pageNumber="194" unit="mm" value="1.466342208E8">146634, 220.8 mm</quantity>
SL, male, off
<collectorName id="26D85334FF86386B8AEAAAFBD1CEFC73" box="[432,505,946,972]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B895CAAF8D240FC74" box="[518,631,945,971]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B89D9AAFBD2F1FC73" box="[643,710,946,972]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B89A1AAF8D3F3FC74" box="[763,964,945,972]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B89A1AAF8D34CFC73" box="[763,891,945,972]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B88B2AAF8D318FC50" collectingDate="2014-09-10" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B88B2AAF8D3CCFC74" box="[1000,1019,945,971]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8F5EAAFBD4A4FC74" box="[1028,1171,946,971]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8F5EAAFBD4A4FC74" box="[1028,1171,946,971]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8FF8AAF8D50CFC74" box="[1186,1339,945,971]" pageId="3" pageNumber="194" value="2014-09-10">
<collectingDate id="EFD7E9CAFF86386B8FF8AAF8D50CFC74" box="[1186,1339,945,971]" pageId="3" pageNumber="194" value="2014-09-10">10 Sep. 2014</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8BCDAA9CD170FC50" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8BCDAA9CD170FC50" box="[151,327,981,1007]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8A0BAA9CD220FC50" box="[337,535,981,1007]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B890AAA9CD318FC50" box="[592,815,981,1007]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8860AA9CD1FEFB88" collectionCode="FAKU" country="Japan" county="Shigetada Fujimoto" location="Rausu" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCode="FAKU 146807-146810, 4" specimenCount="3" specimenCount-female="3" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8860AA9CD475FC50" box="[826,1090,981,1007]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 146807146810</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8F17AA9CD46CFC50" box="[1101,1115,981,1007]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">4</specimenCode>
(
<specimenCount id="9D2BFD6BFF86386B8F31AA9CD4EEFC50" box="[1131,1241,981,1007]" count="3" pageId="3" pageNumber="194" type="female">3 females</specimenCount>
, 1 sex unknown),
<quantity id="4CD59B07FF86386B8BCDAAB0D171FBAB" box="[151,326,1017,1044]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.861" metricValueMin="0.939" pageId="3" pageNumber="194" unit="mm" value="140.0" valueMax="186.1" valueMin="93.9">93.9186.1 mm</quantity>
SL, off
<location id="8EF26039FF86386B8AC4AAB3D1D0FBAB" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8AC4AAB3D1D0FBAB" box="[414,487,1018,1044]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8AAAAAB0D256FBAC" box="[496,609,1017,1043]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8931AAB3D299FBAB" box="[619,686,1018,1044]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, coll. by
<location id="8EF26039FF86386B8854AAB0D38AFBAC" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8854AAB0D38AFBAC" box="[782,957,1017,1043]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B889CAAB0D4BCFBAC" box="[966,1163,1017,1043]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8F98AAB0D5AAFBAC" box="[1218,1437,1017,1043]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
(no further collection data)
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8A8FAD54D4B5FB88" box="[469,1154,1053,1080]" collectingDate="2019-08" collectionCode="FAKU" country="Japan" elevation="650" location="Rausu" municipality="Houyu-maru" pageId="3" pageNumber="194" specimenCode="FAKU 147238" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8A8FAD54D24AFB88" box="[469,637,1053,1079]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147238</specimenCode>
, male,
<quantity id="4CD59B07FF86386B8994AD54D377FB87" box="[718,832,1053,1080]" metricMagnitude="-1" metricUnit="m" metricValue="1.607" pageId="3" pageNumber="194" unit="mm" value="160.7">160.7 mm</quantity>
SL, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8FD4AD54D16CFBC0" collectionCode="FAKU" country="Japan" elevation="645" latitude="44.024" location="Rausu" longLatPrecision="68" longitude="145.308" pageId="3" pageNumber="194" specimenCode="FAKU 147910-147913, 147915, 148026" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8FD4AD54D5AFFB88" box="[1166,1432,1053,1079]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147910147913</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8BCDAD08D0D9FBE3" box="[151,238,1089,1116]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">147915</specimenCode>
,
<specimenCode id="DB8B9E99FF86386B8BA0AD08D166FBE4" box="[250,337,1089,1115]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">148026</specimenCode>
,
<specimenCount id="9D2BFD6BFF86386B8A07AD08D183FBE4" box="[349,436,1089,1115]" count="3" pageId="3" pageNumber="194" type="male">3 males</specimenCount>
and
<specimenCount id="9D2BFD6BFF86386B8AB6AD08D26BFBE4" box="[492,604,1089,1115]" count="3" pageId="3" pageNumber="194" type="female">3 females</specimenCount>
,
<quantity id="4CD59B07FF86386B8932AD08D31FFBE3" box="[616,808,1089,1116]" metricMagnitude="-1" metricUnit="m" metricValue="1.6634999999999998" metricValueMax="1.7759999999999998" metricValueMin="1.551" pageId="3" pageNumber="194" unit="mm" value="166.35" valueMax="177.6" valueMin="155.1">155.1177.6 mm</quantity>
SL,
<geoCoordinate id="EE195025FF86386B8805AD08D3FBFBE4" box="[863,972,1089,1115]" degrees="44.024" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="44.024">44.024ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF86386B8882AD08D478FBE3" box="[984,1103,1089,1116]" degrees="145.308" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="145.308">145.308ºE</geoCoordinate>
, off
<location id="8EF26039FF86386B8FD9AD0BD4FBFBE3" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8FD9AD0BD4FBFBE3" box="[1155,1228,1090,1116]" country="Japan" latitude="44.024" longLatPrecision="68" longitude="145.308" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8F82AD08D57EFBE4" box="[1240,1353,1089,1115]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8E0FAD0BD5AFFBE3" box="[1365,1432,1090,1116]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8BCDAD2CD16CFBC0" box="[151,347,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">
<quantity id="4CD59B07FF86386B8BCDAD2CD123FB3F" box="[151,276,1125,1152]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">540750 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8A26AD2CD5AFFBC0" box="[380,1432,1125,1152]" collectingDate="2020-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8A26AD2CD1B8FBC0" box="[380,399,1125,1151]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8ACEAD2FD214FBC0" box="[404,547,1126,1151]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8ACEAD2FD214FBC0" box="[404,547,1126,1151]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8974AD2CD291FBC0" box="[558,678,1125,1151]" pageId="3" pageNumber="194" value="2020-08">
<collectingDate id="EFD7E9CAFF86386B8974AD2CD291FBC0" box="[558,678,1125,1151]" pageId="3" pageNumber="194" value="2020-08">Aug. 2020</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B885DAD2CD381FBC0" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B885DAD2CD381FBC0" box="[775,950,1125,1151]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B88E5AD2CD4B4FBC0" box="[959,1155,1125,1151]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8FE0AD2CD5AFFBC0" box="[1210,1432,1125,1151]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<collectionCode id="ED3CAE27FF86386B8BCDADC0D0D6FB1C" box="[151,225,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8BB3ADC0D189FB1B" box="[233,446,1161,1188]" metricMagnitude="5" metricUnit="m" metricValue="1.4887219919999999" pageId="3" pageNumber="194" unit="mm" value="1.488721992E8">148872, 199.2 mm</quantity>
SL, male,
<materialsCitation id="3B453CBFFF86386B8960ADC3D571FB1C" box="[570,1350,1161,1188]" collectionCode="HUMZ, FAKU" country="Japan" pageId="3" pageNumber="194" specimenCode="HUMZ 235309, FAKU 148880" specimenCount="2" specimenCount-female="2" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8960ADC3D2DFFB1B" box="[570,744,1161,1188]" collectionCode="HUMZ" country="Japan" httpUri="http://grbio.org/cool/8gr0-wfn9" name="Hokkaido University, Laboratory of Marine Zoology" pageId="3" pageNumber="194">HUMZ 235309</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B8878ADC0D3FBFB1C" box="[802,972,1161,1187]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 148880</specimenCode>
),
<specimenCount id="9D2BFD6BFF86386B88BAADC0D467FB1C" box="[992,1104,1161,1187]" count="2" pageId="3" pageNumber="194" type="female">2 females</specimenCount>
, 169.0
<quantity id="4CD59B07FF86386B8FF0ADC0D52CFB1B" box="[1194,1307,1161,1188]" metricMagnitude="-1" metricUnit="m" metricValue="1.918" pageId="3" pageNumber="194" unit="mm" value="191.8">191.8 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation id="3B453CBFFF86386B8E08ADC0D430FB78" collectionCode="FAKU" collectorName="Rausu" country="Japan" elevation="650" location="Hokkaido" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8E08ADC0D5ABFB1C" box="[1362,1436,1161,1187]" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU</collectionCode>
<quantity id="4CD59B07FF86386B8BCDADE4D158FB77" box="[151,367,1197,1224]" metricMagnitude="5" metricUnit="m" metricValue="1.489801734" pageId="3" pageNumber="194" unit="mm" value="1.489801734E8">148980, 173.4 mm</quantity>
SL, male, off
<collectorName id="26D85334FF86386B8942ADE7D256FB77" box="[536,609,1198,1224]" pageId="3" pageNumber="194">Rausu</collectorName>
,
<collectingRegion id="49E9F800FF86386B8934ADE4D2E8FB78" box="[622,735,1197,1223]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B89B6ADE7D318FB77" box="[748,815,1198,1224]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8866ADE4D430FB78" box="[828,1031,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">
<quantity id="4CD59B07FF86386B8866ADE4D38AFB77" box="[828,957,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="3" pageNumber="194" unit="m" value="650.0" valueMax="800.0" valueMin="500.0">500800 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8F76ADE4D392FB54" collectingDate="2021-07" collectingDateMax="2021-08-31" collectingDateMin="2021-07" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8F76ADE4D408FB78" box="[1068,1087,1197,1223]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8F12ADE7D4E0FB78" box="[1096,1239,1198,1223]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B8F12ADE7D4E0FB78" box="[1096,1239,1198,1223]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B8FBDADE4D5AFFB78" box="[1255,1432,1197,1223]" pageId="3" pageNumber="194" value="2021-07" valueMax="2021-08-31" valueMin="2021-07-00">
<collectingDate id="EFD7E9CAFF86386B8FBDADE4D5AFFB78" box="[1255,1432,1197,1223]" pageId="3" pageNumber="194" value="2021-07" valueMax="2021-08-31" valueMin="2021-07-00">Jul.Aug. 2021</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8BA2AD98D19BFB54" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8BA2AD98D19BFB54" box="[248,428,1233,1259]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8AE3AD98D2B4FB54" box="[441,643,1233,1259]" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8999AD98D392FB54" box="[707,933,1233,1259]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B88E9AD98D219FAB0" collectingDate="2019-08" collectionCode="OS, FAKU" country="Japan" elevation="650" location="Rausu" municipality="Houyu-maru" pageId="3" pageNumber="194" specimenCode="OS 26590, FAKU 147239" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B88E9AD98D410FB53" box="[947,1063,1233,1260]" collectionCode="OS" country="USA" name="Oregon State University" pageId="3" pageNumber="194">OS 26590</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B8F3FAD98D526FB54" box="[1125,1297,1233,1259]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147239</specimenCode>
),
<quantity id="4CD59B07FF86386B8E72AD98D5AAFB53" box="[1320,1437,1233,1260]" metricMagnitude="-1" metricUnit="m" metricValue="1.538" pageId="3" pageNumber="194" unit="mm" value="153.8">153.8 mm</quantity>
SL, female, collected with holotype
</materialsCitation>
;
<materialsCitation id="3B453CBFFF86386B8963ADBCD281FA8C" collectionCode="OS, FAKU" country="Japan" elevation="645" latitude="44.024" location="Rausu" longLatPrecision="68" longitude="145.308" pageId="3" pageNumber="194" specimenCode="OS 26591, FAKU 147914" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<specimenCode id="DB8B9E99FF86386B8963ADBCD29CFAAF" box="[569,683,1269,1296]" collectionCode="OS" country="USA" name="Oregon State University" pageId="3" pageNumber="194">OS 26591</specimenCode>
(ex.
<specimenCode id="DB8B9E99FF86386B89BFADBCD3B8FAB0" box="[741,911,1269,1295]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147914</specimenCode>
),
<quantity id="4CD59B07FF86386B88F9ADBCD421FAAF" box="[931,1046,1269,1296]" metricMagnitude="-1" metricUnit="m" metricValue="1.2429999999999999" pageId="3" pageNumber="194" unit="mm" value="124.3">124.3 mm</quantity>
SL, female,
<geoCoordinate id="EE195025FF86386B8FF2ADBCD522FAB0" box="[1192,1301,1269,1295]" degrees="44.024" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="44.024">44.024ºN</geoCoordinate>
,
<geoCoordinate id="EE195025FF86386B8E7AADBCD5AFFAAF" box="[1312,1432,1269,1296]" degrees="145.308" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="145.308">145.308ºE</geoCoordinate>
, off
<location id="8EF26039FF86386B8B98AC53D13CFA8B" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8B98AC53D13CFA8B" box="[194,267,1306,1332]" country="Japan" latitude="44.024" longLatPrecision="68" longitude="145.308" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B8A43AC50D1BDFA8C" box="[281,394,1305,1331]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B8AC2AC53D1ECFA8B" box="[408,475,1306,1332]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
,
<elevation id="0000D1D1FF86386B8AB3AC50D281FA8C" box="[489,694,1305,1332]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">
<quantity id="4CD59B07FF86386B8AB3AC50D25CFA8B" box="[489,619,1305,1332]" metricMagnitude="2" metricUnit="m" metricValue="6.45" metricValueMax="7.5" metricValueMin="5.4" pageId="3" pageNumber="194" unit="m" value="645.0" valueMax="750.0" valueMin="540.0">540750 m</quantity>
depth
</elevation>
</materialsCitation>
, F/
<materialsCitation id="3B453CBFFF86386B8986AC50D22BFAE8" collectingDate="2020-08" collectionCode="V" collectorName="Houyu-maru" country="Japan" county="Shigetada Fujimoto" location="Koji Matsuzaki" municipality="Shigeki Fujimoto" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
<collectionCode id="ED3CAE27FF86386B8986AC50D2D8FA8C" box="[732,751,1305,1331]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B89A0AC53D3BEFA8C" box="[762,905,1306,1331]" italics="true" pageId="3" pageNumber="194">
<collectorName id="26D85334FF86386B89A0AC53D3BEFA8C" box="[762,905,1306,1331]" pageId="3" pageNumber="194">Houyu-maru</collectorName>
</emphasis>
,
<date id="FF931022FF86386B88C3AC50D422FA8C" box="[921,1045,1305,1331]" pageId="3" pageNumber="194" value="2020-08">
<collectingDate id="EFD7E9CAFF86386B88C3AC50D422FA8C" box="[921,1045,1305,1331]" pageId="3" pageNumber="194" value="2020-08">Aug. 2020</collectingDate>
</date>
, coll. by
<location id="8EF26039FF86386B8FDFAC50D50EFA8C" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B8FDFAC50D50EFA8C" box="[1157,1337,1305,1331]" country="Japan" county="Shigetada Fujimoto" municipality="Shigeki Fujimoto" name="Koji Matsuzaki" pageId="3" pageNumber="194" stateProvince="Hokkaido">Koji Matsuzaki</location>
,
<collectingMunicipality id="6BF6AC98FF86386B8E1DAC50D136FAE8" pageId="3" pageNumber="194">Shigeki Fujimoto</collectingMunicipality>
, and
<collectingCounty id="62F34E6EFF86386B8A61AC74D22BFAE8" box="[315,540,1341,1367]" pageId="3" pageNumber="194">Shigetada Fujimoto</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9236E2FF86386B8B9DAC28D430FA20" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DAC28D1B7FAC4" bold="true" box="[199,384,1377,1403]" pageId="3" pageNumber="194">Other material.</emphasis>
<materialsCitation id="3B453CBFFF86386B8ADCAC28D4F5FAC4" box="[390,1218,1377,1404]" collectionCode="FAKU" country="Japan" elevation="700" location="Rausu" pageId="3" pageNumber="194" specimenCode="FAKU 147740" specimenCount="1" stateProvince="Hokkaido">
<specimenCode id="DB8B9E99FF86386B8ADCAC28D219FAC4" box="[390,558,1377,1403]" collectionCode="FAKU" country="Japan" httpUri="http://grbio.org/cool/ist4-jfcv" name="Kyoto University" pageId="3" pageNumber="194">FAKU 147740</specimenCode>
, C&amp;S, off
<location id="8EF26039FF86386B89F9AC2BD2DBFAC3" LSID="urn:lsid:plazi:treatment:038487F4FF86386E8BCDAB6CD1D6FAC5:8EF26039FF86386B89F9AC2BD2DBFAC3" box="[675,748,1378,1404]" country="Japan" name="Rausu" pageId="3" pageNumber="194" stateProvince="Hokkaido">Rausu</location>
,
<collectingRegion id="49E9F800FF86386B89ACAC28D350FAC4" box="[758,871,1377,1403]" country="Japan" name="Hokkaido" pageId="3" pageNumber="194">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF86386B882BAC2BD383FAC3" box="[881,948,1378,1404]" name="Japan" pageId="3" pageNumber="194">Japan</collectingCountry>
, ca.
<elevation id="0000D1D1FF86386B88BEAC28D49EFAC4" box="[996,1193,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="194" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">
<quantity id="4CD59B07FF86386B88BEAC28D455FAC3" box="[996,1122,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="194" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600800 m</quantity>
depth
</elevation>
, F
</materialsCitation>
/
<materialsCitation id="3B453CBFFF86386B8F91AC28D434FA20" collectingDate="2020-07-20" collectionCode="V" country="Japan" pageId="3" pageNumber="194" specimenCount="1" stateProvince="Hokkaido">
<collectionCode id="ED3CAE27FF86386B8F91AC28D4E9FAC4" box="[1227,1246,1377,1403]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="194" type="Museum">V</collectionCode>
<emphasis id="B959EAF0FF86386B8FBEAC2BD544FAC4" box="[1252,1395,1378,1403]" italics="true" pageId="3" pageNumber="194">Houyu-maru</emphasis>
,
<date id="FF931022FF86386B8EDAAC28D136FA20" pageId="3" pageNumber="194" value="2020-07-20">
<collectingDate id="EFD7E9CAFF86386B8EDAAC28D136FA20" pageId="3" pageNumber="194" value="2020-07-20">20 Jul. 2020</collectingDate>
</date>
, coll. by Koji Matsuzaki, Shigeki Fujimoto, and Shigetada Fujimoto
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386B8B9DACE0D3E1F9EC" pageId="3" pageNumber="194" type="diagnosis">
<paragraph id="8B9236E2FF86386B8B9DACE0D3E1F9EC" blockId="3.[151,1437,729,2016]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DACE0D175FA7C" bold="true" box="[199,322,1449,1475]" pageId="3" pageNumber="194">Diagnosis.</emphasis>
A species of
<taxonomicName id="4C2D4D61FF86386B8ABAACE3D25DFA7B" authorityName="Kroyer" authorityYear="1862" box="[480,618,1450,1476]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="3" pageNumber="194" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF86386B8ABAACE3D25DFA7B" box="[480,618,1450,1476]" italics="true" pageId="3" pageNumber="194">Careproctus</emphasis>
</taxonomicName>
with the following combination of characters: vertebrae 5558; dorsal-fin rays 5053; anal-fin rays 4548; pectoral-fin rays 2734; pectoral fin with shallow notch, lower lobe short, extending slightly posterior to anus; pelvic disk rounded, greatly reduced; cephalic pores 2-6-7-2, chin pores paired in separate pits; gill slit moderately large, extending ventrally to pectoral-fin rays 410; teeth simple or slightly shouldered; anus located below mid-orbit; peritoneum and stomach black.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF86386D8B9DAF14D145F960" lastPageId="5" lastPageNumber="196" pageId="3" pageNumber="194" type="description">
<paragraph id="8B9236E2FF86386B8B9DAF14D5B0F9C8" blockId="3.[151,1437,729,2016]" box="[199,1415,1629,1656]" pageId="3" pageNumber="194">
<emphasis id="B959EAF0FF86386B8B9DAF14D16EF9C8" bold="true" box="[199,345,1629,1655]" pageId="3" pageNumber="194">Description.</emphasis>
Measurements shown in
<tableCitation id="C6AF0359FF86386B892CAF14D2FDF9C8" box="[630,714,1629,1655]" captionStart="TABLE 1" captionStartId="7.[151,239,404,428]" captionText="TABLE 1. Counts and measurements of Careproctus barbatulus sp. nov., C. longibarbatus sp. nov., and C. spinulosus sp. nov." pageId="3" pageNumber="194">Table 1</tableCitation>
.
<typeStatus id="54968840FF86386B898DAF17D30EF9C7" box="[727,825,1630,1656]" pageId="3" pageNumber="194" type="paratype">Paratype</typeStatus>
data given in parentheses if different from
<typeStatus id="54968840FF86386B8E44AF14D5B3F9C8" box="[1310,1412,1629,1655]" pageId="3" pageNumber="194" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B9236E2FF86386C8B9DAFC8D2DCFEA2" blockId="3.[151,1437,729,2016]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="195" pageId="3" pageNumber="194">
Body moderately deep, compressed, tapering posteriorly (
<figureCitation id="13162A67FF86386B883CAFC8D39CF924" box="[870,939,1665,1691]" captionStart="FIGURE 1" captionStartId="4.[151,250,1585,1609]" captionTargetBox="[209,1378,306,1558]" captionTargetId="figure-82@4.[209,1378,306,1558]" captionTargetPageId="4" captionText="FIGURE 1. Careproctus barbatulus sp. nov., FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="3" pageNumber="194">Fig. 1</figureCitation>
). Skin relatively thick, covered by cactus-like prickles; gelatinous tissue under skin moderately developed. Head large, moderately compressed, dorsal profile rounded from nape to snout. Snout rounded, not protruding anterior to mouth. Mouth moderately large, terminal, slightly oblique (or horizontal); maxilla extending to mid orbit; oral cleft extending to anterior rim of orbit. Premaxillary teeth simple or weakly shouldered, arranged in oblique rows of 10 (610) teeth per row forming broad band; inner teeth larger, recurved. Mandibular teeth simple or weakly shouldered, arranged in oblique rows of 9 (49) teeth per row forming broad band; inner teeth larger, recurved. Narrow diastema at symphysis of upper and lower jaws. Orbit of moderate size, rounded; pupil rounded. Nostril single, with short tube, level with mid-orbit. Cephalic sensory pores of moderate size, rims thickened: nasal pores 2, maxillary pores 6, preoperculomandibular pores 7, suprabranchial pores 2; cephalic pore pattern 2-6-7-2. Chin pores paired in separate pits, well separated. Coronal pore absent. Gill slit moderately large, upper margin level with mid-orbit, extending ventrally to pectoral-fin ray 5 (410). Gill rakers 79, short, with tiny spines on tips (not counted in
<typeStatus id="54968840FF81386C8F68A9F2D4AEFF6A" box="[1074,1177,187,213]" pageId="4" pageNumber="195" type="holotype">holotype</typeStatus>
). Opercular flap blunt, pointed posteriorly; tip level with ventral rim of orbit. Pyloric caeca 15 (1422), thick, moderately long, covering half of stomach, on center-left side of visceral cavity.
</paragraph>
<caption id="DF52666AFF81386C8BCDAF78D3E6F92E" ID-DOI="http://doi.org/10.5281/zenodo.13219212" ID-Zenodo-Dep="13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="4" pageNumber="195" startId="4.[151,250,1585,1609]" targetBox="[209,1378,306,1558]" targetPageId="4" targetType="figure">
<paragraph id="8B9236E2FF81386C8BCDAF78D3E6F92E" blockId="4.[151,1437,1585,1681]" pageId="4" pageNumber="195">
<emphasis id="B959EAF0FF81386C8BCDAF78D123F9F6" bold="true" box="[151,276,1585,1609]" pageId="4" pageNumber="195">FIGURE 1.</emphasis>
<taxonomicName id="4C2D4D61FF81386C8A46AF78D239F9F6" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[284,526,1585,1609]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF81386C8A46AF78D239F9F6" box="[284,526,1585,1609]" italics="true" pageId="4" pageNumber="195">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF81386C894CAF78D25FF9F6" bold="true" box="[534,616,1585,1609]" pageId="4" pageNumber="195">
<taxonomicNameLabel id="A26A578BFF81386C894CAF78D25FF9F6" box="[534,616,1585,1609]" pageId="4" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively.
</paragraph>
</caption>
<paragraph id="8B9236E2FF81386C8B9DAFAAD38BF846" blockId="4.[151,1437,1763,2042]" pageId="4" pageNumber="195">
Vertebrae 57 (5558), precaudal 10 (910) and caudal 47 (4649). Pleural ribs minute, thin, in 1 pair (1 or 2 pairs) on abdominal vertebra 9 (810). Dorsal-fin rays 51 (5053), not exserted. Anteriormost dorsal-fin pterygiophore inserted between neural spines 3 and 4 (or 4 and 5), bearing a single ray (sometimes rayless). Anal-fin rays 46 (4548), not exserted. Two (1 or 2) anal-fin pterygiophores anterior to 1
<superScript id="7C589BAAFF81386C88BDAE06D3C5F8E2" attach="left" box="[999,1010,1871,1885]" fontSize="6" pageId="4" pageNumber="195">st</superScript>
haemal spine, each bearing a single ray. Membrane of posterior dorsal- and anal-fin rays attached about equidistant to caudal fin. Caudal fin slightly rounded. Principal caudal-fin rays 7 (79), 3 (34) and 4 (45) rays on upper and lower hypurals, respectively, dorsal procurrent rays 1 (01), ventral procurrent rays 0 (01). Hypural plates fused with terminal vertebral centrum and parhypural, upper and lower hypural plate separated by distinct slit.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DA9DED19BFE36" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="196">
Pectoral fin moderately notched, with 30 (2734) rays; upper lobe with 23 (2025) rays, extending just beyond (or slightly anterior to) level of anal fin origin; lower lobe short, with 7 (710) rays, 4
<superScript id="7C589BAAFF80386D8F1EA9F3D466FF77" attach="left" box="[1092,1105,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
(5
<superScript id="7C589BAAFF80386D8F37A9F3D44DFF77" attach="none" box="[1133,1146,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
7
<superScript id="7C589BAAFF80386D8FCCA9F3D494FF77" attach="none" box="[1174,1187,186,200]" fontSize="6" pageId="5" pageNumber="196">th</superScript>
) ray from ventralmost longest, extending slightly posterior to anus. Rays in notch more widely spaced than rays of lobes. Tip of pectoral-fin rays of upper lobe slightly protruding, rays of lower lobe strongly exserted. Uppermost pectoral-fin base level with mouth corner. Lowermost pectoral-fin base slightly anterior to (or below) anterior rim of orbit. Pelvic disk rounded, greatly reduced (sometimes modified as lobe). Anus much closer to pelvic disk than to anal fin origin, located below mid-orbit.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DA8DAD555FDDE" blockId="5.[151,1437,150,609]" pageId="5" pageNumber="196">
Pectoral girdle (based on FAKU 146634 and 147740:
<figureCitation id="13162A67FF80386D8843A8DAD3A7FE12" box="[793,912,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1159,1183]" captionTargetBox="[157,1432,673,1131]" captionTargetId="figure-242@5.[151,1436,665,1136]" captionTargetPageId="5" captionText="FIGURE 2. Pectoral girdle in five species of Careproctus with a reduced pelvic disk. Careproctus barbatulus sp. nov., FAKU 146634, paratype, 220.8 mm SL (A); Careproctus barbatulus sp. nov., FAKU 147740, non-type (B); Careproctus longibarbatus sp. nov., FAKU 148477, paratype, 223.0 mm SL (C); C. spinulosus sp. nov., FAKU 146632, paratype, 199.8 mm SL (D); C. mederi, FAKU 148478, non-type, 143.6 mm SL (E); C. gilberti, FAKU S4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage." figureDoi="http://doi.org/10.5281/zenodo.13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="5" pageNumber="196">Fig. 2A, B</figureCitation>
): Proximal pectoral radials 3 or 4; upper radials 1 or 2, hour-grass shaped with two notches, equally spaced, radials 2 or 3 (lowermost within upper radials) with one notch, radial 4 rounded squarish, widely separated from upper radials. Interradial fenestrae 2 or 3, extending between scapula and radials 13; fenestra 1 roundish, fenestrae 2 and 3 roundish or elliptical, sometimes with notch. Scapula broadly Y-shaped, with robust helve. Coracoid diamond-shaped or a roundish triangle, with short thin helve. Distal radials present at base of pectoral-fin rays 2 to 2127, absent from base of ventralmost 78 rays.
</paragraph>
<caption id="DF52666AFF80386D8BCDADCED16EFA90" ID-DOI="http://doi.org/10.5281/zenodo.13219214" ID-Zenodo-Dep="13219214" httpUri="https://zenodo.org/record/13219214/files/figure.png" pageId="5" pageNumber="196" startId="5.[151,250,1159,1183]" targetBox="[157,1432,673,1131]" targetPageId="5" targetType="figure">
<paragraph id="8B9236E2FF80386D8BCDADCED16EFA90" blockId="5.[151,1436,1159,1327]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8BCDADCED124FB20" bold="true" box="[151,275,1159,1183]" pageId="5" pageNumber="196">FIGURE 2.</emphasis>
Pectoral girdle in five species of
<taxonomicName id="4C2D4D61FF80386D8932ADC1D2D2FB20" authorityName="Kroyer" authorityYear="1862" box="[616,741,1160,1183]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF80386D8932ADC1D2D2FB20" box="[616,741,1160,1183]" italics="true" pageId="5" pageNumber="196">Careproctus</emphasis>
</taxonomicName>
with a reduced pelvic disk.
<taxonomicName id="4C2D4D61FF80386D8F59ADC1D4C2FB20" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[1027,1269,1159,1183]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8F59ADC1D4C2FB20" box="[1027,1269,1159,1183]" italics="true" pageId="5" pageNumber="196">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8FA1ADCED57BFB20" bold="true" box="[1275,1356,1159,1183]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8FA1ADCED57BFB20" box="[1275,1356,1159,1183]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 146634, paratype, 220.8 mm SL (A);
<taxonomicName id="4C2D4D61FF80386D8954ADE5D2C9FB7C" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[526,766,1195,1219]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8954ADE5D2C9FB7C" box="[526,766,1195,1219]" italics="true" pageId="5" pageNumber="196">Careproctus barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8859ADE2D364FB7C" bold="true" box="[771,851,1195,1219]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8859ADE2D364FB7C" box="[771,851,1195,1219]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 147740, non-type (B);
<taxonomicName id="4C2D4D61FF80386D8FD6ADE5D5ABFB7C" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[1164,1436,1195,1219]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="longibarbatus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8FD6ADE5D5ABFB7C" box="[1164,1436,1195,1219]" italics="true" pageId="5" pageNumber="196">Careproctus longibarbatus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8BCDAD86D0DDFB58" bold="true" box="[151,234,1231,1255]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8BCDAD86D0DDFB58" box="[151,234,1231,1255]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 148477, paratype, 223.0 mm SL (C);
<taxonomicName id="4C2D4D61FF80386D8989AD99D368FB58" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[723,863,1231,1255]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="spinulosus" status="sp. nov.">
<emphasis id="B959EAF0FF80386D8989AD99D2D3FB58" box="[723,740,1232,1255]" italics="true" pageId="5" pageNumber="196">C</emphasis>
.
<emphasis id="B959EAF0FF80386D89AEAD86D368FB58" box="[756,863,1231,1255]" italics="true" pageId="5" pageNumber="196">spinulosus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF80386D8832AD86D38CFB58" bold="true" box="[872,955,1231,1255]" pageId="5" pageNumber="196">
<taxonomicNameLabel id="A26A578BFF80386D8832AD86D38CFB58" box="[872,955,1231,1255]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, FAKU 146632, paratype, 199.8 mm SL (D);
<taxonomicName id="4C2D4D61FF80386D8BCDADBDD178FAB3" authority=", FAKU" authorityName="FAKU" box="[151,335,1267,1292]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF80386D8BCDADBDD0CAFAB4" box="[151,253,1267,1291]" italics="true" pageId="5" pageNumber="196">C. mederi</emphasis>
, FAKU
</taxonomicName>
148478, non-type, 143.6 mm SL (E);
<taxonomicName id="4C2D4D61FF80386D89BBADBDD384FAB4" authority=", FAKU S" authorityName="FAKU S" box="[737,947,1267,1292]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF80386D89BBADBDD37BFAB4" box="[737,844,1267,1291]" italics="true" pageId="5" pageNumber="196">C. gilberti</emphasis>
, FAKU S
</taxonomicName>
4869, non-type, 67.2 mm SL (F). Shaded areas represent cartilage.
</paragraph>
</caption>
<paragraph id="8B9236E2FF80386D8B9DACC8D158F994" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
Coloration (
<figureCitation id="13162A67FF80386D8A0AACC8D1A3FA23" box="[336,404,1409,1436]" captionStart="FIGURE 1" captionStartId="4.[151,250,1585,1609]" captionTargetBox="[209,1378,306,1558]" captionTargetId="figure-82@4.[209,1378,306,1558]" captionTargetPageId="4" captionText="FIGURE 1. Careproctus barbatulus sp. nov., FAKU 148026, paratype, 152.7 mm SL, (fresh) (A); FAKU 147240, holotype, 157.7 mm SL, (preserved) (B); ventral view of head region of FAKU 147240 (C), and FAKU 148026 (D) (both preserved). Dotted red line and arrows indicate position of anus and pelvic disk, respectively." figureDoi="http://doi.org/10.5281/zenodo.13219212" httpUri="https://zenodo.org/record/13219212/files/figure.png" pageId="5" pageNumber="196">Fig. 1</figureCitation>
): When fresh (based on FAKU 148026), head and body pale to dark reddish; tip of nostril dark brown; postorbital region and belly somewhat paler. Iris dark silvery. Dorsal, anal, and caudal fins pale to dark reddish; pectoral fin dark red, basal part paler. Pelvic disk white. When preserved, head and body white; upper and lower jaws mottled with black; nostril margined with black. Eyes black. Oral and branchial cavities pale; peritoneum and stomach black.
</paragraph>
<paragraph id="8B9236E2FF80386D8B9DAF7CD145F960" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8B9DAF7CD144F9F0" bold="true" box="[199,371,1589,1615]" pageId="5" pageNumber="196">Reproduction.</emphasis>
The
<typeStatus id="54968840FF80386D8AF4AF7CD227F9F0" box="[430,528,1589,1615]" pageId="5" pageNumber="196" type="holotype">holotype</typeStatus>
(FAKU 147240) was an immature female. FAKU 147911 (
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SL) was a spent female with yolked (
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diameter) and small immature (less than
<quantity id="4CD59B07FF80386D8F6EAF10D4AFF9CB" box="[1076,1176,1625,1652]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="196" unit="mm" value="0.5">0.50 mm</quantity>
diameter) eggs. FAKU 148880 (169.0 mm SL and
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SL) were also spent females with yolked (
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diameter and
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diameter, respectively) and small immature (less than
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diameter in both specimens) eggs. Yolked eggs were rounded.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF80386D8B9DAFA0D138F897" pageId="5" pageNumber="196" type="distribution">
<paragraph id="8B9236E2FF80386D8B9DAFA0D138F897" blockId="5.[151,1437,1409,1832]" pageId="5" pageNumber="196">
<emphasis id="B959EAF0FF80386D8B9DAFA0D156F8BC" bold="true" box="[199,353,1769,1795]" pageId="5" pageNumber="196">Distribution.</emphasis>
Known only from the southern Sea of Okhotsk, off Rausu,
<collectingRegion id="49E9F800FF80386D88B4AFA0D468F8BC" box="[1006,1119,1769,1795]" country="Japan" name="Hokkaido" pageId="5" pageNumber="196">Hokkaido</collectingRegion>
,
<collectingCountry id="F33A7672FF80386D8F33AFA3D49BF8BB" box="[1129,1196,1770,1796]" name="Japan" pageId="5" pageNumber="196">Japan</collectingCountry>
, at depths of
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(
<figureCitation id="13162A67FF80386D8BE6AE44D136F897" box="[188,257,1805,1832]" captionStart="FIGURE 3" captionStartId="6.[151,250,729,753]" captionTargetBox="[175,1410,188,699]" captionTargetId="figure-20@6.[171,1416,181,706]" captionTargetPageId="6" captionText="FIGURE 3. Distributional records of species of Careproctus with a reduced pelvic disk based on the present and previous studies (see text)." figureDoi="http://doi.org/10.5281/zenodo.13219216" httpUri="https://zenodo.org/record/13219216/files/figure.png" pageId="5" pageNumber="196">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF52666AFF83386E8BCDAB90D17EFCA9" ID-DOI="http://doi.org/10.5281/zenodo.13219216" ID-Zenodo-Dep="13219216" httpUri="https://zenodo.org/record/13219216/files/figure.png" pageId="6" pageNumber="197" startId="6.[151,250,729,753]" targetBox="[175,1410,188,699]" targetPageId="6" targetType="figure">
<paragraph id="8B9236E2FF83386E8BCDAB90D17EFCA9" blockId="6.[151,1436,729,790]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8BCDAB90D122FD4D" bold="true" box="[151,277,729,754]" pageId="6" pageNumber="197">FIGURE 3.</emphasis>
Distributional records of species of
<taxonomicName id="4C2D4D61FF83386E89CCAB93D324FD4E" authorityName="Kroyer" authorityYear="1862" box="[662,787,730,753]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF83386E89CCAB93D324FD4E" box="[662,787,730,753]" italics="true" pageId="6" pageNumber="197">Careproctus</emphasis>
</taxonomicName>
with a reduced pelvic disk based on the present and previous studies (see text).
</paragraph>
</caption>
<subSubSection id="C3376569FF83386E8B9DAA21D38FFC19" pageId="6" pageNumber="197" type="etymology">
<paragraph id="8B9236E2FF83386E8B9DAA21D38FFC19" blockId="6.[151,1436,872,1402]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8B9DAA21D179FC3D" bold="true" box="[199,334,872,898]" pageId="6" pageNumber="197">Etymology.</emphasis>
The specific name
<taxonomicName id="4C2D4D61FF83386E897CAA21D29BFC3D" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[550,684,872,898]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus">
<emphasis id="B959EAF0FF83386E8969AA21D29BFC3D" box="[563,684,872,898]" italics="true" pageId="6" pageNumber="197">barbatulus</emphasis>
</taxonomicName>
” derived from the diminutive form of Latin “barba” (barbel), refers to the short lower lobe of the pectoral fin, reminiscent of a short barbel.
</paragraph>
</subSubSection>
<subSubSection id="C3376569FF83386E8B9DAAF9D1D6FAC5" pageId="6" pageNumber="197" type="discussion">
<paragraph id="8B9236E2FF83386E8B9DAAF9D1D6FAC5" blockId="6.[151,1436,872,1402]" pageId="6" pageNumber="197">
<emphasis id="B959EAF0FF83386E8B9DAAF9D10CFC75" bold="true" box="[199,315,944,970]" pageId="6" pageNumber="197">Remarks.</emphasis>
Within the
<taxonomicName id="4C2D4D61FF83386E8A98AAF9D201FC75" box="[450,566,944,970]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="gilberti">
<emphasis id="B959EAF0FF83386E8A98AAF9D201FC75" box="[450,566,944,970]" italics="true" pageId="6" pageNumber="197">C. gilberti</emphasis>
</taxonomicName>
species group,
<taxonomicName id="4C2D4D61FF83386E89B2AAF9D3B4FC75" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[744,899,944,970]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus" status="sp. nov.">
<emphasis id="B959EAF0FF83386E89B2AAF9D3B4FC75" box="[744,899,944,970]" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
<emphasis id="B959EAF0FF83386E88D1AAF9D3D1FC75" bold="true" box="[907,998,944,970]" pageId="6" pageNumber="197">
<taxonomicNameLabel id="A26A578BFF83386E88D1AAF9D3D1FC75" box="[907,998,944,970]" pageId="6" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares some morphological characters with
<taxonomicName id="4C2D4D61FF83386E8B88AA9DD173FC51" box="[210,324,980,1006]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF83386E8B88AA9DD173FC51" box="[210,324,980,1006]" italics="true" pageId="6" pageNumber="197">C. mederi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D61FF83386E8A25AA9DD230FC51" authorityName="Gilbert" authorityYear="1896" box="[383,519,980,1006]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF83386E8A25AA9DD230FC51" box="[383,519,980,1006]" italics="true" pageId="6" pageNumber="197">C. ostentum</emphasis>
</taxonomicName>
, including a short lower lobe of the pectoral fin, extending slightly posterior to the anus, and relatively thick skin with strong prickles. The new species can be distinguished from the latter two species by the gill slit length (extending to pectoral-fin ray 410 vs.
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<taxonomicName id="4C2D4D61FF83386E8F52AD55D440FB89" box="[1032,1143,1052,1078]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF83386E8F52AD55D440FB89" box="[1032,1143,1052,1078]" italics="true" pageId="6" pageNumber="197">C. mederi</emphasis>
</taxonomicName>
and
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<taxonomicName id="4C2D4D61FF83386E8E56AD55D5A4FB89" authorityName="Gilbert" authorityYear="1896" box="[1292,1427,1052,1078]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="ostentum">
<emphasis id="B959EAF0FF83386E8E56AD55D5A4FB89" box="[1292,1427,1052,1078]" italics="true" pageId="6" pageNumber="197">C. ostentum</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFBC4B13FF83386E8BC5AD09D18EFBE5" author="Mecklenburg, C. W. &amp; Mecklenburg, T. A. &amp; Thorsteinson, L. K." box="[159,441,1088,1114]" pageId="6" pageNumber="197" refId="ref18345" refString="Mecklenburg, C. W., Mecklenburg, T. A. &amp; Thorsteinson, L. K. (2002) Fishes of Alaska. American Fisheries Society, Bethesda, Maryland, 1037 pp." type="book" year="2002">
Mecklenburg
<emphasis id="B959EAF0FF83386E8A67AD08D141FBE5" box="[317,374,1088,1114]" italics="true" pageId="6" pageNumber="197">et al.</emphasis>
2002
</bibRefCitation>
; this study) and coloration of the peritoneum (dark or black vs. pale with black dots). In addition,
<taxonomicName id="4C2D4D61FF83386E8A5AAD2DD1E8FBC1" box="[256,479,1124,1150]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="mederi">
<emphasis id="B959EAF0FF83386E8A5AAD2DD1E8FBC1" box="[256,479,1124,1150]" italics="true" pageId="6" pageNumber="197">Careproctus mederi</emphasis>
</taxonomicName>
differs from the new species by having the lower end of the pectoral girdle protruding anteriorly (vs. not protruding, see description below). Although
<taxonomicName id="4C2D4D61FF83386E882FADC1D5AFFB1D" authority="Gilbert &amp; Burke, 1912" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[885,1432,1160,1186]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="acanthodes">
<emphasis id="B959EAF0FF83386E882FADC1D4BFFB1D" box="[885,1160,1160,1186]" italics="true" pageId="6" pageNumber="197">Careproctus acanthodes</emphasis>
<bibRefCitation id="EFBC4B13FF83386E8FCBADC1D5AFFB1D" author="Gilbert, C. H. &amp; Burke, C. V." box="[1169,1432,1160,1186]" pageId="6" pageNumber="197" pagination="31 - 96" refId="ref17636" refString="Gilbert, C. H. &amp; Burke, C. V. (1912) Fishes from Bering Sea and Kamchatka. Bulletin of the Bureau of Fisheries, 30, 31 - 96." type="journal article" year="1912">Gilbert &amp; Burke, 1912</bibRefCitation>
</taxonomicName>
, a member of the
<taxonomicName id="4C2D4D61FF83386E8A39ADE5D1DBFB79" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[355,492,1196,1222]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="rastrinus">
<emphasis id="B959EAF0FF83386E8A39ADE5D1DBFB79" box="[355,492,1196,1222]" italics="true" pageId="6" pageNumber="197">C. rastrinus</emphasis>
</taxonomicName>
species group, has the shortest pectoral fin lower lobe among the species group, the present new species clearly differs from
<taxonomicName id="4C2D4D61FF83386E89D5AD99D306FB55" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[655,817,1232,1258]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="acanthodes">
<emphasis id="B959EAF0FF83386E89D5AD99D306FB55" box="[655,817,1232,1258]" italics="true" pageId="6" pageNumber="197">C. acanthodes</emphasis>
</taxonomicName>
in having a smaller pelvic disk (0.40.9% SL vs. 4.2 7.6% SL) (
<bibRefCitation id="EFBC4B13FF83386E8A48ADBDD1F7FAB0" author="Orr, J. W. &amp; Kai, Y. &amp; Nakabo, T." box="[274,448,1268,1295]" pageId="6" pageNumber="197" pagination="301 - 348" refId="ref18697" refString="Orr, J. W., Kai, Y. &amp; Nakabo, T. (2015) Snailfishes of the Careproctus rastrinus complex (Liparidae): redescriptions of seven species in the North Pacific Ocean region, with the description of a new species from the Beaufort Sea. Zootaxa, 4018 (3), 301 - 348. https: // doi. org / 10.11646 / zootaxa. 4018.3.1" type="journal article" year="2015">
Orr
<emphasis id="B959EAF0FF83386E8A19ADBCD143FAB1" box="[323,372,1268,1294]" italics="true" pageId="6" pageNumber="197">et al</emphasis>
. 2015
</bibRefCitation>
). Among the species of
<taxonomicName id="4C2D4D61FF83386E8994ADBDD368FAB1" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[718,863,1268,1294]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="genus">
<emphasis id="B959EAF0FF83386E8994ADBDD368FAB1" box="[718,863,1268,1294]" italics="true" pageId="6" pageNumber="197">Elassodiscus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2D4D61FF83386E8837ADBDD5ABFAB1" authority="Gilbert &amp; Burke, 1912" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[877,1436,1268,1294]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="tremebundus">
<emphasis id="B959EAF0FF83386E8837ADBDD4A1FAB1" box="[877,1174,1268,1294]" italics="true" pageId="6" pageNumber="197">Elassodiscus tremebundus</emphasis>
<bibRefCitation id="EFBC4B13FF83386E8FC7ADBDD5ABFAB1" author="Gilbert, C. H. &amp; Burke, C. V." box="[1181,1436,1268,1294]" pageId="6" pageNumber="197" pagination="31 - 96" refId="ref17636" refString="Gilbert, C. H. &amp; Burke, C. V. (1912) Fishes from Bering Sea and Kamchatka. Bulletin of the Bureau of Fisheries, 30, 31 - 96." type="journal article" year="1912">Gilbert &amp; Burke, 1912</bibRefCitation>
</taxonomicName>
also has a relatively short pectoral fin lower lobe. However, together with the diagnostic characters of the genus,
<taxonomicName id="4C2D4D61FF83386E8BCDAC74D17EFAE9" authorityName="Gilbert &amp; Burke" authorityYear="1912" box="[151,329,1340,1366]" family="Liparidae" genus="Elassodiscus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="tremebundus">
<emphasis id="B959EAF0FF83386E8BCDAC74D17EFAE9" box="[151,329,1340,1366]" italics="true" pageId="6" pageNumber="197">E. tremebundus</emphasis>
</taxonomicName>
is further distinguishable from
<taxonomicName id="4C2D4D61FF83386E89EFAC75D366FAE9" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" box="[693,849,1340,1366]" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus">
<emphasis id="B959EAF0FF83386E89EFAC75D366FAE9" box="[693,849,1340,1366]" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
in having a single suprabranchial pore (vs.
<quantity id="4CD59B07FF83386E8E16AC75D54EFAE9" box="[1356,1401,1340,1366]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="197" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C2D4D61FF83386E8ED9AC75D127FAC5" authorityName="Kai &amp; Matsuzaki &amp; Mori &amp; Pitruk &amp; Misawa &amp; Tashiro" authorityYear="2024" family="Liparidae" genus="Careproctus" kingdom="Animalia" order="Scorpaeniformes" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="barbatulus">
<emphasis id="B959EAF0FF83386E8ED9AC75D127FAC5" italics="true" pageId="6" pageNumber="197">C. barbatulus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFBC4B13FF83386E8A73AC29D1E1FAC5" author="Kai, Y. &amp; Matsuzaki, K. &amp; Orr, J. W. &amp; Mori, T. &amp; Kamiunten, M." box="[297,470,1376,1402]" pageId="6" pageNumber="197" pagination="55 - 66" refId="ref17899" refString="Kai, Y., Matsuzaki, K., Orr, J. W., Mori, T. &amp; Kamiunten, M. (2020) A new species of Elassodiscus (Cottoidei: Liparidae) from the North Pacific with an emended diagnosis of the genus. Ichthyological Research, 68, 55 - 66. https: // doi. org / 10.1007 / s 10228 - 020 - 00764 - 4" type="journal article" year="2020">
Kai
<emphasis id="B959EAF0FF83386E8A00AC28D1BCFAC5" box="[346,395,1376,1402]" italics="true" pageId="6" pageNumber="197">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>