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<document id="E16B16AEE05A1BABF4D4749BF7A986EE" ID-CLB-Dataset="31448" ID-DOI="10.11646/zootaxa.4085.1.1" ID-GBIF-Dataset="429cf011-ec4c-4a61-b894-2bd32b2ffc9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052381" ID-ZooBank="7F819439-FD77-4EF2-8E26-7B46F9B5FAAB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510872834225" checkinUser="plazi" docAuthor="Golovatch, Sergei, Evsyukov, Aleksandr &amp; Reip, Hans" docDate="2016" docId="03D23F44F6189D31FF075774FB36C006" docLanguage="en" docName="zootaxa.4085.1.1.pdf" docOrigin="Zootaxa 4085 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Picea (Nomarchus) mediterraneus Daday 1889" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FFEB473CF6129D3BFF905518FFA3C500" masterDocTitle="The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="11" updateTime="1698498788515" updateUser="ExternalLinkService">
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<mods:title id="40E05B974517C95856938184BB5D1FD2">The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)</mods:title>
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<mods:namePart id="0B00A458806FF831B6AEA108CDB284C7">Evsyukov, Aleksandr</mods:namePart>
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<mods:namePart id="FA83A85D44D26F9EC49DC486EF26C262">Reip, Hans</mods:namePart>
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<treatment id="03D23F44F6189D31FF075774FB36C006" ID-DOI="http://doi.org/10.5281/zenodo.6079491" ID-GBIF-Taxon="137308727" ID-Zenodo-Dep="6079491" LSID="urn:lsid:plazi:treatment:03D23F44F6189D31FF075774FB36C006" httpUri="http://treatment.plazi.org/id/03D23F44F6189D31FF075774FB36C006" lastPageNumber="11" pageId="10" pageNumber="11">
<subSubSection id="C361DDD9F6189D31FF075774FE8CC7A7" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BC48E52F6189D31FF075774FC86C786" blockId="10.[151,805,620,679]" box="[151,805,620,646]" pageId="10" pageNumber="11">
<heading id="D08C393EF6189D31FF075774FC86C786" bold="true" box="[151,805,620,646]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B90F5240F6189D31FF075774FC86C786" bold="true" box="[151,805,620,646]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF075774FE85C786" bold="true" box="[151,294,620,646]" italics="true" pageId="10" pageNumber="11">Polydesmus</emphasis>
<taxonomicName id="4C7BF5D1F6189D31FEBD5774FC86C786" authority="Daday, 1889" authorityName="Daday" authorityYear="1889" box="[301,805,620,646]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="mediterraneus" subGenus="Nomarchus">
(
<emphasis id="B90F5240F6189D31FEA75774FE66C786" bold="true" box="[311,453,620,646]" italics="true" pageId="10" pageNumber="11">Nomarchus</emphasis>
)
<emphasis id="B90F5240F6189D31FE465774FD27C786" bold="true" box="[470,644,620,646]" italics="true" pageId="10" pageNumber="11">mediterraneus</emphasis>
Daday, 1889
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC48E52F6189D31FF075796FE8CC7A7" blockId="10.[151,805,620,679]" box="[151,303,654,679]" pageId="10" pageNumber="11">
<figureCitation id="134092D7F6189D31FF075796FF79C7A7" box="[151,218,654,679]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
, Map 1
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F6189D31FF0757CEFCB9C7EC" box="[151,794,726,749]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BC48E52F6189D31FF0757CEFCB9C7EC" blockId="10.[151,794,726,749]" box="[151,794,726,749]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB6BA97CF6189D31FF0757CEFCB9C7EC" box="[151,794,726,749]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF0757CEFDB5C7ED" box="[151,534,726,749]" italics="true" pageId="10" pageNumber="11">
Polydesmus
<taxonomicName id="4C7BF5D1F6189D31FE8657CEFE0BC7EC" authorityName="Daday" authorityYear="1889" box="[278,424,726,748]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="mediterraneus">mediterraneus</taxonomicName>
oertzeni—
</emphasis>
<treatmentCitation id="0ADAA843F6189D31FD8757CEFD48C7EC" author="Talikadze" box="[535,747,726,748]" page="143" pageId="10" pageNumber="11" year="1984">
<bibRefCitation id="EFEAF3A3F6189D31FD8757CEFD18C7EC" author="Talikadze" box="[535,699,726,748]" pageId="10" pageNumber="11" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii Zhurnal, 63 (1), 124 - 145. [in Russian, a summary in English]" type="journal article" year="1984">Talikadze, 1984</bibRefCitation>
: 143
</treatmentCitation>
(R).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F6189D31FF075601FCF9C19A" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BC48E52F6189D31FF075601FC35C67A" blockId="10.[151,1436,793,1286]" pageId="10" pageNumber="11">
<materialsCitation id="3B13840FF6189D31FF075601FE5FC656" ID-GBIF-Occurrence="1701200769" collectingDate="1986-03-29" collectionCode="ZMUM" collectorName="A. Markossian" location="Yashtukha" municipality="Sukhumi Distr." pageId="10" pageNumber="11" specimenCount="12" specimenCount-female="9" specimenCount-male="3" stateProvince="ABKHAZIA">
<emphasis id="B90F5240F6189D31FF075601FE2BC632" bold="true" box="[151,392,793,818]" pageId="10" pageNumber="11">Material examined.</emphasis>
<specimenCount id="9D7D45DBF6189D31FE065602FE54C632" box="[406,503,793,818]" pageId="10" pageNumber="11" type="male">3 males</specimenCount>
,
<specimenCount id="9D7D45DBF6189D31FD9A5602FDDEC632" box="[522,637,793,818]" pageId="10" pageNumber="11" type="female">9 females</specimenCount>
(
<collectionCode id="ED6A1697F6189D31FD045602FD52C632" box="[660,753,794,818]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="11">ZMUM</collectionCode>
),
<collectingRegion id="49BF40B0F6189D31FC9B5601FC00C632" box="[779,931,793,818]" country="Georgia" name="Abkhazia" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FC9B5601FC00C632" bold="true" box="[779,931,793,818]" pageId="10" pageNumber="11">ABKHAZIA</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6BA01428F6189D31FC295601FBCDC632" box="[953,1134,793,818]" pageId="10" pageNumber="11">Sukhumi Distr.</collectingMunicipality>
,
<location id="8EA4D889F6189D31FB105601FB5BC632" LSID="urn:lsid:plazi:treatment:03D23F44F6189D31FF075774FB36C006:8EA4D889F6189D31FB105601FB5BC632" box="[1152,1272,793,818]" municipality="Sukhumi Distr." name="Yashtukha" pageId="10" pageNumber="11" stateProvince="ABKHAZIA">Yashtukha</location>
, tree nursery,
<collectingDate id="EF81517AF6189D31FF075625FEB4C656" box="[151,279,829,854]" pageId="10" pageNumber="11" value="1986-03-29">
<date id="FFC5A892F6189D31FF075625FEB4C656" box="[151,279,829,854]" pageId="10" pageNumber="11" value="1986-03-29">29.III.1986</date>
</collectingDate>
, leg.
<collectorName id="268EEB84F6189D31FEC35626FE5FC656" box="[339,508,829,854]" pageId="10" pageNumber="11">A. Markossian</collectorName>
</materialsCitation>
;
<materialsCitation id="3B13840FF6189D31FD975626FAE6C656" ID-GBIF-Occurrence="1701200782" box="[519,1349,829,854]" collectingDate="1978-10-02" collectionCode="ZMUM" location="Sukhumi Botanical Garden" pageId="10" pageNumber="11" specimenCount="2" specimenCount-subadult="2" stateProvince="ABKHAZIA">
<specimenCount id="9D7D45DBF6189D31FD975626FDD8C656" box="[519,635,829,854]" pageId="10" pageNumber="11" type="subadult">2 subadult</specimenCount>
females (
<collectionCode id="ED6A1697F6189D31FD785626FCE6C656" box="[744,837,830,854]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="11">ZMUM</collectionCode>
),
<location id="8EA4D889F6189D31FCC85625FB2EC656" LSID="urn:lsid:plazi:treatment:03D23F44F6189D31FF075774FB36C006:8EA4D889F6189D31FCC85625FB2EC656" box="[856,1165,829,854]" name="Sukhumi Botanical Garden" pageId="10" pageNumber="11" stateProvince="ABKHAZIA">Sukhumi Botanical Garden</location>
, litter,
<collectingDate id="EF81517AF6189D31FB4A5626FAE6C656" box="[1242,1349,830,854]" pageId="10" pageNumber="11" value="1978-10-02">
<date id="FFC5A892F6189D31FB4A5626FAE6C656" box="[1242,1349,830,854]" pageId="10" pageNumber="11" value="1978-10-02">2.X.1978</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B13840FF6189D31FAC15626FC35C67A" ID-GBIF-Occurrence="1701200780" collectingDate="1978-10-09" collectionCode="ZMUM" collectorName="S. Golovatch." location="Sukhumi Botanical Garden" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="ABKHAZIA">
<specimenCount id="9D7D45DBF6189D31FAC15626FA3FC656" box="[1361,1436,829,854]" pageId="10" pageNumber="11" type="male">1 male</specimenCount>
(slide only) (
<collectionCode id="ED6A1697F6189D31FEB6567AFE20C67A" box="[294,387,866,890]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="11">ZMUM</collectionCode>
), same locality
<collectingDate id="EF81517AF6189D31FDA1567AFD3EC67A" box="[561,669,866,890]" pageId="10" pageNumber="11" value="1978-10-09">
<date id="FFC5A892F6189D31FDA1567AFD3EC67A" box="[561,669,866,890]" pageId="10" pageNumber="11" value="1978-10-09">9.X.1978</date>
</collectingDate>
; all leg.
<collectorName id="268EEB84F6189D31FD6C567AFC35C67A" box="[764,918,865,890]" pageId="10" pageNumber="11">S. Golovatch.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BC48E52F6189D31FF57569DFEC3C6E6" blockId="10.[151,1436,793,1286]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF57569DFE38C69E" bold="true" box="[199,411,901,926]" pageId="10" pageNumber="11">Descriptive notes.</emphasis>
Length ca
<quantity id="4C8323B7F6189D31FD89569EFD25C69D" box="[537,646,901,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="10" pageNumber="11" unit="mm" value="12.0" valueMax="16.0" valueMin="8.0">
8
<emphasis id="B90F5240F6189D31FDB6569DFD97C69E" bold="true" box="[550,564,901,926]" pageId="10" pageNumber="11"></emphasis>
16 mm
</quantity>
, width of midbody metazonae 1.5
<emphasis id="B90F5240F6189D31FB94569DFBB1C69E" bold="true" box="[1028,1042,901,926]" pageId="10" pageNumber="11"></emphasis>
2.0 mm (male, female), metazonite to prozonite width ratio &lt;1.6 (
<figureCitation id="134092D7F6189D31FE6156B1FDEFC6C2" box="[497,588,937,962]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Live coloration usually uniformly light brown to brown, about same as in alcohol (
<figureCitation id="134092D7F6189D31FF6956D5FEF0C6E6" box="[249,339,973,998]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BC48E52F6189D31FF5756E9FCF9C19A" blockId="10.[151,1436,793,1286]" pageId="10" pageNumber="11">
All characters as in
<taxonomicName id="4C7BF5D1F6189D31FE3E56EAFD9FC10A" authorityName="Attems" authorityYear="1898" box="[430,572,1009,1034]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="abchasius">
<emphasis id="B90F5240F6189D31FE3E56EAFD9FC10A" box="[430,572,1009,1034]" italics="true" pageId="10" pageNumber="11">P. abchasius</emphasis>
</taxonomicName>
(
<figureCitation id="134092D7F6189D31FDDD56E9FD30C10A" box="[589,659,1009,1034]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
), except as follows. Collum much narrower than head, transversely oval in shape (
<figureCitation id="134092D7F6189D31FEAB510DFE36C12E" box="[315,405,1045,1070]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Paraterga not upturned, subhorizontal, much more strongly rounded laterally, at least until segment 9 obtuse-angled and drawn behind rear tergal margin starting with segment 14 or 15 (
<figureCitation id="134092D7F6189D31FB7D5121FAE8C152" box="[1261,1355,1081,1106]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Male prefemora strongly bulging laterad (
<figureCitation id="134092D7F6189D31FDBA5145FD22C176" box="[554,641,1117,1142]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5B</figureCitation>
). Gonopod (
<figureCitation id="134092D7F6189D31FC9C5145FCC1C176" box="[780,866,1117,1142]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
) with a stout and stump-shaped endomere (
<emphasis id="B90F5240F6189D31FAD85145FAC7C176" bold="true" box="[1352,1380,1117,1142]" pageId="10" pageNumber="11">en</emphasis>
) and only a slightly curved and apically strongly bifid exomere (
<emphasis id="B90F5240F6189D31FCA05199FCE9C19A" bold="true" box="[816,842,1153,1178]" pageId="10" pageNumber="11">ex</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F6189D31FF5751BDFB36C006" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BC48E52F6189D31FF5751BDFB36C006" blockId="10.[151,1436,793,1286]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF5751BDFE98C1BE" bold="true" box="[199,315,1189,1214]" pageId="10" pageNumber="11">Remarks.</emphasis>
The only previous record in the region (
<bibRefCitation id="EFEAF3A3F6189D31FC9A51BDFC19C1BE" author="Talikadze" box="[778,954,1189,1214]" pageId="10" pageNumber="11" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii Zhurnal, 63 (1), 124 - 145. [in Russian, a summary in English]" type="journal article" year="1984">Talikadze 1984</bibRefCitation>
) concerned “
<collectingRegion id="49BF40B0F6189D31FBC051BDFB61C1BE" box="[1104,1218,1189,1214]" country="Georgia" name="Abkhazia" pageId="10" pageNumber="11">Abkhazia</collectingRegion>
”, without a precise locality. This eastern Mediterranean species/subspecies is quite common in the adjacent
<collectingRegion id="49BF40B0F6189D31FBEE51D1FB77C1E2" box="[1150,1236,1225,1250]" country="Ukraine" name="Crimea" pageId="10" pageNumber="11">Crimea</collectingRegion>
, apparently being introduced to the Caucasus. New illustrations are provided to facilitate recognition (
<figureCitation id="134092D7F6189D31FBD351F5FB24C006" box="[1091,1159,1261,1286]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>