treatments-xml/data/AA/6E/87/AA6E87D5BE42C81B8FFDF8E3FE49FA64.xml

<document id="E9699C8F7C51E226E970E5F243ED3CF0" ID-DOI="10.11646/phytotaxa.302.2.10" ID-ISSN="1179-3163" ID-Zenodo-Dep="13695212" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725557682662" checkinUser="felipe" docAuthor="Ramasubbu, R., Divya, C., Kala, N. Sasi, Surendran, Anjana &amp; Sreekala, A. K." docDate="2017" docId="AA6E87D5BE42C81B8FFDF8E3FE49FA64" docLanguage="en" docName="phytotaxa.302.2.10.pdf" docOrigin="Phytotaxa 302 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.302.2.10" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Impatiens megamalayana Ramasubbu 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="196" masterDocId="5657FFADBE42C8188F75FFF7FFC4FFC1" masterDocTitle="Impatiens megamalayana, a new species of Impatiens from the Western Ghats, Tamil Nadu, India" masterLastPageNumber="197" masterPageNumber="193" pageNumber="193" updateTime="1725558065160" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CE5AEAECC26985170ABE050EABFB640C">Impatiens megamalayana, a new species of Impatiens from the Western Ghats, Tamil Nadu, India</mods:title>
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<mods:namePart id="E3DA2248B60922D66B631B487482B65B">Ramasubbu, R.</mods:namePart>
<mods:affiliation id="80444A4B0C532C7995859CA9CE6F123D">Department of Biology, The Gandhigram Rural Institute- Deemed University Dindigul, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="D08E76EB1F7682840D6C413B010187FA">Divya, C.</mods:namePart>
<mods:affiliation id="FF2DB93E106CDD91DB0AEEE6F1E4B6C1">Department of Biology, The Gandhigram Rural Institute- Deemed University Dindigul, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="C13BF92B8A598D4D5FE7F680B86FFD39">Kala, N. Sasi</mods:namePart>
<mods:affiliation id="31B3604EEEB219AE01ABEB916121FB9A">Department of Biology, The Gandhigram Rural Institute- Deemed University Dindigul, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="9B486AF890D7A4419101A85E32A52C6A">Surendran, Anjana</mods:namePart>
<mods:affiliation id="DF20AB12ED28CF7F286BA38B66549A32">Department of Biology, The Gandhigram Rural Institute- Deemed University Dindigul, Tamil Nadu, India</mods:affiliation>
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<mods:namePart id="58BFBEDF989705572236A228958130A6">Sreekala, A. K.</mods:namePart>
<mods:affiliation id="4000491378BA74E2380C8458102F81B1">Conservation Biology Division, Jawaharlal Nehru Tropical Botanic Garden and Research Institute Thiruvananthapuram, Kerala, India</mods:affiliation>
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<paragraph id="227836C3BE42C8188FFDF8E3FCD4F8EE" blockId="0.[136,784,1812,1839]" box="[136,784,1812,1839]" pageId="0" pageNumber="193">
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<taxonomicName id="E5C74D40BE42C8188FFDF8E3FDF9F8EE" ID-CoL="3PK4Q" authority="Ramasubbu" authorityName="Ramasubbu" authorityYear="2017" box="[136,573,1812,1839]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="0" pageNumber="193" phylum="Tracheophyta" rank="species" species="megamalayana" status="sp. nov.">
<emphasis id="10B3EAD1BE42C8188FFDF8E3FE76F8EE" bold="true" box="[136,434,1812,1839]" italics="true" pageId="0" pageNumber="193">Impatiens megamalayana</emphasis>
Ramasubbu
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<emphasis id="10B3EAD1BE42C8188D31F8E1FD5DF8EE" box="[580,665,1814,1839]" italics="true" pageId="0" pageNumber="193">
<taxonomicNameLabel id="0B8057AABE42C8188D31F8E1FD5DF8EE" box="[580,665,1814,1839]" pageId="0" pageNumber="193" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="BAFC2A46BE42C8188DDDF8E1FD2EF8EE" box="[680,746,1814,1839]" captionStart="FIGURE 1" captionStartId="1.[136,229,1915,1936]" captionTargetBox="[164,1423,195,1879]" captionTargetId="figure-14@1.[151,1436,188,1888]" captionTargetPageId="1" captionText="FIGURE 1. Impatiens megamalayana Ramasubbu sp. nov. A. Upper portion of adult plant, B. Leaves, adaxial view, C. Petiole with pairs of extra-petiolar glands, D. Leaf, lateral view, showing hairs, E. Leaves, adaxial view, showing main nerve with single-celled hairs, F. Flower, ventral view, G. Flower lateral view, H. Lower sepal, ventral view, I. SEM of pollen grain, J. Ovary, K. Fruit, L. Seeds, M. Ornamentation of seed coat. (Photographs by R. Ramasubbu)." figureDoi="http://doi.org/10.5281/zenodo.13695216" httpUri="https://zenodo.org/record/13695216/files/figure.png" pageId="0" pageNumber="193">Fig. 1</figureCitation>
, 
<figureCitation id="BAFC2A46BE42C8188D8DF8E1FCC3F8EE" box="[760,775,1814,1839]" captionStart="FIGURE 2" captionStartId="2.[136,229,1953,1974]" captionTargetBox="[164,1427,215,1917]" captionTargetId="figure-13@2.[156,1431,190,1929]" captionTargetPageId="2" captionText="FIGURE 2. Impatiens megamalayana Ramasubbu sp. nov.A. Habit, B. Leaf blade, lateral (top) view and adaxial (bottom) view, C. Stem and leaves showing pairs of extra-petiolar glands, D. Flower, ventral view, E. Flower, lateral view, F. Lateral view of lower sepal, G. dorsal view of lateral sepals, H. Adaxial view of dorsal petal, I. Top view of lateral petals, J. Androecium, K. Ovary, L. Seed. (Illustrated by Dr. D. Felix Irudhyaraj based on fresh specimens)." figureDoi="http://doi.org/10.5281/zenodo.13695218" httpUri="https://zenodo.org/record/13695218/files/figure.png" pageId="0" pageNumber="193">2</figureCitation>
)
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</paragraph>
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<paragraph id="227836C3BE42C8188FFDF897FBC2F858" blockId="0.[136,1452,1887,2053]" pageId="0" pageNumber="193">
Type:─ 
<collectingCountry id="5AD07653BE42C8188FBAF897FED2F8B4" box="[207,278,1888,1909]" name="India" pageId="0" pageNumber="193">INDIA</collectingCountry>
. 
<collectingRegion id="E003F821BE42C8188E68F897FE55F8B4" box="[285,401,1888,1909]" country="India" name="Tamil Nadu" pageId="0" pageNumber="193">Tamil Nadu</collectingRegion>
: Theni District, Megamalai Hills way to Highwavies, 
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09 
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39′06.36″N
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, 
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77 
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21′05.08″E
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, 
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a.s.l, 
<date id="56791003BE42C8188A5FF897FF2BF858" pageId="0" pageNumber="193" value="2015-09-11">11 September 2015</date>
, 
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<collectionCode id="44D6AE06BE42C8188F8CF873FECEF858" box="[249,266,1924,1945]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="193">R</collectionCode>
. Ramasubbu 609
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(
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<collectionCode id="44D6AE06BE42C8188D62F873FD80F858" box="[535,580,1924,1945]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="0" pageNumber="193">MH</collectionCode>
!, 
<typeStatus id="FD7C8861BE42C8188D27F873FD53F858" box="[594,663,1924,1945]" pageId="0" pageNumber="193" type="isotype">isotype</typeStatus>
Gandhigram University Herbarium!).
</paragraph>
<paragraph id="227836C3BE42C8188FFDF85FFCB6F7C4" blockId="0.[136,1452,1887,2053]" pageId="0" pageNumber="193">
Diagnosis: This species is closely allied to 
<taxonomicName id="E5C74D40BE42C8188D47F85FFC6AF87C" authority="Hook. f. (1911: 354)" authorityName="Hook. f." authorityPageNumber="354" authorityYear="1911" box="[562,942,1960,1981]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="0" pageNumber="193" phylum="Tracheophyta" rank="species" species="herbicola">
<emphasis id="10B3EAD1BE42C8188D47F85FFD31F87C" box="[562,757,1960,1981]" italics="true" pageId="0" pageNumber="193">Impatiens herbicola</emphasis>
Hook.f.(1911:354)
</taxonomicName>
and 
<taxonomicName id="E5C74D40BE42C8188CAEF85FFB69F87C" authority="Benth." authorityName="Benth." box="[987,1197,1960,1981]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="0" pageNumber="193" phylum="Tracheophyta" rank="species" species="inconspicua">
<emphasis id="10B3EAD1BE42C8188CAEF85FFBA1F87C" box="[987,1125,1960,1981]" italics="true" pageId="0" pageNumber="193">I. inconspicua</emphasis>
Benth.
</taxonomicName>
in Wall. (1831:4741), the latter with three varieties, but it differs by its ridged stem, ensiform and villous leaves, a wrinkled boat-shaped lower sepal with outwardly curved tip, ornamented seeds and strictly ovate pollen grains.
</paragraph>
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<paragraph id="227836C3BE43C8198FFDF88CFD19F83D" blockId="1.[136,1452,1915,2044]" pageId="1" pageNumber="194">
<emphasis id="10B3EAD1BE43C8198FFDF88CFEC4F851" bold="true" box="[136,256,1915,1936]" pageId="1" pageNumber="194">FIGURE 1.</emphasis>
<taxonomicName id="E5C74D40BE43C8198E7DF88CFDB8F851" authority="Ramasubbu" authorityName="Ramasubbu" authorityYear="2017" box="[264,636,1915,1936]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="194" phylum="Tracheophyta" rank="species" species="megamalayana" status="sp. nov.">
<emphasis id="10B3EAD1BE43C8198E7DF88CFDC6F851" box="[264,514,1915,1936]" italics="true" pageId="1" pageNumber="194">Impatiens megamalayana</emphasis>
Ramasubbu
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<emphasis id="10B3EAD1BE43C8198DF1F88CFD14F851" box="[644,720,1915,1936]" italics="true" pageId="1" pageNumber="194">
<taxonomicNameLabel id="0B8057AABE43C8198DF1F88CFD14F851" box="[644,720,1915,1936]" pageId="1" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Upper portion of adult plant, B. Leaves, adaxial view, C. Petiole with pairs of extra-petiolar glands, D. Leaf, lateral view, showing hairs, E. Leaves, adaxial view, showing main nerve with single-celled hairs, F. Flower, ventral view, G. Flower lateral view, H. Lower sepal, ventral view, I. SEM of pollen grain, J. Ovary, K. Fruit, L. Seeds, M. Ornamentation of seed coat. (Photographs by R. Ramasubbu).
</paragraph>
</caption>
<caption id="76B8664BBE40C81A8FFDF856FD8EF7E3" ID-DOI="http://doi.org/10.5281/zenodo.13695218" ID-Zenodo-Dep="13695218" httpUri="https://zenodo.org/record/13695218/files/figure.png" pageId="2" pageNumber="195" startId="2.[136,229,1953,1974]" targetBox="[164,1427,215,1917]" targetPageId="2" targetType="figure">
<paragraph id="227836C3BE40C81A8FFDF856FD8EF7E3" blockId="2.[136,1452,1953,2082]" pageId="2" pageNumber="195">
<emphasis id="10B3EAD1BE40C81A8FFDF856FF39F877" bold="true" box="[136,253,1953,1974]" pageId="2" pageNumber="195">FIGURE 2.</emphasis>
<taxonomicName id="E5C74D40BE40C81A8E76F856FDB5F877" authority="Ramasubbu" authorityName="Ramasubbu" authorityYear="2017" box="[259,625,1953,1974]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="megamalayana" status="sp. nov.">
<emphasis id="10B3EAD1BE40C81A8E76F856FE3EF877" box="[259,506,1953,1974]" italics="true" pageId="2" pageNumber="195">Impatiens megamalayana</emphasis>
Ramasubbu
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<emphasis id="10B3EAD1BE40C81A8D02F856FD04F877" box="[631,704,1953,1974]" italics="true" pageId="2" pageNumber="195">
<taxonomicNameLabel id="0B8057AABE40C81A8D02F856FD04F877" box="[631,704,1953,1974]" pageId="2" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Habit, B. Leaf blade, lateral (top) view and adaxial (bottom) view, C. Stem and leaves showing pairs of extra-petiolar glands, D. Flower, ventral view, E. Flower, lateral view, F. Lateral view of lower sepal, G. dorsal view of lateral sepals, H. Adaxial view of dorsal petal, I. Top view of lateral petals, J. Androecium, K. Ovary, L. Seed. (Illustrated by Dr. 
<taxonomicName id="E5C74D40BE40C81A8FFDF7FAFE85F7E3" authority="Irudhyaraj" authorityName="Irudhyaraj" box="[136,321,2061,2082]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="195" phylum="Tracheophyta" rank="species" species="felix">D. Felix Irudhyaraj</taxonomicName>
based on fresh specimens).
</paragraph>
</caption>
<paragraph id="227836C3BE41C81B8FFDFF57FEE9FD70" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
Annual, succulent, erect, glabrous herbs 
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tall; stem purplish, cylindrical with 3–5 ridges, often branched, branches erect, nodes slightly swollen, internode elongate, 
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. Leaves in a pair, opposite decussate, shortly petiolate, petiole 
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, densely hairy, ensiform, acuminate, entire, base truncate, slightly cordate, reflexed upwards, 2.5–3.5 × 
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, densely hairy above, smooth below, completely purplish at base when mature, lower part of main nerve densely hairy, leaf margin distinctly serrate, leaf blade teeth 
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, extra petiolar glands present. Inflorescence 1–2 per angle (4 per node), flowers simple, axillary, 
<quantity id="E53F9B26BE41C81B8DABFEA3FCA2FEAC" box="[734,870,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.7" metricValueMin="3.8" pageId="3" pageNumber="196" unit="mm" value="4.25" valueMax="4.7" valueMin="3.8">3.8–4.7 mm</quantity>
wide, pale purple with deep reddish-yellow spot at base; pedicellate, pedicel 
<quantity id="E53F9B26BE41C81B8ED2FE8FFDE0FE50" box="[423,548,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="3" pageNumber="196" unit="cm" value="1.55" valueMax="1.8" valueMin="1.3">1.3–1.8 cm</quantity>
at anthesis, 
<quantity id="E53F9B26BE41C81B8DDAFE8FFCEFFE50" box="[687,811,376,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="3" pageNumber="196" unit="cm" value="2.0" valueMax="2.3" valueMin="1.7">1.7–2.3 cm</quantity>
in fruit, with one row of long hairs. Sepals: lateral sepals 2, sickle-shaped, apex acuminate, 
<quantity id="E53F9B26BE41C81B8D66FE6BFD59FE74" box="[531,669,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.6" metricValueMin="1.8" pageId="3" pageNumber="196" unit="mm" value="2.2" valueMax="2.6" valueMin="1.8">1.8–2.6 mm</quantity>
long, 1–2 nerved, upwardly curved, purplish, densely hairy. Petals: dorsal petal orbicular, 2–2.3 × 
<quantity id="E53F9B26BE41C81B8E99FE37FDBCFE18" box="[492,632,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="3.8" metricValueMin="3.2" pageId="3" pageNumber="196" unit="mm" value="3.5" valueMax="3.8" valueMin="3.2">3.2–3.8 mm</quantity>
, recurved, dorsally keeled, apiculate, keel densely covered with singlecelled hairs, lateral united petals oblong-spherical, deeply clawed, 2-lobed, 2.6–3.1× 
<quantity id="E53F9B26BE41C81B8B21FE13FB1BFE3C" box="[1108,1247,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.6" pageId="3" pageNumber="196" unit="mm" value="3.95" valueMax="4.3" valueMin="3.6">3.6–4.3 mm</quantity>
, basal lobe small, oblong, distal lobe round or spherical, pale orange with deep reddish-yellow spot, dorsal auricle prominent, obtuse; lower sepal boat-shaped, 3.2–4.3 × 
<quantity id="E53F9B26BE41C81B8D69FDDBFD62FD84" box="[540,678,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="3" pageNumber="196" unit="mm" value="1.2" valueMax="1.6" valueMin="0.8">0.8–1.6 mm</quantity>
, with 4–7 wrinkles, tip of lower sepal outwardly curved, horizontal, spur absent, densely hairy, purple. Ovary elongate, 
<quantity id="E53F9B26BE41C81B8DB2FDA7FC8AFDA8" box="[711,846,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="3" pageNumber="196" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
long. Fruit a capsule, ovoid, turgid, 11–13 × 
<quantity id="E53F9B26BE41C81B8A3DFDA7FA63FDA8" box="[1352,1447,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="196" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, acute, beaked, 5–9-seeded. Seed round with prominent caruncle, compressed, 0.8–1.3 × 
<quantity id="E53F9B26BE41C81B8BF3FD83FAD6FD4C" box="[1158,1298,628,653]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="3" pageNumber="196" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
, ornamented, black, shining.
</paragraph>
<paragraph id="227836C3BE41C81B8FC8FD4DFD52FD14" blockId="3.[136,1453,160,1445]" box="[189,662,698,725]" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FD4DFE0CFD15" bold="true" box="[189,456,698,725]" italics="true" pageId="3" pageNumber="196">Flowering and fruiting</emphasis>
—July to October.
</paragraph>
</subSubSection>
<subSubSection id="6ADD6548BE41C81B8FC8FD29FBF1FC80" pageId="3" pageNumber="196" type="description">
<paragraph id="227836C3BE41C81B8FC8FD29FBF1FC80" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FD29FD1DFD38" box="[189,729,734,761]" italics="true" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FD29FE5BFD38" bold="true" box="[189,415,734,761]" italics="true" pageId="3" pageNumber="196">Conservation status</emphasis>
— 
<taxonomicName id="E5C74D40BE41C81B8ECEFD17FD1DFD38" authorityName="Ramasubbu" authorityYear="2017" box="[443,729,736,761]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="megamalayana">Impatiens megamalayana</taxonomicName>
</emphasis>
is assessed as Critically Endangered (CR B1ab(i,ii,v); 2ab(i,ii,iv); D) in accordance with the IUCN guidelines(
<bibRefCitation id="46564B32BE41C81B8D1AFCF3FD31FCDC" author="IUCN" box="[623,757,772,797]" pageId="3" pageNumber="196" refId="ref2318" refString="IUCN (2016) Guidelines for using the IUCN list categories and Criteria version 12. Prepared by IUCN standards and petitions committee. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 29 March 2016)" type="book" year="2016">IUCN 2016</bibRefCitation>
).However, more field studies are required to observe details as to the extent of decline. The habitat of the species is severely affected by landslides.
</paragraph>
</subSubSection>
<subSubSection id="6ADD6548BE41C81B8FC8FCBDFF08FC48" pageId="3" pageNumber="196" type="etymology">
<paragraph id="227836C3BE41C81B8FC8FCBDFF08FC48" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FCBDFEFCFCA4" bold="true" box="[189,312,842,869]" italics="true" pageId="3" pageNumber="196">Etymology</emphasis>
—The species for its occurrence in the Megamalai Wildlife Sanctuary, a part of Western Ghats, 
<collectingRegion id="E003F821BE41C81B8A1EFCBBFF0CFC48" country="India" name="Tamil Nadu" pageId="3" pageNumber="196">Tamil Nadu</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="6ADD6548BE41C81B8FC8FC65FE49FA64" pageId="3" pageNumber="196" type="distribution">
<paragraph id="227836C3BE41C81B8FC8FC65FE79FC10" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FC65FE8DFC6C" bold="true" box="[189,329,914,941]" italics="true" pageId="3" pageNumber="196">Distribution</emphasis>
—Rare on hill tops at 
<quantity id="E53F9B26BE41C81B8D33FC63FD58FC6C" box="[582,668,916,941]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="3" pageNumber="196" unit="m" value="1451.0">1451 m</quantity>
a.s.l., only found on the way to Highwavies, Megamalai Hills, Theni District, 
<collectingRegion id="E003F821BE41C81B8F9FFC4FFEB5FC10" box="[234,369,952,977]" country="India" name="Tamil Nadu" pageId="3" pageNumber="196">Tamil Nadu</collectingRegion>
, 
<collectingCountry id="5AD07653BE41C81B8E09FC4FFE7DFC10" box="[380,441,952,977]" name="India" pageId="3" pageNumber="196">India</collectingCountry>
.
</paragraph>
<paragraph id="227836C3BE41C81B8FC8FC2DFC4CFBA0" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
<materialsCitation id="92AF3C9EBE41C81B8FC8FC2DFD9BFBFC" collectingDate="2016-10-16" collectionCode="G" collectorName="Manikandan &amp; Anjana Surendran" country="India" county="Theni District" elevation="1451" latitude="9.651766" location="Highwavies" longLatPrecision="1" longitude="77.35141" municipality="Megamalai Hills" pageId="3" pageNumber="196" specimenCode="GU 627" specimenCount="2" stateProvince="Tamil Nadu">
<emphasis id="10B3EAD1BE41C81B8FC8FC2DFD06FC35" bold="true" box="[189,706,986,1013]" italics="true" pageId="3" pageNumber="196">
<typeStatus id="FD7C8861BE41C81B8FC8FC2DFDF1FC34" box="[189,565,986,1013]" isOtherTypeStatus="true" pageId="3" pageNumber="196">Additional specimens examined</typeStatus>
(
<typeStatus id="FD7C8861BE41C81B8D3CFC2DFD7CFC35" box="[585,696,986,1012]" pageId="3" pageNumber="196" type="paratype">paratypes</typeStatus>
)
</emphasis>
— 
<collectingCountry id="5AD07653BE41C81B8DAAFC2BFCF4FC34" box="[735,816,988,1013]" name="India" pageId="3" pageNumber="196">INDIA</collectingCountry>
, 
<collectingRegion id="E003F821BE41C81B8C4AFC2BFC0FFC34" box="[831,971,988,1013]" country="India" name="Tamil Nadu" pageId="3" pageNumber="196">Tamil Nadu</collectingRegion>
, 
<collectingCounty id="CB194E4FBE41C81B8CAEFC2BFBBBFC34" box="[987,1151,988,1013]" pageId="3" pageNumber="196">Theni District</collectingCounty>
, 
<collectingMunicipality id="C21CACB9BE41C81B8BFAFC2BFA8BFC34" box="[1167,1359,988,1013]" pageId="3" pageNumber="196">Megamalai Hills</collectingMunicipality>
way to 
<location id="27186018BE41C81B8FFDFBF7FED5FBD8" LSID="urn:lsid:plazi:treatment:AA6E87D5BE42C81B8FFDF8E3FE49FA64:27186018BE41C81B8FFDFBF7FED5FBD8" box="[136,273,1024,1049]" country="India" county="Theni District" latitude="9.651766" longLatPrecision="1" longitude="77.35141" municipality="Megamalai Hills" name="Highwavies" pageId="3" pageNumber="196" stateProvince="Tamil Nadu">Highwavies</location>
, 
<geoCoordinate id="47F35004BE41C81B8E69FBF7FE05FBD8" box="[284,449,1022,1049]" degrees="09" direction="north" minutes="39" orientation="latitude" pageId="3" pageNumber="196" precision="1" seconds="06.36" value="9.651766">
09 
<superScript id="D5B29B8BBE41C81B8E4DFC09FEF9FBCD" attach="none" box="[312,317,1022,1036]" fontSize="6" pageId="3" pageNumber="196">º</superScript>
39′06.36″N
</geoCoordinate>
, 
<geoCoordinate id="47F35004BE41C81B8EB9FBF7FDA9FBD8" box="[460,621,1022,1049]" degrees="77" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="196" precision="1" seconds="05.08" value="77.35141">
77 
<superScript id="D5B29B8BBE41C81B8E9DFC09FE29FBCD" attach="none" box="[488,493,1022,1036]" fontSize="6" pageId="3" pageNumber="196">º</superScript>
21′05.08″E
</geoCoordinate>
, 
<elevation id="A9EAD1F0BE41C81B8D0CFBF7FCCEFBD8" box="[633,778,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="3" pageNumber="196" unit="m" value="1451.0">
<quantity id="E53F9B26BE41C81B8D0CFBF7FD0AFBD8" box="[633,718,1024,1049]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="3" pageNumber="196" unit="m" value="1451.0">1451 m</quantity>
a.s.l.
</elevation>
, 
<date id="56791003BE41C81B8C62FBF7FC12FBD8" box="[791,982,1024,1049]" pageId="3" pageNumber="196" value="2016-10-16">
<collectingDate id="463DE9EBBE41C81B8C62FBF7FC12FBD8" box="[791,982,1024,1049]" pageId="3" pageNumber="196" value="2016-10-16">16 October 2016</collectingDate>
</date>
, 
<collectionCode id="44D6AE06BE41C81B8C94FBF7FC3CFBD8" box="[993,1016,1024,1049]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="3" pageNumber="196" type="Herbarium">G</collectionCode>
. 
<collectorName id="8F325315BE41C81B8B76FBF7FB4BFBD8" box="[1027,1167,1024,1049]" pageId="3" pageNumber="196">Manikandan</collectorName>
&amp; 
<collectorName id="8F325315BE41C81B8BC4FBF7FAB8FBD8" box="[1201,1404,1024,1049]" pageId="3" pageNumber="196">Anjana Surendran</collectorName>
<specimenCode id="72619EB8BE41C81B8AF6FBF7FF76FBFC" collectionCode="GU" country="Sweden" lsid="urn:lsid:biocol.org:col:15204" name="Gotland University" pageId="3" pageNumber="196" type="Herbarium">GU 627</specimenCode>
(Gandhigram University Herbarium)
</materialsCitation>
; 
<materialsCitation id="92AF3C9EBE41C81B8D19FBD3FC41FBA0" collectingDate="2016-11-08" collectionCode="C, N" collectorName="Divya &amp; Sasi Kala" country="India" county="Theni District" elevation="1451" latitude="9.651774" location="Highwavies" longLatPrecision="1" longitude="77.351425" municipality="Megamalai Hills" pageId="3" pageNumber="196" specimenCode="GU 636" specimenCount="1" stateProvince="Tamil Nadu">
ibid., 
<geoCoordinate id="47F35004BE41C81B8DDAFBD3FC97FBFC" box="[687,851,1058,1085]" degrees="09" direction="north" minutes="39" orientation="latitude" pageId="3" pageNumber="196" precision="1" seconds="06.39" value="9.651774">
09 
<superScript id="D5B29B8BBE41C81B8DBEFBD5FD14FBF1" attach="none" box="[715,720,1058,1072]" fontSize="6" pageId="3" pageNumber="196">º</superScript>
39′06.39″N
</geoCoordinate>
, 
<geoCoordinate id="47F35004BE41C81B8C14FBD3FBC5FBFC" box="[865,1025,1058,1085]" degrees="77" direction="east" minutes="21" orientation="longitude" pageId="3" pageNumber="196" precision="1" seconds="05.13" value="77.351425">
77 
<superScript id="D5B29B8BBE41C81B8C08FBD5FC46FBF1" attach="none" box="[893,898,1058,1072]" fontSize="6" pageId="3" pageNumber="196">º</superScript>
21′05.13″E
</geoCoordinate>
, 
<elevation id="A9EAD1F0BE41C81B8B65FBD3FB61FBFC" box="[1040,1189,1060,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="3" pageNumber="196" unit="m" value="1451.0">
<quantity id="E53F9B26BE41C81B8B65FBD3FBA3FBFC" box="[1040,1127,1060,1085]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="3" pageNumber="196" unit="m" value="1451.0">1451 m</quantity>
a.s.l.
</elevation>
, 
<date id="56791003BE41C81B8BC6FBD3FA41FBFC" box="[1203,1413,1060,1085]" pageId="3" pageNumber="196" value="2016-11-08">
<collectingDate id="463DE9EBBE41C81B8BC6FBD3FA41FBFC" box="[1203,1413,1060,1085]" pageId="3" pageNumber="196" value="2016-11-08">8 November 2016</collectingDate>
</date>
, 
<collectionCode id="44D6AE06BE41C81B8AE7FBD3FA6CFBFC" box="[1426,1448,1060,1085]" country="Denmark" name="University of Copenhagen" pageId="3" pageNumber="196" type="Herbarium">C</collectionCode>
. 
<collectorName id="8F325315BE41C81B8FFDFBBFFF08FBA0" box="[136,204,1096,1121]" pageId="3" pageNumber="196">Divya</collectorName>
&amp; 
<collectionCode id="44D6AE06BE41C81B8F85FBBFFEC3FBA0" box="[240,263,1096,1121]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="196" type="Herbarium">N</collectionCode>
. 
<collectorName id="8F325315BE41C81B8E67FBBFFEB9FBA0" box="[274,381,1096,1121]" pageId="3" pageNumber="196">Sasi Kala</collectorName>
<specimenCode id="72619EB8BE41C81B8EF1FBBFFE19FBA0" box="[388,477,1096,1121]" collectionCode="GU" country="Sweden" lsid="urn:lsid:biocol.org:col:15204" name="Gotland University" pageId="3" pageNumber="196" type="Herbarium">GU 636</specimenCode>
(Gandhigram University Herbarium)
</materialsCitation>
.
</paragraph>
<paragraph id="227836C3BE41C81B8FC8FB9CFE49FA64" blockId="3.[136,1453,160,1445]" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FB9CFD84FB44" box="[189,576,1131,1157]" italics="true" pageId="3" pageNumber="196">
<emphasis id="10B3EAD1BE41C81B8FC8FB9CFEDFFB44" bold="true" box="[189,283,1131,1157]" italics="true" pageId="3" pageNumber="196">Notes—</emphasis>
<taxonomicName id="E5C74D40BE41C81B8E6EFB9BFD84FB44" authorityName="Ramasubbu" authorityYear="2017" box="[283,576,1132,1157]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="megamalayana">Impatiens megamalayana</taxonomicName>
</emphasis>
grows on rocky slopes and in grasslands near streams and marshy areas. This species is associated with 
<taxonomicName id="E5C74D40BE41C81B8E87FB67FC76FB68" authority="J. Graham" authorityName="J. Graham" box="[498,946,1168,1193]" class="Liliopsida" family="Orchidaceae" genus="Habenaria" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="longicorniculata">
<emphasis id="10B3EAD1BE41C81B8E87FB67FCF4FB68" box="[498,816,1168,1193]" italics="true" pageId="3" pageNumber="196">Habenaria longicorniculata</emphasis>
J. Graham
</taxonomicName>
, 
<taxonomicName id="E5C74D40BE41C81B8CCAFB67FF5EFB0C" authority="(Spreng.) Schult. &amp; Schult. f." authorityName="Schult. &amp; Schult. f." baseAuthorityName="Spreng." class="Magnoliopsida" family="Asteraceae" genus="Cynotis" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="10B3EAD1BE41C81B8CCAFB67FBAEFB68" box="[959,1130,1168,1193]" italics="true" pageId="3" pageNumber="196">Cynotis villosa</emphasis>
(Spreng.) Schult. &amp; Schult. f.
</taxonomicName>
, 
<taxonomicName id="E5C74D40BE41C81B8FDCFB43FE7DFB0C" authority="Thunb." authorityName="Thunb." box="[169,441,1204,1229]" class="Magnoliopsida" family="Droseraceae" genus="Drosera" kingdom="Plantae" order="Caryophyllales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="peltata">
<emphasis id="10B3EAD1BE41C81B8FDCFB43FE99FB0C" box="[169,349,1204,1229]" italics="true" pageId="3" pageNumber="196">Drosera peltata</emphasis>
Thunb.
</taxonomicName>
, 
<taxonomicName id="E5C74D40BE41C81B8EB3FB43FC9CFB0C" authority="(Hack.) Kuntze" authorityName="Kuntze" baseAuthorityName="Hack." box="[454,856,1204,1229]" class="Liliopsida" family="Poaceae" genus="Eulalia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="phaeothrix">
<emphasis id="10B3EAD1BE41C81B8EB3FB43FD5FFB0C" box="[454,667,1204,1229]" italics="true" pageId="3" pageNumber="196">Eulalia phaeothrix</emphasis>
(Hack.) Kuntze
</taxonomicName>
, 
<taxonomicName id="E5C74D40BE41C81B8C10FB43FA63FB0C" authority="(Wight &amp; Arn. ex Steud.) Miq." authorityName="Miq." baseAuthorityName="Steud." box="[869,1447,1204,1229]" class="Liliopsida" family="Poaceae" genus="Isachne" kingdom="Plantae" order="Poales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="kunthiana">
<emphasis id="10B3EAD1BE41C81B8C10FB43FBF2FB0C" box="[869,1078,1204,1229]" italics="true" pageId="3" pageNumber="196">Isachne kunthiana</emphasis>
(Wight &amp; Arn. ex Steud.) Miq.
</taxonomicName>
, and 
<taxonomicName id="E5C74D40BE41C81B8FCCFB2FFC6BFB30" authority="(Ard.) P. Beauv. ex Roem. &amp; Schult. In" authorityName="Roem. &amp; Schult. In" baseAuthorityName="Ard." box="[185,943,1240,1265]" class="Liliopsida" family="Poaceae" genus="Oplismenus" kingdom="Plantae" order="Poales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="undulatifolius">
<emphasis id="10B3EAD1BE41C81B8FCCFB2FFE27FB30" box="[185,483,1240,1265]" italics="true" pageId="3" pageNumber="196">Oplismenus undulatifolius</emphasis>
(Ard.) P. Beauv. ex Roem. &amp; Schult. In
</taxonomicName>
general floral structure, 
<taxonomicName id="E5C74D40BE41C81B8BBEFB2FFA54FB30" authorityName="Ramasubbu" authorityYear="2017" box="[1227,1424,1240,1265]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="megamalayana">
<emphasis id="10B3EAD1BE41C81B8BBEFB2FFA54FB30" box="[1227,1424,1240,1265]" italics="true" pageId="3" pageNumber="196">I. megamalayana</emphasis>
</taxonomicName>
is closely allied to 
<taxonomicName id="E5C74D40BE41C81B8E3CFB0BFE0AFAD4" box="[329,462,1276,1301]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="herbicola">
<emphasis id="10B3EAD1BE41C81B8E3CFB0BFE0AFAD4" box="[329,462,1276,1301]" italics="true" pageId="3" pageNumber="196">I. herbicola</emphasis>
</taxonomicName>
but it differs by its ridged stem, ensiform and villous leaves, a wrinkled boat-shaped lower sepal with outwardly curved tip, ornamented seeds and strictly ovate pollen grains. 
<taxonomicName id="E5C74D40BE41C81B8BFCFAD7FA6FFAF8" authorityName="Ramasubbu" authorityYear="2017" box="[1161,1451,1312,1337]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="megamalayana">
<emphasis id="10B3EAD1BE41C81B8BFCFAD7FA6FFAF8" box="[1161,1451,1312,1337]" italics="true" pageId="3" pageNumber="196">Impatiens megamalayana</emphasis>
</taxonomicName>
is phenotypically similar to 
<taxonomicName id="E5C74D40BE41C81B8EB1FAB3FDA7FA9C" box="[452,611,1348,1373]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="196" phylum="Tracheophyta" rank="species" species="inconspicua">
<emphasis id="10B3EAD1BE41C81B8EB1FAB3FDA7FA9C" box="[452,611,1348,1373]" italics="true" pageId="3" pageNumber="196">I. inconspicua</emphasis>
</taxonomicName>
, the latter with three varieties (
<bibRefCitation id="46564B32BE41C81B8CB0FAB3FBA0FA9C" author="Bhaskar, V." box="[965,1124,1348,1373]" pageId="3" pageNumber="196" refId="ref2071" refString="Bhaskar, V. (2012) Taxonomic Monograph on Impatiens L. (Balsaminaceae) of Western Ghats, South India: The Key Genus for Endemism. Centre for Plant Taxonomic Studies, Bangalore, India, 283 pp." type="book" year="2012">Bhaskar 2012</bibRefCitation>
) but vary from characteristic sickle shaped lateral sepals, densely hairy, wrinkled boat-shaped lower sepals, ornamented seeds and 3-colpate, ovate pollen grains (
<tableCitation id="6F450378BE41C81B8E5DFA7BFEB8FA64" box="[296,380,1420,1445]" captionStart="TABLE 1" captionStartId="3.[136,224,1493,1516]" captionText="TABLE 1. Comparative morphology of Impatiens inconspicua, I. herbicola and I. megamalayana." pageId="3" pageNumber="196">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>