170 lines
19 KiB
XML
170 lines
19 KiB
XML
<document id="8BE8F4AEB67440C0B3FA1691AAEA982B" ID-CLB-Dataset="46615" ID-DOI="10.11646/zootaxa.3755.2.1" ID-GBIF-Dataset="e02bbd39-b7ee-4ddf-8109-3407dd6d2ea2" ID-ISSN="1175-5326" ID-Zenodo-Dep="250959" ID-ZooBank="1872ECAB-3C5C-4D76-93A0-A8626F75B96E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460372737068" checkinUser="plazi" docAuthor="Helwerda, Renate Ariane, Wesselingh, Frank Pieter & Williams, Suzanne T." docDate="2014" docId="490087B3FFB36D2EFF1FFDA85B6DF874" docLanguage="en" docName="zt03755p135.pdf" docOrigin="Zootaxa 3755 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calliotropis arenosa" docType="treatment" docVersion="10" lastPageNumber="115" masterDocId="B539FFCBFFBD6D20FF88FFB45A06FFFF" masterDocTitle="On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species" masterLastPageNumber="135" masterPageNumber="101" pageNumber="115" updateTime="1698295924955" updateUser="ExternalLinkService">
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<mods:mods id="ABFA80F016F0ABCEDFABF49118819DC4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="19CE8F21F67D8B2866DEA6ED31908451">
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<mods:title id="90388B9657F1D584D502F8205A41E0DD">On some Vetigastropoda (Mollusca, Gastropoda) from the Plio-Pleistocene of the Philippines with descriptions of three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="FC96951586A99002FDFD433EA5E28CD8" type="personal">
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<mods:role id="8B48B8B687BBC66572EFB71A98FC7830">
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<mods:roleTerm id="2E63429F9688C96920E18331C095DBD2">Author</mods:roleTerm>
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<mods:namePart id="15659A77F00CA35EE3F0C045EE9F5CCD">Helwerda, Renate Ariane</mods:namePart>
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</mods:name>
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<mods:name id="CE1B6A9273A4D55CEE9772D822E5B6AA" type="personal">
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<mods:roleTerm id="448EB3B75F681AD65A8C595C6AF33FE6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="59002A7B19F046B3FD15B0B6600EBAC7">Wesselingh, Frank Pieter</mods:namePart>
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</mods:name>
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<mods:name id="4573E755D6E72BD3CE8EE13D655F5FFF" type="personal">
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<mods:role id="40ED451DF6CA13F8B320130B9A399C18">
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<mods:roleTerm id="493B29FDB1AAE52F7ED50B191111E66D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFA8E4EE2986AB30EB6D45752CDC30E1">Williams, Suzanne T.</mods:namePart>
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<mods:typeOfResource id="5CB23790BC658D6B1D2793E96A3833C9">text</mods:typeOfResource>
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<mods:titleInfo id="637BFB94B4A47360D80CB7CF317AFE1B">
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<mods:title id="489E3E83BE67B521B33A96615202F7F9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2A5C1EC7CA6FCDC4D31A7A65245C004C">
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<mods:date id="60B96855012B54A472C92F264761F5BA">2014</mods:date>
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<mods:detail id="E65F62F90CDEDCA58553DD8C5BDFA5B6" type="volume">
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<mods:number id="20E275651D7D2AD1FCB4E467556B293D">3755</mods:number>
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</mods:detail>
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<mods:detail id="FDC3E62666A18DBCAA95AFC1C64902E2" type="issue">
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<mods:number id="1993CDCCFD6776462BD745A040F646E6">2</mods:number>
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<mods:start id="CB4A004CA7B4F6EB63C9A7B23D4584AA">101</mods:start>
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<mods:end id="B05DDBD197C4A815EC2B5CFD98913CBF">135</mods:end>
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<mods:classification id="82F3B8F5AA3B99E21AE31564ADBD3A05">journal article</mods:classification>
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<mods:identifier id="2C610CE5F2AE14AEA4A4524DB07C9AC8" type="CLB-Dataset">46615</mods:identifier>
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<mods:identifier id="5D9FEE8FED9F43DE2CD69EE2F443A08D" type="DOI">10.11646/zootaxa.3755.2.1</mods:identifier>
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<mods:identifier id="AC1EE33624BF971540D9BC84DD6D50AF" type="GBIF-Dataset">e02bbd39-b7ee-4ddf-8109-3407dd6d2ea2</mods:identifier>
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<treatment id="490087B3FFB36D2EFF1FFDA85B6DF874" ID-DOI="http://doi.org/10.5281/zenodo.5671281" ID-GBIF-Taxon="119412421" ID-Zenodo-Dep="5671281" LSID="urn:lsid:plazi:treatment:490087B3FFB36D2EFF1FFDA85B6DF874" httpUri="http://treatment.plazi.org/id/490087B3FFB36D2EFF1FFDA85B6DF874" lastPageNumber="115" pageId="14" pageNumber="115">
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<subSubSection id="89B3652EFFB36D2EFF1FFDA85B4DFDA8" pageId="14" pageNumber="115" type="nomenclature">
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<paragraph id="C11636A5FFB36D2EFF1FFDA8580DFDC9" blockId="14.[151,523,540,599]" box="[151,523,540,566]" pageId="14" pageNumber="115">
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<heading id="9A5E81C9FFB36D2EFF1FFDA8580DFDC9" bold="true" box="[151,523,540,566]" fontSize="11" level="1" pageId="14" pageNumber="115" reason="1">
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<emphasis id="F3DDEAB7FFB36D2EFF1FFDA8580DFDC9" bold="true" box="[151,523,540,566]" pageId="14" pageNumber="115">
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<taxonomicName id="06A94D26FFB36D2EFF1FFDA85B8AFDC9" ID-CoL="dc3d7180-fb3a-4bd0-bfd3-f33cc950bf1b" box="[151,396,540,566]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="arenosa">
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<emphasis id="F3DDEAB7FFB36D2EFF1FFDA85B8AFDC9" bold="true" box="[151,396,540,566]" italics="true" pageId="14" pageNumber="115">Calliotropis arenosa</emphasis>
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</taxonomicName>
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nov. spec.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C11636A5FFB36D2EFF1FFD8A5B4DFDA8" blockId="14.[151,523,540,599]" box="[151,331,574,599]" pageId="14" pageNumber="115">
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(
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<figureCitation id="59922A20FFB36D2EFF16FD8A5B45FDA8" box="[158,323,574,599]" captionStart="FIGURES 26 – 32" captionStartId="15.[151,264,1793,1815]" captionTargetBox="[259,1325,197,1767]" captionTargetId="figure@15.[257,1328,195,1768]" captionTargetPageId="15" captionText="FIGURES 26 – 32. Calliotropidae. 26. Calliotropis arenosa nov. spec. RGM 608.272 (holotype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 27. Calliotropis arenosa nov. spec. RGM 608.273 (paratype). Locality AndaDeVos. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 28. Calliotropis arenosa nov. spec. RGM 608.274 (paratype). Locality Anda 3. (a) rear view, (b) apertural view, (c) apical view. 29. Calliotropis arenosa nov. spec. RGM 608.275 (paratype). Locality AndaDeVos. SEM detail of protoconch. 30. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.281. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 31. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.282. Locality Anda 2. (a) rear view, (b) apertural view, (c) basal view, (d) apical view. 32. Calliotropis cf. pyramoeides Vilvens, 2007. RGM 608.283. Locality Anda 1. (a) SEM, side view, (b) SEM detail of protoconch." httpUri="https://zenodo.org/record/250966/files/figure.png" pageId="14" pageNumber="115">Figures 26–29</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF1FFD325B18FCF4" pageId="14" pageNumber="115" type="materials_examined">
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<paragraph id="C11636A5FFB36D2EFF1FFD325B62FD3C" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF1FFD325B42FD60" bold="true" box="[151,324,646,671]" pageId="14" pageNumber="115">
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<typeStatus id="1E128807FFB36D2EFF1FFD325AD4FD60" box="[151,210,646,671]" pageId="14" pageNumber="115">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="1E128807FFB36D2EFEC3FD325BABFD60" box="[331,429,646,671]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
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<collectionCode id="A7B8AE60FFB36D2EFE3BFD335BF5FD60" box="[435,499,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
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608.272.
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<typeStatus id="1E128807FFB36D2EFDEAFD3358C2FD60" box="[610,708,647,671]" pageId="14" pageNumber="115" type="paratype">Paratype</typeStatus>
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<collectionCode id="A7B8AE60FFB36D2EFD43FD33590DFD60" box="[715,779,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
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608.273 and
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<collectionCode id="A7B8AE60FFB36D2EFC2AFD3359E4FD60" box="[930,994,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
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608.275 (
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<typeStatus id="1E128807FFB36D2EFBD9FD335E85FD60" box="[1105,1155,647,671]" pageId="14" pageNumber="115">type</typeStatus>
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locality);
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<typeStatus id="1E128807FFB36D2EFB7DFD335F53FD60" box="[1269,1365,647,671]" pageId="14" pageNumber="115" type="paratype">paratype</typeStatus>
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<collectionCode id="A7B8AE60FFB36D2EFAD4FD335F9AFD60" box="[1372,1436,647,671]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="115">RGM</collectionCode>
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608.274 (Anda 3).
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</paragraph>
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<paragraph id="C11636A5FFB36D2EFF4FFD7A5B18FCF4" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFD7A5B76FD18" bold="true" box="[199,368,718,743]" pageId="14" pageNumber="115">
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<typeStatus id="1E128807FFB36D2EFF4FFD7A5B04FD18" box="[199,258,718,743]" pageId="14" pageNumber="115">Type</typeStatus>
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locality.
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</emphasis>
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AndaDeVos, Bolinao province,
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<collectingCountry id="B9BE7635FFB36D2EFC8FFD7A598EFD18" box="[775,904,718,743]" name="Philippines" pageId="14" pageNumber="115">Philippines</collectingCountry>
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, Santa Cruz Formation, Pliocene or Early Pleistocene.
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF4FFCA259C7FCD0" box="[199,961,790,815]" pageId="14" pageNumber="115" type="etymology">
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<paragraph id="C11636A5FFB36D2EFF4FFCA259C7FCD0" blockId="14.[151,1436,646,1931]" box="[199,961,790,815]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFCA25BA2FCD0" bold="true" box="[199,420,790,815]" pageId="14" pageNumber="115">Derivatio nominis.</emphasis>
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Named after the sediment covering adult shells.
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF4FFC8E5EBCFCAC" box="[199,1210,826,851]" pageId="14" pageNumber="115" type="materials_examined">
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<paragraph id="C11636A5FFB36D2EFF4FFC8E5EBCFCAC" blockId="14.[151,1436,646,1931]" box="[199,1210,826,851]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFC8E5B87FCAC" bold="true" box="[199,385,826,851]" pageId="14" pageNumber="115">Other material.</emphasis>
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Anda 2 (1); Anda 5 (1); Anda 6 (49); AndaDeVos (102); AndaClif3 (18).
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF4FFCEA58F6FC41" pageId="14" pageNumber="115" type="diagnosis">
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<paragraph id="C11636A5FFB36D2EFF4FFCEA58F6FC41" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFCEA5B47FC88" bold="true" box="[199,321,862,887]" pageId="14" pageNumber="115">Diagnosis.</emphasis>
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Shell conical, covered in sediment, H
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<quantity id="06519B40FFB36D2EFD74FCEB5950FC89" box="[764,854,863,887]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="115" unit="mm" value="3.4">3.4 mm</quantity>
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,
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<geoCoordinate id="A49D5062FFB36D2EFCEBFCEB59AFFC88" box="[867,937,863,887]" direction="west" orientation="longitude" pageId="14" pageNumber="115" precision="5555" value="-2.7">
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W
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<quantity id="06519B40FFB36D2EFC0EFCEB59AFFC88" box="[902,937,863,887]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="115" unit="mm" value="2.7">2.7</quantity>
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</geoCoordinate>
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mm; P large, DN
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<quantity id="06519B40FFB36D2EFBF5FCEB5EE1FC88" box="[1149,1255,863,887]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="115" unit="mm" value="0.19">0.19 mm</quantity>
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); T sculpture of multiple axial ribs and two spiral ribs, with granules at intersections; suture canaliculate; aperture with three lamellae and a columellar denticle; umbilicus narrow.
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF4FFC7E5919F970" pageId="14" pageNumber="115" type="description">
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<paragraph id="C11636A5FFB36D2EFF4FFC7E59B2FA25" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFC7E5B5FFC1C" bold="true" box="[199,345,970,995]" pageId="14" pageNumber="115">Description.</emphasis>
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This species has a conical-shaped shell that is covered with a thin, even layer of fine sand from the early teleoconch onwards. This sand is most likely agglutinated by the live animal. The protoconch has a large, globose nucleus (DN 185–192 µm) and it is finely granular and consists of 1.0 whorls. The P/T boundary is marked by a thin varix. The teleoconch bears microscopic pustules. The first teleoconch whorl is sculpted with 15 axial ribs. Their interspaces are about three times as wide as their width. Sediment cover develops within the first 0.5 teleoconch whorl. Two spiral ribs appear on the second teleoconch whorl. The axial ribs develop granules at intersections with these spiral ribs. The whorl profile changes from round to angular as the spiral ribs become more pronounced. The area between the spiral ribs becomes slightly concave. The suture is canaliculate. The body whorl bears four additional spiral ribs. The area between the middle rib and basal rib is also slightly concave. The three basal ribs are slightly weaker than the three ribs on the preceding whorls. The base is slightly convex. The aperture is weakly nacreous and subcircular to subquadrate. The parietal margin is slightly convex and contains a thin callus and a weak knob on its upper part. The general shape of the outer lip is rounded, but the spiral ribs produce local angulations. The inner side of the outer lip contains two weak lamellae and a stronger basal lamella. The columellar lip is also rounded but the columella is nearly straight. A weak denticle is present at midpoint of columella in
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<typeStatus id="1E128807FFB36D2EFF1FFA765AFAFA24" box="[151,252,1474,1499]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
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, but absent in most other specimens. The umbilicus is narrow.
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</paragraph>
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<paragraph id="C11636A5FFB36D2EFF4FFA525919F970" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
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<emphasis id="F3DDEAB7FFB36D2EFF4FFA525B79FA00" bold="true" box="[199,383,1510,1535]" pageId="14" pageNumber="115">Differentiation.</emphasis>
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<taxonomicName id="06A94D26FFB36D2EFE04FA52587FFA01" box="[396,633,1510,1535]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="arenosa">
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<emphasis id="F3DDEAB7FFB36D2EFE04FA52587FFA01" box="[396,633,1510,1535]" italics="true" pageId="14" pageNumber="115">Calliotropis arenosa</emphasis>
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</taxonomicName>
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is most similar to the slightly taller
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<taxonomicName id="06A94D26FFB36D2EFBCAFA535F9AFA00" authority="Marshall, 1979" authorityName="Marshall" authorityYear="1979" box="[1090,1436,1510,1535]" class="Gastropoda" family="Calliotropidae" genus="Calliotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="14" pageNumber="115" phylum="Mollusca" rank="species" species="acherontis">
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<emphasis id="F3DDEAB7FFB36D2EFBCAFA535ED8FA00" box="[1090,1246,1510,1535]" italics="true" pageId="14" pageNumber="115">C. acherontis</emphasis>
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<bibRefCitation id="A5384B54FFB36D2EFB64FA525F9AFA00" author="Marshall" box="[1260,1436,1510,1535]" pageId="14" pageNumber="115" refString="Marshall, B. A. (1979) The Trochidae and Turbinidae of the Kermadec Ridge (Mollusca: Gastropoda). New Zealand Journal of Zoology, 6, 521 - 552. http: // dx. doi. org / 10.1080 / 03014223.1979.10428396" type="journal article" year="1979">
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||
<collectingCountry id="B9BE7635FFB36D2EFB64FA525F54FA00" box="[1260,1362,1510,1535]" name="Marshall Islands" pageId="14" pageNumber="115">Marshall</collectingCountry>
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, 1979
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</bibRefCitation>
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</taxonomicName>
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(Recent, southwest-Pacific), but the latter species does not agglutinate sand to the shell. Furthermore, the specimens figured by
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<bibRefCitation id="A5384B54FFB36D2EFE11F99A5854F9B8" author="Marshall" box="[409,594,1582,1607]" pageId="14" pageNumber="115" refString="Marshall, B. A. (1979) The Trochidae and Turbinidae of the Kermadec Ridge (Mollusca: Gastropoda). New Zealand Journal of Zoology, 6, 521 - 552. http: // dx. doi. org / 10.1080 / 03014223.1979.10428396" type="journal article" year="1979">
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<collectingCountry id="B9BE7635FFB36D2EFE11F99A5BFBF9B8" box="[409,509,1582,1607]" name="Marshall Islands" pageId="14" pageNumber="115">Marshall</collectingCountry>
|
||
(1979)
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</bibRefCitation>
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and
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<bibRefCitation id="A5384B54FFB36D2EFD18F99A593AF9B8" author="Vilvens" box="[656,828,1582,1607]" pageId="14" pageNumber="115" refString="Vilvens, C. (2007) New species and new records of Calliotropis (Gastropoda: Chilodontidae: Calliotropinae) from Indo- Pacific. Novapex: Trimestriel de la Societe Belge de Malacologie, 8 (Hors Serie 5), 1 - 72." type="journal article" year="2007">Vilvens (2007)</bibRefCitation>
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do not appear to have lamellae or a denticle in the aperture, while the larger specimens in the studied material, in which the aperture is not filled with sediment, always have lamellae and sometimes also have a denticle.
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</paragraph>
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</subSubSection>
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<subSubSection id="89B3652EFFB36D2EFF4FF92E5B6DF874" pageId="14" pageNumber="115" type="discussion">
|
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<paragraph id="C11636A5FFB36D2EFF4FF92E5B6DF874" blockId="14.[151,1436,646,1931]" pageId="14" pageNumber="115">
|
||
<emphasis id="F3DDEAB7FFB36D2EFF4FF92E5B3DF94C" bold="true" box="[199,315,1690,1715]" pageId="14" pageNumber="115">Remarks.</emphasis>
|
||
The layer of sediment on the adults of this species is considered here as a characteristic of the species, because it is present in all adult material and its appearance, thickness and evenness is constant and similar in all adults. Furthermore, such a characteristic layer is not present in any other species of shells in the studied material. The origin of this layer of sediment remains speculative, but is perhaps facilitated by periostracal fluids, as it is in the Caenogastropod genus
|
||
<taxonomicName id="06A94D26FFB36D2EFDC9F89E5E4EF8BC" authority="Adams, 1860 (Healy & Wells 1998)" authorityName="Adams, 1860 (Healy & Wells" authorityYear="1998" box="[577,1096,1834,1859]" class="Gastropoda" family="Scaliolidae" genus="Scaliola" kingdom="Animalia" pageId="14" pageNumber="115" phylum="Mollusca" rank="genus">
|
||
<emphasis id="F3DDEAB7FFB36D2EFDC9F89E589BF8BC" box="[577,669,1834,1859]" italics="true" pageId="14" pageNumber="115">Scaliola</emphasis>
|
||
Adams, 1860 (
|
||
<bibRefCitation id="A5384B54FFB36D2EFCC6F89E5E46F8BC" author="Healy" box="[846,1088,1834,1859]" pageId="14" pageNumber="115" refString="Healy, J. M. & Wells, F. E. (1998) Superfamily Cerithioidea. In: Beesley, P. L., Ross, G. J. B. & Wells, A. (Eds.), Mollusca: The Southern Synthesis. Fauna of Australia. Vol. 5. CSIRO Publishing, Melbourne, Part B, pp. 707 - 733." type="book" year="1998">Healy & Wells 1998</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. One specimen in the studied material has grown wider whorls after a break in the shell, so injuries to snails of this species may sometimes alter shell development.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |