treatments-xml/data/BE/31/10/BE31101C9A718D625EBC2FD513F09F04.xml

<document ID-DOI="10.11646/zootaxa.5258.3.1" ID-GBIF-Dataset="9e20f525-2e14-424c-b808-fe4389c291a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7780211" ID-ZooBank="240213D7-D155-4C08-BF56-EABBAA4AC00F" approvalRequired="2" approvalRequired_for_taxonomicNames="2" checkinTime="1680072242305" checkinUser="plazi" docAuthor="Boyko, Christopher B. &amp; Williams, Jason D." docDate="2023" docId="BE31101C9A718D625EBC2FD513F09F04" docLanguage="en" docName="zootaxa.5258.3.1.pdf" docOrigin="Zootaxa 5258 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5258.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anathelges foliosus" docType="treatment" docVersion="3" lastPageNumber="252" masterDocId="420868649A708D635E2B2C0612749862" masterDocTitle="Nomenclatural and taxonomic changes in parasitic isopods (Isopoda: Epicaridea) including two new families and note on the questionable association between monogeneans and bopyrids" masterLastPageNumber="269" masterPageNumber="251" pageNumber="252" updateTime="1680095728178" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Nomenclatural and taxonomic changes in parasitic isopods (Isopoda: Epicaridea) including two new families and note on the questionable association between monogeneans and bopyrids</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Boyko, Christopher B.</mods:namePart>
<mods:affiliation>Department of Biology, Hofstra University, Hempstead, NY 11549, U. S. A. and Division of Invertebrate Zoology, American Museum of Natural History, Central Park West @ 79 Street, New York, NY, U. S. A.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Williams, Jason D.</mods:namePart>
<mods:affiliation>Department of Biology, Hofstra University, Hempstead, NY 11549, U. S. A.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-29</mods:number>
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<mods:detail type="volume">
<mods:number>5258</mods:number>
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<mods:number>3</mods:number>
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<subSubSection box="[151,866,979,1006]" pageId="1" pageNumber="252" type="nomenclature">
<paragraph blockId="1.[151,866,979,1041]" box="[151,866,979,1006]" pageId="1" pageNumber="252">
<heading bold="true" box="[151,866,979,1006]" fontSize="11" level="1" pageId="1" pageNumber="252" reason="1">
<emphasis bold="true" box="[151,866,979,1006]" pageId="1" pageNumber="252">
<taxonomicName box="[151,389,979,1006]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis bold="true" box="[151,389,979,1006]" italics="true" pageId="1" pageNumber="252">Anathelges foliosus</emphasis>
</taxonomicName>
(Kröyer 
<emphasis bold="true" box="[503,527,979,1005]" italics="true" pageId="1" pageNumber="252">in</emphasis>
Gaimard, [1842]) n. comb.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="252" type="description">
<paragraph blockId="1.[151,866,979,1041]" box="[151,228,1015,1041]" pageId="1" pageNumber="252">
(
<figureCitation box="[159,217,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[151,250,774,798]" captionTargetBox="[177,1412,203,729]" captionTargetId="figure-24@2.[151,1436,181,751]" captionTargetPageId="2" captionText="FIGURE 1. Bopyrus foliosus Kr̂yer in Gaimard, [1842]. A, female, ventral view. B, female, dorsal view. C, male, dorsal view. D, pereopod (presumed from male). Stegophrixus [sic] thompsoni Nierstrasz &amp; Brender à Brandis, 1931. E, female, dorsal view (oostegites not drawn). F, male, dorsal view. Figures not to scale. A–D from Kr̂yer in Gaimard [1842]; E, F from Nierstrasz &amp; Brender à Brandis (1931)." figureDoi="http://doi.org/10.5281/zenodo.7780213" httpUri="https://zenodo.org/record/7780213/files/figure.png" pageId="1" pageNumber="252">Fig 1</figureCitation>
)
</paragraph>
<paragraph blockId="1.[151,933,1084,1138]" box="[151,620,1084,1108]" pageId="1" pageNumber="252">
= 
<taxonomicName authority="Kr" authorityName="Kr" box="[172,377,1084,1108]" class="Malacostraca" family="Bopyridae" genus="Bopyrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[172,342,1084,1108]" italics="true" pageId="1" pageNumber="252">Bopyrus foliosus</emphasis>
Kr
</taxonomicName>
̂yer 
<emphasis box="[426,446,1084,1107]" italics="true" pageId="1" pageNumber="252">in</emphasis>
Gaimard, [1842]
</paragraph>
<paragraph blockId="1.[151,933,1084,1138]" box="[151,933,1114,1138]" pageId="1" pageNumber="252">
= 
<taxonomicName authority="Nierstrasz &amp; Brender" authorityName="Nierstrasz &amp; Brender" box="[172,691,1114,1138]" class="Malacostraca" family="Bopyridae" genus="Stegophrixus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[172,304,1114,1138]" italics="true" pageId="1" pageNumber="252">Stegophrixus</emphasis>
(
<emphasis box="[319,347,1115,1138]" italics="true" pageId="1" pageNumber="252">sic</emphasis>
) 
<emphasis box="[362,468,1114,1138]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
Nierstrasz &amp; Brender
</taxonomicName>
à Brandis, 1931 
<emphasis bold="true" box="[863,933,1114,1138]" pageId="1" pageNumber="252">n. syn.</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[151,956,1184,1211]" pageId="1" pageNumber="252" type="distribution">
<paragraph blockId="1.[151,1437,1184,1895]" box="[151,956,1184,1211]" pageId="1" pageNumber="252">
<emphasis bold="true" box="[151,307,1184,1210]" pageId="1" pageNumber="252">Distribution:</emphasis>
<collectingRegion box="[314,435,1184,1210]" country="Chile" name="Valparaiso" pageId="1" pageNumber="252">Valparaíso</collectingRegion>
, 
<collectingCountry box="[446,507,1184,1210]" name="Chile" pageId="1" pageNumber="252">Chile</collectingCountry>
(Nierstrasz &amp; Brender à Brandis 1931).
</paragraph>
</subSubSection>
<subSubSection box="[199,890,1220,1247]" pageId="1" pageNumber="252" type="biology_ecology">
<paragraph blockId="1.[151,1437,1184,1895]" box="[199,890,1220,1247]" pageId="1" pageNumber="252">
<emphasis bold="true" box="[199,275,1221,1247]" pageId="1" pageNumber="252">Hosts:</emphasis>
“ 
<taxonomicName box="[295,390,1221,1246]" class="Malacostraca" family="Paguridae" genus="Pagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis box="[295,390,1221,1246]" italics="true" pageId="1" pageNumber="252">Pagurus</emphasis>
</taxonomicName>
” sp. (Nierstrasz &amp; Brender à Brandis 1931).
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="252" type="discussion">
<paragraph blockId="1.[151,1437,1184,1895]" pageId="1" pageNumber="252">
<emphasis bold="true" box="[199,317,1257,1283]" pageId="1" pageNumber="252">Remarks:</emphasis>
There was never a text description published for 
<taxonomicName box="[867,1054,1256,1282]" class="Malacostraca" family="Bopyridae" genus="Bopyrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[867,1054,1256,1282]" italics="true" pageId="1" pageNumber="252">Bopyrus foliosus</emphasis>
</taxonomicName>
and the availability of the name is from Kr̂yer’s plate 29, fig. 2, although the dating of this and the other plates in the volume is contentious (see note under Kr̂yer 
<emphasis box="[307,329,1329,1354]" italics="true" pageId="1" pageNumber="252">in</emphasis>
Gaimard [1842] in References). As far as can be determined, this species has only been cited in the literature one time subsequent to its introduction as a brief mention by Spence Bate &amp; Westwood (1867). The 
<typeStatus box="[1388,1436,1365,1391]" pageId="1" pageNumber="252">type</typeStatus>
specimens (ZMUC CRU-006445) are, unfortunately, missing from the ZMUC collection.
</paragraph>
<paragraph blockId="1.[151,1437,1184,1895]" pageId="1" pageNumber="252">
A comparison of the figures of Kr̂yer 
<emphasis box="[656,678,1437,1462]" italics="true" pageId="1" pageNumber="252">in</emphasis>
Gaimard ([1842]) (
<figureCitation box="[907,1030,1436,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,774,798]" captionTargetBox="[177,1412,203,729]" captionTargetId="figure-24@2.[151,1436,181,751]" captionTargetPageId="2" captionText="FIGURE 1. Bopyrus foliosus Kr̂yer in Gaimard, [1842]. A, female, ventral view. B, female, dorsal view. C, male, dorsal view. D, pereopod (presumed from male). Stegophrixus [sic] thompsoni Nierstrasz &amp; Brender à Brandis, 1931. E, female, dorsal view (oostegites not drawn). F, male, dorsal view. Figures not to scale. A–D from Kr̂yer in Gaimard [1842]; E, F from Nierstrasz &amp; Brender à Brandis (1931)." figureDoi="http://doi.org/10.5281/zenodo.7780213" httpUri="https://zenodo.org/record/7780213/files/figure.png" pageId="1" pageNumber="252">Fig. 1A–D</figureCitation>
herein) with those of Nierstrasz &amp; Brender à Brandis (1931: figs. 87–89) (
<figureCitation box="[587,699,1472,1499]" captionStart="FIGURE 1" captionStartId="2.[151,250,774,798]" captionTargetBox="[177,1412,203,729]" captionTargetId="figure-24@2.[151,1436,181,751]" captionTargetPageId="2" captionText="FIGURE 1. Bopyrus foliosus Kr̂yer in Gaimard, [1842]. A, female, ventral view. B, female, dorsal view. C, male, dorsal view. D, pereopod (presumed from male). Stegophrixus [sic] thompsoni Nierstrasz &amp; Brender à Brandis, 1931. E, female, dorsal view (oostegites not drawn). F, male, dorsal view. Figures not to scale. A–D from Kr̂yer in Gaimard [1842]; E, F from Nierstrasz &amp; Brender à Brandis (1931)." figureDoi="http://doi.org/10.5281/zenodo.7780213" httpUri="https://zenodo.org/record/7780213/files/figure.png" pageId="1" pageNumber="252">Fig. 1E, F</figureCitation>
herein) shows that 
<taxonomicName box="[914,1101,1472,1498]" class="Malacostraca" family="Bopyridae" genus="Bopyrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[914,1101,1472,1498]" italics="true" pageId="1" pageNumber="252">Bopyrus foliosus</emphasis>
</taxonomicName>
is identical with 
<taxonomicName authority="Nierstrasz &amp; Brender" authorityName="Nierstrasz &amp; Brender" class="Malacostraca" family="Bopyridae" genus="Stegophrixus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[1290,1436,1472,1498]" italics="true" pageId="1" pageNumber="252">Stegophrixus</emphasis>
(
<emphasis box="[161,192,1509,1534]" italics="true" pageId="1" pageNumber="252">sic</emphasis>
) 
<emphasis box="[212,329,1508,1534]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
Nierstrasz &amp; Brender
</taxonomicName>
à Brandis, 1931 (currently 
<taxonomicName baseAuthorityName="Nierstrasz &amp; Brender a Brandis" baseAuthorityYear="1931" box="[916,1168,1508,1534]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[916,1168,1508,1534]" italics="true" pageId="1" pageNumber="252">Anathelges thompsoni</emphasis>
</taxonomicName>
(Nierstrasz &amp; Brender à Brandis, 1931)); the 
<typeStatus box="[413,461,1545,1571]" pageId="1" pageNumber="252">type</typeStatus>
material of 
<taxonomicName baseAuthorityName="Nierstrasz &amp; Brender a Brandis" baseAuthorityYear="1931" box="[606,754,1544,1571]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[606,620,1545,1571]" italics="true" pageId="1" pageNumber="252">S</emphasis>
. 
<emphasis box="[637,754,1544,1570]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
</taxonomicName>
was examined by 
<bibRefCitation author="Boyko, C. B. &amp; Williams, J. D." box="[973,1278,1544,1571]" pageId="1" pageNumber="252" pagination="795 - 813" refId="ref10782" refString="Boyko, C. B. &amp; Williams, J. D. (2003) A revision of Anathelges and Stegophryxus (Isopoda: Bopyridae: Athelginae) with descriptions of two new genera and one new species. Journal of Crustacean Biology, 23 (4), 795 - 813. https: // doi. org / 10.1651 / C- 2398" type="journal article" year="2003">Boyko &amp; Williams (2003)</bibRefCitation>
. Nierstrasz &amp; Brender à Brandis (1931) based their description of 
<taxonomicName baseAuthorityName="Nierstrasz &amp; Brender a Brandis" baseAuthorityYear="1931" box="[743,889,1580,1607]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[743,757,1581,1607]" italics="true" pageId="1" pageNumber="252">S</emphasis>
. 
<emphasis box="[772,889,1580,1606]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
</taxonomicName>
on a pair of Kr̂yer specimens from 
<collectingRegion box="[1311,1432,1580,1606]" country="Chile" name="Valparaiso" pageId="1" pageNumber="252">Valparaíso</collectingRegion>
, 
<collectingCountry box="[151,214,1616,1642]" name="Chile" pageId="1" pageNumber="252">Chile</collectingCountry>
, which may be the same specimens that Kr̂yer used as the basis for the figures of 
<taxonomicName box="[1143,1261,1616,1643]" class="Malacostraca" family="Bopyridae" genus="Bopyrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[1143,1160,1617,1642]" italics="true" pageId="1" pageNumber="252">B</emphasis>
. 
<emphasis box="[1174,1261,1616,1642]" italics="true" pageId="1" pageNumber="252">foliosus</emphasis>
</taxonomicName>
, although there are some slight differences that make this not completely certain. As 
<taxonomicName box="[933,1262,1652,1679]" class="Malacostraca" family="Bopyridae" genus="Stegophrixus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[933,1079,1652,1678]" italics="true" pageId="1" pageNumber="252">Stegophrixus</emphasis>
(
<emphasis box="[1097,1128,1653,1678]" italics="true" pageId="1" pageNumber="252">sic</emphasis>
) 
<emphasis box="[1145,1262,1652,1678]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
</taxonomicName>
/ 
<taxonomicName authorityName="Nierstrasz &amp; Brender a Brandis" authorityYear="1931" class="Malacostraca" family="Bopyridae" genus="Stegophryxus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis italics="true" pageId="1" pageNumber="252">Stegophryxus thompsoni</emphasis>
</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Nierstrasz &amp; Brender a Brandis" baseAuthorityYear="1931" box="[290,539,1688,1714]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[290,539,1688,1714]" italics="true" pageId="1" pageNumber="252">Anathelges thompsoni</emphasis>
</taxonomicName>
has only been mentioned in the literature a few times subsequent to the original description (
<bibRefCitation author="Markham, J. C." box="[293,487,1724,1750]" pageId="1" pageNumber="252" pagination="33 - 41" refId="ref13596" refString="Markham, J. C. (1974 b) Parasitic bopyrid isopods of the amphi-American genus Stegophryxus Thompson with the description of a new species from California. Bulletin of the Southern California Academy of Sciences, 73 (1), 33 - 41." type="journal article" year="1974">Markham 1974b</bibRefCitation>
; 
<bibRefCitation author="Boyko, C. B. &amp; Williams, J. D." box="[501,788,1724,1751]" pageId="1" pageNumber="252" pagination="795 - 813" refId="ref10782" refString="Boyko, C. B. &amp; Williams, J. D. (2003) A revision of Anathelges and Stegophryxus (Isopoda: Bopyridae: Athelginae) with descriptions of two new genera and one new species. Journal of Crustacean Biology, 23 (4), 795 - 813. https: // doi. org / 10.1651 / C- 2398" type="journal article" year="2003">Boyko &amp; Williams 2003</bibRefCitation>
; 
<bibRefCitation author="Markham, J. C." box="[803,980,1724,1751]" pageId="1" pageNumber="252" pagination="71 - 77" refId="ref13773" refString="Markham, J. C. (2003) A worldwide list of hermit crabs and their relatives (Anomura: Paguroidea) reported as hosts of Isopoda Bopyridae. In: Lemaitre, R. &amp; Tudge, C. C. (Eds.), Biology of the Anomura. Proceedings of a Symposium at the Fifth International Crustacean Congress, Melbourne, Australia, 9 - 13 July 2001. Memoirs of Museum Victoria, 60 (1), pp. 71 - 77. https: // doi. org / 10.24199 / j. mmv. 2003.60.10" type="journal article" year="2003">Markham 2003</bibRefCitation>
; 
<bibRefCitation author="Diaz, M. V. &amp; Roccatagliata, D." box="[995,1312,1724,1751]" pageId="1" pageNumber="252" pagination="331 - 340" refId="ref11826" refString="Diaz, M. V. &amp; Roccatagliata, D. (2006) Remarks on the genus Anathelges (Isopoda: Bopyridae), with a new record from the Beagle Channel, Argentina. Journal of Crustacean Biology, 26 (3), 331 - 340. https: // doi. org / 10.1651 / C- 2593.1" type="journal article" year="2006">Diaz &amp; Roccatagliata 2006</bibRefCitation>
; EspinosaPérez &amp; Hendrickx 2006; 
<bibRefCitation author="Pardo, L. M. &amp; Boyko, C. B. &amp; Mantelatto, F." box="[454,654,1760,1787]" pageId="1" pageNumber="252" pagination="2041 - 2055" refId="ref14444" refString="Pardo, L. M., Boyko, C. B. &amp; Mantelatto, F. (2009) Description of a new species of Asymmetrione (Isopoda: Bopyridae: Pseudioninae) infesting the hermit crab Paguristes tomentosus (Anomura: Diogenidae) from Peru, with a key to species and a review of southeastern Pacific bopyrids. Journal of Natural History, 43 (33 - 34), 2041 - 2055. https: // doi. org / 10.1080 / 00222930903094639" type="journal article" year="2009">
Pardo 
<emphasis box="[528,586,1760,1786]" italics="true" pageId="1" pageNumber="252">et al.</emphasis>
2009
</bibRefCitation>
; 
<bibRefCitation author="McDermott, J. J. &amp; Williams, J. D. &amp; Boyko, C. B." box="[668,933,1760,1787]" pageId="1" pageNumber="252" pagination="2 - 44" refId="ref13972" refString="McDermott, J. J., Williams, J. D. &amp; Boyko, C. B. (2010) The unwanted guests of hermits: a worldwide review of the diversity and natural history of hermit crab parasites. Journal of Experimental Marine Biology, 394 (1 - 2), 2 - 44. https: // doi. org / 10.1016 / j. jembe. 2010.06.022" type="journal article" year="2010">
McDermott 
<emphasis box="[808,866,1760,1786]" italics="true" pageId="1" pageNumber="252">et al.</emphasis>
2010
</bibRefCitation>
) and new material of this species has never been reported, there is no issue in recognizing 
<taxonomicName box="[684,803,1796,1823]" class="Malacostraca" family="Bopyridae" genus="Bopyrus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[684,701,1797,1822]" italics="true" pageId="1" pageNumber="252">B</emphasis>
. 
<emphasis box="[716,803,1796,1822]" italics="true" pageId="1" pageNumber="252">foliosus</emphasis>
</taxonomicName>
as the senior synonym of 
<taxonomicName authority="Nierstrasz &amp; Brender" authorityName="Nierstrasz &amp; Brender" class="Malacostraca" family="Bopyridae" genus="Stegophiyxus" isUncertain="true" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="thompsoni">
<emphasis box="[1105,1253,1796,1822]" italics="true" pageId="1" pageNumber="252">Stegophiyxus</emphasis>
(
<emphasis box="[1271,1302,1797,1822]" italics="true" pageId="1" pageNumber="252">sic</emphasis>
) 
<emphasis box="[1320,1437,1796,1822]" italics="true" pageId="1" pageNumber="252">thompsoni</emphasis>
Nierstrasz &amp; Brender
</taxonomicName>
à Brandis, 1931, making 
<taxonomicName box="[678,897,1832,1858]" class="Malacostraca" family="Bopyridae" genus="Anathelges" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="foliosus">
<emphasis box="[678,897,1832,1858]" italics="true" pageId="1" pageNumber="252">Anathelges foliosus</emphasis>
</taxonomicName>
(Kr̂yer in Gaimard, [1842]) 
<emphasis bold="true" box="[1230,1332,1833,1859]" pageId="1" pageNumber="252">n. comb.</emphasis>
the valid name for the species.
</paragraph>
</subSubSection>
</treatment>
</document>