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<mods:title id="74072A5BDBB83D54181BC1BA3F1F397D">A revision of the rare flower beetle genus Macronotops Krikken (Coleoptera: Scarabaeidae: Cetoniinae) from Asia with biological notes</mods:title>
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<treatment id="03CA3425FFB5A40EC2E9FFA9155A1128" ID-DOI="http://doi.org/10.5281/zenodo.5934019" ID-GBIF-Taxon="152633815" ID-Zenodo-Dep="5934019" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128" httpUri="http://treatment.plazi.org/id/03CA3425FFB5A40EC2E9FFA9155A1128" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
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<taxonomicName id="4C63FEB0FFB5A40CC2E9FFA91661128F" ID-CoL="3X78Z" authority="(Bourgoin, 1916)" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[151,618,151,178]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
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<emphasis id="B9175921FFB5A40CC2E9FFA9159A128C" bold="true" box="[151,401,151,177]" italics="true" pageId="37" pageNumber="38">Macronotops vuilleti</emphasis>
(
<bibRefCitation id="EFF2F8C2FFB5A40CC3DEFFA61669128F" author="Bourgoin, A." box="[416,610,152,178]" pageId="37" pageNumber="38" pagination="133 - 137" refId="ref34313" refString="Bourgoin, A. (1916) Description de trois Macronota nouveaux (Col. Scarabaeidae). Bulletin de la Societe Entomologique de France Paris, 1916 (7), 133 - 137." type="journal article" year="1916">Bourgoin, 1916</bibRefCitation>
)
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<paragraph id="8BDC8533FFB5A40CC2E9FF8410C212EE" blockId="37.[151,1225,151,211]" box="[151,1225,186,211]" pageId="37" pageNumber="38">
(
<figureCitation id="135899B6FFB5A40CC2E1FF84152C12EE" box="[159,295,186,211]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="37" pageNumber="38">Figs. 1617</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC34CFF85157112EE" box="[306,378,187,211]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="37" pageNumber="38">3940</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC3F8FF8415C412EE" box="[390,463,186,211]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="37" pageNumber="38">6364</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC3A4FF85162812EE" box="[474,547,187,211]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="37" pageNumber="38">9497</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC051FF85165012EE" box="[559,603,187,211]" captionStart="FIGURES 111122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="37" pageNumber="38">119</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC01BFF85169A12EE" box="[613,657,187,211]" captionStart="FIGURES 122134" captionStartId="17.[151,264,1757,1780]" captionTargetBox="[168,1424,207,1720]" captionTargetId="figure@17.[151,1436,193,1737]" captionTargetPageId="17" captionText="FIGURES 122134. Male genitalia of Macronotops (lateral view). 123, M. biserratus new species; 124, M. dianensis new species; 125, M. fulvoguttatus; 126, M. fulvopilosus; 127, M. miksici new species; 128, M. nigropubescens; 129, M. olivaceofuscus; 130, M. sexmaculatus; 131, M. vuilleti; 132, M. curvimarginatus; 133, M. medogensis new species; 134, M. ovaliceps." httpUri="https://zenodo.org/record/2585222/files/figure.png" pageId="37" pageNumber="38">131</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC0E3FF84170912EE" box="[669,770,186,211]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">169172</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC173FF85177912EE" box="[781,882,187,211]" captionStart="FIGURES 215226" captionStartId="51.[151,264,1940,1963]" captionTargetBox="[165,1421,187,1916]" captionTargetId="figure@51.[151,1436,181,1920]" captionTargetPageId="51" captionText="FIGURES 215226. Habitus of Macronotops (male in left, female in right; oblique view). 215216, M. olivaceofuscus; 217 218, M. sexmaculatus; 219220, M. vuilleti; 221222, M. curvimarginatus; 223224, M. medogensis new species; 225226, M. ovaliceps." httpUri="https://zenodo.org/record/2585256/files/figure.png" pageId="37" pageNumber="38">219220</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC103FF8517E912EE" box="[893,994,187,211]" captionStart="FIGURES 235242" captionStartId="53.[151,264,1260,1283]" captionTargetBox="[186,1419,199,1224]" captionTargetId="figure@53.[151,1436,193,1239]" captionTargetPageId="53" captionText="FIGURES 235242. Type specimens and labels of Macronotops. 235237, M. olivaceofuscus, male holotype (photograph by Enrico Ruzzier); 238242, M. vuilleti, male holotype (photograph by Maxwell Barclay)." httpUri="https://zenodo.org/record/2585260/files/figure.png" pageId="37" pageNumber="38">238242</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC193FF84101112EE" box="[1005,1050,186,211]" captionStart="FIGURES 258260" captionStartId="55.[151,264,1245,1268]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@55.[151,1436,193,1224]" captionTargetPageId="55" captionText="FIGURES 258260. Known distribution of Macronotops. 258, Distribution for the whole genus; 259, map of SE Xizang, China; 260, map of W Sichuan, China. Syncretic markers indicate identical or very close localities; type localities of the known species are underlined, that of M. olivaceofuscus is missing because the precise type locality is unknown." httpUri="https://zenodo.org/record/2585264/files/figure.png" pageId="37" pageNumber="38">258</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC65BFF85108112EE" box="[1061,1162,187,211]" captionStart="FIGURES 276281" captionStartId="58.[151,264,1812,1835]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@58.[151,1436,193,1792]" captionTargetPageId="58" captionText="FIGURES 276281. Habitat of Macronotops. 276277, habitat of M. sexmaculatus and M. medogensis new species in Valley near Bononggong of Mêdog (Xizang, China); 278, an adult of M. sexmaculatus feeding on sap of bamboo (photograph by Yang He); 279281, habitat of M. vuilleti in Mount Huanglianshan (Yunnan, China), and larva in fallen rotten wood." httpUri="https://zenodo.org/record/2585270/files/figure.png" pageId="37" pageNumber="38">279281</figureCitation>
,
<figureCitation id="135899B6FFB5A40CC6EBFF8510CB12EE" box="[1173,1216,187,211]" captionStart="FIGURES 295309" captionStartId="61.[151,264,1775,1798]" captionTargetBox="[151,1436,193,1755]" captionTargetId="figure@61.[151,1436,193,1755]" captionTargetPageId="61" captionText="FIGURES 295309. Living adults and pupae of the Macronotops sexmaculatus species group in artificial conditions. 295297, adult of M. dianensis new species feeding on banana and newly emerged adults in pupal cells; 298300, adult of M. fulvopilosus feeding on banana and newly emerged adults in pupal cells; 301302, pupa of M. fulvoguttatus and newly emerged adult in pupal cell; 303304, newly emerged adults of M. miksici new species in pupal cells; 305308, pupae of M. olivaceofuscus and newly emerged adults in pupal cells; 309, newly emerged adults of M. vuilleti in pupal cells." httpUri="https://zenodo.org/record/2585276/files/figure.png" pageId="37" pageNumber="38">309</figureCitation>
)
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<treatmentCitationGroup id="AB73A21DFFB5A40CC2E9FE3D11901325" box="[151,1435,258,280]" pageId="37" pageNumber="38">
<taxonomicName id="4C63FEB0FFB5A40CC2E9FE3D16251325" ID-CoL="3X78H" authority="Bourgoin, 1916: 135" authorityName="Bourgoin" authorityPageNumber="135" authorityYear="1916" box="[151,558,258,280]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB5A40CC2E9FE3D15581325" box="[151,339,258,280]" italics="true" pageId="37" pageNumber="38">Macronota vuilleti</emphasis>
<treatmentCitation id="0AC2A322FFB5A40CC325FE3C16251325" author="Bourgoin, A." box="[347,558,258,280]" page="135" pageId="37" pageNumber="38" year="1916">
<bibRefCitation id="EFF2F8C2FFB5A40CC325FE3C15F61325" author="Bourgoin, A." box="[347,509,258,280]" pageId="37" pageNumber="38" pagination="133 - 137" refId="ref34313" refString="Bourgoin, A. (1916) Description de trois Macronota nouveaux (Col. Scarabaeidae). Bulletin de la Societe Entomologique de France Paris, 1916 (7), 133 - 137." type="journal article" year="1916">Bourgoin, 1916</bibRefCitation>
: 135
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</taxonomicName>
(
<typeStatus id="54D83B91FFB5A40CC042FE3C16621325" box="[572,617,258,280]" pageId="37" pageNumber="38">type</typeStatus>
locality: Chapa, Lao-Kay,
<collectingCountry id="F374C5A3FFB5A40CC103FE3C17CF1325" box="[893,964,258,280]" name="Vietnam" pageId="37" pageNumber="38">Tonkin</collectingCountry>
[=Sa Pa,
<collectingRegion id="49A74BD1FFB5A40CC654FE3D10761325" box="[1066,1149,258,280]" country="Vietnam" name="Lao Cai" pageId="37" pageNumber="38">Lao Cai</collectingRegion>
, N.
<collectingCountry id="F374C5A3FFB5A40CC6D6FE3C110C1325" box="[1192,1287,258,280]" name="Vietnam" pageId="37" pageNumber="38">Vietnam</collectingCountry>
])*;
<bibRefCitation id="EFF2F8C2FFB5A40CC757FE3C11901325" author="Schenkling, S." box="[1321,1435,258,280]" pageId="37" pageNumber="38" pagination="1 - 431" refId="ref35805" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III Pars. 72. W. Junk, Berlin, pp. 1 - 431." type="book chapter" year="1921">Schenkling</bibRefCitation>
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1921: 143.
<taxonomicName id="4C63FEB0FFB5A40CC2E9FE7E15CF136B" authority="(Bourgoin)" baseAuthorityName="Bourgoin" box="[151,452,320,342]" class="Insecta" family="Scarabaeidae" genus="Coilodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB5A40CC2E9FE7E1541136B" box="[151,330,320,342]" italics="true" pageId="37" pageNumber="38">Coilodera vuilleti</emphasis>
(Bourgoin)
</taxonomicName>
:
<bibRefCitation id="EFF2F8C2FFB5A40CC3B3FE7E165D136B" author="Paulian, R." box="[461,598,320,342]" pageId="37" pageNumber="38" pagination="1 - 87" refId="ref35514" refString="Paulian, R. (1960) Coleopteres Scarabeides de L'Indochine (Rutelines et Cetonines) (Suite). Annales de la Societe Entomologique de France, 129, 1 - 87." type="journal article" year="1960">Paulian 1960</bibRefCitation>
: 18.
<taxonomicName id="4C63FEB0FFB5A40CC2E9FE6015C41349" authority="(Bourgoin)" baseAuthorityName="Bourgoin" box="[151,463,350,372]" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB5A40CC2E9FE60155F1349" box="[151,340,350,372]" italics="true" pageId="37" pageNumber="38">Pleuronota vuilleti</emphasis>
(Bourgoin)
</taxonomicName>
:
<bibRefCitation id="EFF2F8C2FFB5A40CC3A6FE6016501349" author="Miksic, R." box="[472,603,350,372]" pageId="37" pageNumber="38" pagination="208 - 210" refId="ref35268" refString="Miksic, R. (1971) Notizien uber ostasiatische Cetoniiden. (72. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique, Conseil des Academies de la RPF Yougoslavie. Section A. Sciences Naturelles, Techniques et Medicales, 16 (7 - 8), 208 - 210." type="journal article" year="1971">Mikšić 1971</bibRefCitation>
: 208;
<bibRefCitation id="EFF2F8C2FFB5A40CC0E6FE6017101349" author="Miksic, R." box="[664,795,350,372]" pageId="37" pageNumber="38" refId="ref35365" refString="Miksic, R. (1976) Monographie der Cetonniiae der Palaarktischen und Orientalischen Region. Coleoptera: Lamellicornia. Band 1. Institut za sumarstvo, Sarajevo, 444 pp." type="book" year="1976">Mikšić 1976</bibRefCitation>
: 170 (
<typeStatus id="54D83B91FFB5A40CC126FE6017BC1349" box="[856,951,350,372]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
).
<taxonomicName id="4C63FEB0FFB5A40CC2E9FE40163A13AE" ID-CoL="3X78Z" authority="vuilleti (Bourgoin)" authorityName="vuilleti (Bourgoin" box="[151,561,381,403]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="vuilleti" subSpecies="vuilleti">
<emphasis id="B9175921FFB5A40CC2E9FE4015BD13AE" box="[151,438,381,403]" italics="true" pageId="37" pageNumber="38">Macronotops vuilleti vuilleti</emphasis>
(Bourgoin)
</taxonomicName>
:
<bibRefCitation id="EFF2F8C2FFB5A40CC045FE4316C113AE" author="Krikken, J." box="[571,714,381,403]" pageId="37" pageNumber="38" pagination="199 - 209" refId="ref34797" refString="Krikken, J. (1977) The Asian genus Pleuronota Kraatz and allied forms: a clarification (Coleoptera: Cetoniidae). Zoologische Mededelingen, 51 (13), 199 - 209." type="journal article" year="1977">Krikken 1977</bibRefCitation>
: 208;
<bibRefCitation id="EFF2F8C2FFB5A40CC176FE43179A13AE" author="Krajcik, M." box="[776,913,381,403]" pageId="37" pageNumber="38" refId="ref34717" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
: 90;
<bibRefCitation id="EFF2F8C2FFB5A40CC1BCFE43103C13AE" author="Sakai, K." box="[962,1079,381,403]" pageId="37" pageNumber="38" pagination="11 - 22" refId="ref35646" refString="Sakai, K. (2008) Record of Cetoniid beetles from the central and northern Laos (Coleoptera, Scarabaeidae). Saikaku-Tsushin, 16, 11 - 22. [in Japanese]" type="journal article" year="2008">Sakai 2008</bibRefCitation>
: 20 (Mount Phu Pan &amp; Xam Neua,
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB5A40CC2B9FEA216F2138F" box="[199,761,412,434]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BDC8533FFB5A40CC2B9FEA216F2138F" blockId="37.[151,1436,258,434]" box="[199,761,412,434]" pageId="37" pageNumber="38">
<collectingCountry id="F374C5A3FFB5A40CC2B9FEA214F6138F" box="[199,253,412,434]" name="Laos" pageId="37" pageNumber="38">Laos</collectingCountry>
)*, fig.
<specimenCount id="9D654EBAFFB5A40CC33DFEA2157C138C" box="[323,375,412,433]" pageId="37" pageNumber="38" type="male">72 ♂</specimenCount>
;
<bibRefCitation id="EFF2F8C2FFB5A40CC3FDFEA21601138F" author="Krajcik, M." box="[387,522,412,434]" pageId="37" pageNumber="38" pagination="1 - 113" refId="ref34739" refString="Krajcik, M. (2011) Illustrated catalogue of Cetoniinae, Trichiinae and Valginae of China (Coleoptera, Cetoniidae). Animma. x, Supplement, 1 - 113." type="book chapter" year="2011">Krajčík 2011</bibRefCitation>
: 59;
<bibRefCitation id="EFF2F8C2FFB5A40CC044FEA216CF138F" author="Krajcik, M." box="[570,708,412,434]" pageId="37" pageNumber="38" pagination="1 - 278" refId="ref34772" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">Krajčík 2012</bibRefCitation>
: 152.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB5A40CC2E9FEE017E51100" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BDC8533FFB5A40CC2E9FEE010861002" blockId="37.[151,1437,478,1982]" pageId="37" pageNumber="38">
<emphasis id="B9175921FFB5A40CC2E9FEE015B613CA" bold="true" box="[151,445,478,503]" pageId="37" pageNumber="38">
Type material.
<materialsCitation id="3B0B8F6EFFB5A40CC32EFEE015B613CA" ID-GBIF-Occurrence="2004970386" box="[336,445,478,503]" collectionCode="BMNH" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D83B91FFB5A40CC32EFEE015B613CA" box="[336,445,478,503]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
(by monotypy,
<specimenCount id="9D654EBAFFB5A40CC008FEE1168613CB" box="[630,653,479,502]" pageId="37" pageNumber="38" type="male"></specimenCount>
, BMNH,
<figureCitation id="135899B6FFB5A40CC17EFEE017A213CA" box="[768,937,478,503]" captionStart="FIGURES 235242" captionStartId="53.[151,264,1260,1283]" captionTargetBox="[186,1419,199,1224]" captionTargetId="figure@53.[151,1436,193,1239]" captionTargetPageId="53" captionText="FIGURES 235242. Type specimens and labels of Macronotops. 235237, M. olivaceofuscus, male holotype (photograph by Enrico Ruzzier); 238242, M. vuilleti, male holotype (photograph by Maxwell Barclay)." httpUri="https://zenodo.org/record/2585260/files/figure.png" pageId="37" pageNumber="38">Figs. 238242</figureCitation>
):
<taxonomicName id="4C63FEB0FFB5A40CC1C3FEE1110D13CA" authority="Bourgoin" authorityName="Bourgoin" authorityYear="1916" box="[957,1286,478,503]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB5A40CC1C3FEE1109B13CA" box="[957,1168,478,503]" italics="true" pageId="37" pageNumber="38">Macronota vuilleti</emphasis>
Bourgoin
</taxonomicName>
, Type [white label with red right edge]//
<materialsCitation id="3B0B8F6EFFB5A40CC3B5FD3C17591002" ID-GBIF-Occurrence="2004970391" collectionCode="B, M, E" determinerName="Bacchus" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D83B91FFB5A40CC3B5FD3C16381026" box="[459,563,514,539]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
[white label with red border] // Bourgoin Coll.,
<collectionCode id="ED721DF6FFB5A40CC62BFD3D10611027" box="[1109,1130,515,538]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="37" pageNumber="38">B</collectionCode>
.
<collectionCode id="ED721DF6FFB5A40CC608FD3D10991027" box="[1142,1170,515,538]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="37" pageNumber="38">M</collectionCode>
. 1938-252 [pale yellow label] //
<taxonomicName id="4C63FEB0FFB5A40CC28AFD1916131002" authority="Bourg." authorityName="Bourgoin" authorityYear="1916" box="[244,536,550,575]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB5A40CC28AFD1915CF1002" box="[244,452,550,575]" italics="true" pageId="37" pageNumber="38">Macronota vuilleti</emphasis>
Bourg.
</taxonomicName>
<specimenCount id="9D654EBAFFB5A40CC061FD19163D1003" box="[543,566,551,574]" pageId="37" pageNumber="38" type="male"></specimenCount>
,
<collectionCode id="ED721DF6FFB5A40CC03CFD1916551003" box="[578,606,551,574]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="37" pageNumber="38">M</collectionCode>
.
<collectionCode id="ED721DF6FFB5A40CC017FD1916761003" box="[617,637,551,574]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="37" pageNumber="38">E</collectionCode>
.
<determinerName id="7E0069F7FFB5A40CC0F6FD1816EC1002" box="[648,743,550,575]" pageId="37" pageNumber="38">Bacchus</determinerName>
det 1974
</materialsCitation>
,
<typeStatus id="54D83B91FFB5A40CC123FD1917FA1002" box="[861,1009,551,575]" pageId="37" pageNumber="38" type="holotype">HOLOTYPE</typeStatus>
[white label].
</paragraph>
<paragraph id="8BDC8533FFB5A40CC2B9FD7417E51100" blockId="37.[151,1437,478,1982]" pageId="37" pageNumber="38">
<emphasis id="B9175921FFB5A40CC2B9FD74163F105E" bold="true" box="[199,564,586,611]" pageId="37" pageNumber="38">Additional material examined</emphasis>
(
<specimenCount id="9D654EBAFFB5A40CC036FD75169A105E" box="[584,657,587,611]" pageId="37" pageNumber="38" type="male">20♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB5A40CC0DEFD7416CF105E" box="[672,708,586,611]" pageId="37" pageNumber="38" type="female">5♀</specimenCount>
).
<materialsCitation id="3B0B8F6EFFB5A40CC0A1FD7417BB105E" ID-GBIF-Occurrence="2004970380" box="[735,944,586,611]" country="China" location="Yunnan" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Yunnan">
<emphasis id="B9175921FFB5A40CC0A1FD741737105E" bold="true" box="[735,828,586,611]" pageId="37" pageNumber="38">
<collectingCountry id="F374C5A3FFB5A40CC0A1FD741737105E" box="[735,828,586,611]" name="China" pageId="37" pageNumber="38">CHINA</collectingCountry>
</emphasis>
:
<emphasis id="B9175921FFB5A40CC130FD7417BB105E" bold="true" box="[846,944,586,611]" pageId="37" pageNumber="38">
<collectingRegion id="49A74BD1FFB5A40CC130FD7417BB105E" box="[846,944,586,611]" country="China" name="Yunnan" pageId="37" pageNumber="38">Yunnan</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B0B8F6EFFB5A40CC1BFFD741084105E" ID-GBIF-Occurrence="2004970381" box="[961,1167,586,611]" collectionCode="QCCC" country="China" pageId="37" pageNumber="38" specimenCount="7" specimenCount-female="1" specimenCount-male="6">
<specimenCount id="9D654EBAFFB5A40CC1BFFD7417F0105E" box="[961,1019,586,611]" pageId="37" pageNumber="38" type="male">6♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB5A40CC675FD751028105E" box="[1035,1059,587,611]" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
(
<collectionCode id="ED721DF6FFB5A40CC652FD75108C105E" box="[1068,1159,587,611]" pageId="37" pageNumber="38">QCCC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B0B8F6EFFB5A40CC6E0FD75119310B7" ID-GBIF-Occurrence="2004970393" collectingDate="2018-02-21" collectionCode="QIU" collectorName="Hao XU" country="China" county="Luchun County" elevation="2" location="Jian-Yue" municipality="Mount Huanglianshan Nature Reserve" pageId="37" pageNumber="38" specimenCount="1">
<collectingMunicipality id="6BB81F49FFB5A40CC6E0FD75154210B4" pageId="37" pageNumber="38">Mount Huanglianshan Nature Reserve</collectingMunicipality>
,
<collectingCounty id="62BDFDBFFFB5A40CC328FD4F160310B7" box="[342,520,625,650]" pageId="37" pageNumber="38">Lüchun County</collectingCounty>
,
<quantity id="4C9B28D6FFB5A40CC06BFD4C167E10B4" box="[533,629,626,650]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="37" pageNumber="38" unit="m" value="1.9">
<elevation id="004E6200FFB5A40CC06BFD4C167E10B4" box="[533,629,626,650]" metricMagnitude="0" metricUnit="m" metricValue="1.9" pageId="37" pageNumber="38" unit="m" value="1.9">1,900 m</elevation>
</quantity>
,
<location id="8EBCD3E8FFB5A40CC0FCFD4F16EF10B7" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC0FCFD4F16EF10B7" box="[642,740,625,650]" country="China" county="Luchun County" municipality="Mount Huanglianshan Nature Reserve" name="Jian-Yue" pageId="37" pageNumber="38">Jian-Yue</location>
<collectionCode id="ED721DF6FFB5A40CC093FD4C171410B7" box="[749,799,626,650]" pageId="37" pageNumber="38">QIU</collectionCode>
&amp;
<collectorName id="2696E0E5FFB5A40CC138FD4C17A210B4" box="[838,937,626,649]" pageId="37" pageNumber="38">Hao XU</collectorName>
, 3
<superScript id="7C16287BFFB5A40CC1BAFD5317DF1046" attach="left" box="[964,980,621,635]" fontSize="6" pageId="37" pageNumber="38">nd</superScript>
instar larvae in rotten wood
<collectingDate id="EF995A1BFFB5A40CC75EFD4C119310B7" box="[1312,1432,626,650]" pageId="37" pageNumber="38" value="2018-02-21">
<date id="FFDDA3F3FFB5A40CC75EFD4C119310B7" box="[1312,1432,626,650]" pageId="37" pageNumber="38" value="2018-02-21">21.II.2018</date>
</collectingDate>
</materialsCitation>
, adult emerged
<date id="FFDDA3F3FFB5A40CC340FDA815991093" box="[318,402,662,686]" pageId="37" pageNumber="38" value="2018-05">V.2018</date>
.
<materialsCitation id="3B0B8F6EFFB5A40CC3E3FDAB172B1093" ID-GBIF-Occurrence="2004970397" box="[413,800,661,686]" collectingDate="2015-05" collectionCode="QCCC, V" country="Vietnam" location="VIETNAM" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
<emphasis id="B9175921FFB5A40CC3E3FDAB16261093" bold="true" box="[413,557,661,686]" pageId="37" pageNumber="38">
<collectingCountry id="F374C5A3FFB5A40CC3E3FDAB16221093" box="[413,553,661,686]" name="Vietnam" pageId="37" pageNumber="38">VIETNAM</collectingCountry>
:
</emphasis>
<specimenCount id="9D654EBAFFB5A40CC04BFDA816531093" box="[565,600,662,686]" pageId="37" pageNumber="38" type="male">1♂</specimenCount>
(
<collectionCode id="ED721DF6FFB5A40CC016FDA816B11093" box="[616,698,662,686]" pageId="37" pageNumber="38">QCCC</collectionCode>
),
<date id="FFDDA3F3FFB5A40CC0B0FDAB172B1093" box="[718,800,661,686]" pageId="37" pageNumber="38" value="2015-05">
<collectingDate id="EF995A1BFFB5A40CC0B0FDAB172B1093" box="[718,800,661,686]" pageId="37" pageNumber="38" value="2015-05">
<collectionCode id="ED721DF6FFB5A40CC0B0FDAB16E81093" box="[718,739,661,686]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="37" pageNumber="38">V</collectionCode>
.2015
</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B0B8F6EFFB5A40CC153FDA810571090" ID-GBIF-Occurrence="2004970400" box="[813,1116,661,686]" country="Vietnam" location="Mount Sapa" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Lao Cai">
<location id="8EBCD3E8FFB5A40CC153FDA817B31093" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC153FDA817B31093" box="[813,952,662,686]" country="Vietnam" name="Mount Sapa" pageId="37" pageNumber="38" stateProvince="Lao Cai">Mount Sapa</location>
,
<collectingRegion id="49A74BD1FFB5A40CC1BAFDA810141093" box="[964,1055,661,686]" country="Vietnam" name="Lao Cai" pageId="37" pageNumber="38">Lao Cai</collectingRegion>
prov
</materialsCitation>
.
<materialsCitation id="3B0B8F6EFFB5A40CC619FDAB14E210EF" ID-GBIF-Occurrence="2004970402" collectingDate="2013-05" collectionCode="QCCC, V" country="Laos" location="LAOS" pageId="37" pageNumber="38" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<emphasis id="B9175921FFB5A40CC619FDAB10B71093" bold="true" box="[1127,1212,661,686]" pageId="37" pageNumber="38">
<collectingCountry id="F374C5A3FFB5A40CC619FDAB10B31093" box="[1127,1208,661,686]" name="Laos" pageId="37" pageNumber="38">LAOS</collectingCountry>
:
</emphasis>
<specimenCount id="9D654EBAFFB5A40CC6BAFDA810F41093" box="[1220,1279,662,686]" pageId="37" pageNumber="38" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB5A40CC775FDA811251093" box="[1291,1326,662,686]" pageId="37" pageNumber="38" type="female">1♀</specimenCount>
(
<collectionCode id="ED721DF6FFB5A40CC740FDA8119B1093" box="[1342,1424,662,686]" pageId="37" pageNumber="38">QCCC</collectionCode>
),
<date id="FFDDA3F3FFB5A40CC2E9FD8414E210EF" box="[151,233,698,722]" pageId="37" pageNumber="38" value="2013-05">
<collectingDate id="EF995A1BFFB5A40CC2E9FD8414E210EF" box="[151,233,698,722]" pageId="37" pageNumber="38" value="2013-05">
<collectionCode id="ED721DF6FFB5A40CC2E9FD8414A710EF" box="[151,172,698,722]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="37" pageNumber="38">V</collectionCode>
.2013
</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B0B8F6EFFB5A40CC28AFD84166310EC" ID-GBIF-Occurrence="2004970384" box="[244,616,697,722]" country="Laos" location="Mount Phu Pane" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Houaphan">
<location id="8EBCD3E8FFB5A40CC28AFD8415BA10EC" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC28AFD8415BA10EC" box="[244,433,697,722]" country="Laos" name="Mount Phu Pane" pageId="37" pageNumber="38" stateProvince="Houaphan">Mount Phu Pane</location>
,
<collectingRegion id="49A74BD1FFB5A40CC3C2FD84162210EF" box="[444,553,697,722]" country="Laos" name="Houaphan" pageId="37" pageNumber="38">Hua Phan</collectingRegion>
prov
</materialsCitation>
.;
<materialsCitation id="3B0B8F6EFFB5A40CC009FD87178A10EF" ID-GBIF-Occurrence="2004970401" box="[631,897,697,722]" collectingDate="2014-06" collectionCode="QCCC, VI" country="Laos" pageId="37" pageNumber="38" specimenCount="5" specimenCount-male="5">
<specimenCount id="9D654EBAFFB5A40CC009FD8716A410EF" box="[631,687,697,722]" pageId="37" pageNumber="38" type="male">5♂♂</specimenCount>
(
<collectionCode id="ED721DF6FFB5A40CC0C3FD84170410EF" box="[701,783,698,722]" pageId="37" pageNumber="38">QCCC</collectionCode>
),
<date id="FFDDA3F3FFB5A40CC15CFD84178A10EF" box="[802,897,698,722]" pageId="37" pageNumber="38" value="2014-06">
<collectingDate id="EF995A1BFFB5A40CC15CFD84178A10EF" box="[802,897,698,722]" pageId="37" pageNumber="38" value="2014-06">
<collectionCode id="ED721DF6FFB5A40CC15CFD84174A10EF" box="[802,833,698,722]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="37" pageNumber="38">VI</collectionCode>
.2014
</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B0B8F6EFFB5A40CC1F2FD84116510EC" ID-GBIF-Occurrence="2004970390" box="[908,1390,697,722]" collectorName="Prov." country="Laos" elevation="2" location="Mount Phu Pane" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Houaphan">
<location id="8EBCD3E8FFB5A40CC1F2FD84104210EC" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC1F2FD84104210EC" box="[908,1097,697,722]" country="Laos" name="Mount Phu Pane" pageId="37" pageNumber="38" stateProvince="Houaphan">Mount Phu Pane</location>
,
<collectingRegion id="49A74BD1FFB5A40CC62AFD8410CA10EF" box="[1108,1217,697,722]" country="Laos" name="Houaphan" pageId="37" pageNumber="38">Hua Phan</collectingRegion>
<collectorName id="2696E0E5FFB5A40CC6B6FD84110E10EC" box="[1224,1285,698,721]" pageId="37" pageNumber="38">Prov.</collectorName>
,
<quantity id="4C9B28D6FFB5A40CC771FD87116510EC" box="[1295,1390,697,722]" metricMagnitude="0" metricUnit="m" metricValue="2.06" pageId="37" pageNumber="38" unit="m" value="2.06">
<elevation id="004E6200FFB5A40CC771FD87116510EC" box="[1295,1390,697,722]" metricMagnitude="0" metricUnit="m" metricValue="2.06" pageId="37" pageNumber="38" unit="m" value="2.06">2,060 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="3B0B8F6EFFB5A40CC707FD84154D10CB" ID-GBIF-Occurrence="2004970392" collectionCode="KSCJ" country="Laos" pageId="37" pageNumber="38" specimenCount="1" specimenCount-male="1">
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,
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<collectionCode id="ED721DF6FFB5A40CC386FC3C16131127" box="[504,536,770,794]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="37" pageNumber="38">VI</collectionCode>
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(
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),
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Prov.,
<location id="8EBCD3E8FFB5A40CC190FC3C104E1127" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC190FC3C104E1127" box="[1006,1093,769,794]" country="Laos" county="Phou Pan" municipality="Mt." name="St. Jakl" pageId="37" pageNumber="38" stateProvince="Houa Phan">St. Jakl</location>
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(
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),
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<collectionCode id="ED721DF6FFB5A40CC6A0FC3C10F91127" box="[1246,1266,770,794]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="37" pageNumber="38">V</collectionCode>
.2012
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,
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<location id="8EBCD3E8FFB5A40CC742FC3C11931124" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC742FC3C11931124" box="[1340,1432,770,793]" country="Laos" name="Mt. Pan" pageId="37" pageNumber="38">Mt. Pan</location>
, Sam Neua,
<collectorName id="2696E0E5FFB5A40CC367FC1815CC1100" box="[281,455,806,829]" pageId="37" pageNumber="38">Houa Pan Prov</collectorName>
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.;
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(
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),
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<collectingDate id="EF995A1BFFB5A40CC01CFC1816BF1103" box="[610,692,806,830]" pageId="37" pageNumber="38" value="2007-05">
<collectionCode id="ED721DF6FFB5A40CC01CFC18167C1103" box="[610,631,806,830]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="37" pageNumber="38">V</collectionCode>
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,
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<location id="8EBCD3E8FFB5A40CC0BEFC18173F1100" LSID="urn:lsid:plazi:treatment:03CA3425FFB5A40EC2E9FFA9155A1128:8EBCD3E8FFB5A40CC0BEFC18173F1100" box="[704,820,806,829]" country="Laos" name="Xan Neua" pageId="37" pageNumber="38" stateProvince="Houaphan">Xan Neua</location>
,
<collectingRegion id="49A74BD1FFB5A40CC141FC1817A71103" box="[831,940,805,830]" country="Laos" name="Houaphan" pageId="37" pageNumber="38">Hua Phan</collectingRegion>
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.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB5A40EC2B9FC7715A712E8" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8BDC8533FFB5A40CC2B9FC7710B6156C" blockId="37.[151,1437,478,1982]" pageId="37" pageNumber="38">
<emphasis id="B9175921FFB5A40CC2B9FC771631115F" bold="true" box="[199,570,841,866]" pageId="37" pageNumber="38">Redescription (male). General:</emphasis>
Body length 16.518.0 mm; width 7.07.5 mm, widest at humeral umbone, gradually narrowed backward. Body reddish-brown to dark brown. Surface with fulvous, light yellow and black setae, and yellow tomentous maculae.
<emphasis id="B9175921FFB5A40CC037FCAF16991197" bold="true" box="[585,658,913,938]" pageId="37" pageNumber="38">Head:</emphasis>
Brown. Dorsal surface with dense, setiferous, punctures; setae short, fulvous, but long on frons. Anterior margin of clypeus nearly straight and slightly raised. Frons flat or with an indistinct longitudinal ridge (
<figureCitation id="135899B6FFB5A40CC3A3FCE7162611CF" box="[477,557,985,1010]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
). Maxillary palpus and labial palpus dark brown. Antenna yellowish-brown to dark brown; antennal club long, about 1.5 times of antennomeres 27 combined; inner side of antennomere 8 with dense, fulvous setae (
<figureCitation id="135899B6FFB5A40CC3F9FB1F15D31607" box="[391,472,1057,1082]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
). Ventral surface clad with long, fulvous setae (
<figureCitation id="135899B6FFB5A40CC18EFB1F10451607" box="[1008,1102,1057,1082]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Fig. 170</figureCitation>
).
<emphasis id="B9175921FFB5A40CC618FB1F10E31607" bold="true" box="[1126,1256,1057,1082]" pageId="37" pageNumber="38">Pronotum:</emphasis>
Reddish-brown to brown. Widest at base, basomedian area depressed. Lateral margin curved; posterior margin distinctly protruded. Surface densely clad with long, fulvous and black setae (
<figureCitation id="135899B6FFB5A40CC16EFB57176516BF" box="[784,878,1129,1154]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Fig. 169</figureCitation>
).
<emphasis id="B9175921FFB5A40CC1FBFB57100D16BF" bold="true" box="[901,1030,1129,1154]" pageId="37" pageNumber="38">Scutellum:</emphasis>
Reddish-brown to dark brown, with sparse, fulvous setae.
<emphasis id="B9175921FFB5A40CC3DFFBB3160D169B" bold="true" box="[417,518,1165,1190]" pageId="37" pageNumber="38">Elytron:</emphasis>
Humeral umbone, disc, lateral declivity, and distal declivity reddish-brown; juxtascutellar area dark brown; with posthumeral macula, lateral macula, median macula, and distal macula. Surface densely clad with setae; setae long between sutural and discolateral costae; setae black on lateral declivity, light yellow on maculae; setae fulvous on the rest of elytron; humeral umbone glabrous (
<figureCitation id="135899B6FFB5A40CC602FBC710D0172F" box="[1148,1243,1273,1298]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Fig. 169</figureCitation>
).
<emphasis id="B9175921FFB5A40CC68CFBC71624170B" bold="true" pageId="37" pageNumber="38">Mesepimeron, metepisternum and metepimeron:</emphasis>
Dark brown, with fulvous and/or dark brown setae.
<emphasis id="B9175921FFB5A40CC601FA2310E4170B" bold="true" box="[1151,1263,1309,1334]" pageId="37" pageNumber="38">Sternum:</emphasis>
Preprosternum reddish, with sparse, fulvous setae on sides. Mesosternum dark reddish-brown, with sparse, fulvous setae. Mesometasternal process short, glabrous, dark reddish-brown, apex rounded; mesometasternal suture depressed, with long, fulvous setae. Metasternum black, with long, fulvous setae; middle portion glabrous (
<figureCitation id="135899B6FFB5A40CC758FAB71187179F" box="[1318,1420,1417,1442]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Fig. 170</figureCitation>
).
<emphasis id="B9175921FFB5A40CC2E9FA93151817FB" bold="true" box="[151,275,1453,1478]" pageId="37" pageNumber="38">Pygidium:</emphasis>
Brown. With a small yellow macula. Surface densely clad with setae; setae fulvous in middle, dark brown on sides (
<figureCitation id="135899B6FFB5A40CC31CFAEF15B317D7" box="[354,440,1489,1514]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="37" pageNumber="38">Fig. 94</figureCitation>
).
<emphasis id="B9175921FFB5A40CC3ABFAEF165817D7" bold="true" box="[469,595,1489,1514]" pageId="37" pageNumber="38">Abdomen:</emphasis>
Dark reddish-brown. With a slightly depressed longitudinal groove in median. Six abdominal sternites visible; sternites surface clad with fulvous setae; setae on sternite II long, dense; setae on sternites IIIV longer on sides, but almost glabrous in median; setae near posterior margin of sternite VI long, dense; sternite VII with sparse, short setae on sides. Sternites IIV densely clad with black and fulvous setae in dorsal and lateral portion (
<figureCitation id="135899B6FFB5A40CC398F95F164C1447" box="[486,583,1633,1658]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Fig. 170</figureCitation>
).
<emphasis id="B9175921FFB5A40CC01FF95F16A91447" bold="true" box="[609,674,1633,1658]" pageId="37" pageNumber="38">Legs:</emphasis>
Reddish-brown, slender, simple. Metacoxa, femora, and tibia clad with long, fulvous and light yellow setae. Protibia without tiny tooth (
<figureCitation id="135899B6FFB5A40CC1B6F9BB101714A3" box="[968,1052,1669,1694]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="37" pageNumber="38">Fig. 39</figureCitation>
); mesotibia and metatibia with a sharp spine near the middle of outer margin; metatibia with a row of brush-like, fulvous setae along inner margin (
<figureCitation id="135899B6FFB5A40CC2E1F9F3154A14DB" box="[159,321,1741,1766]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Figs. 169170</figureCitation>
); dorsal tooth of metatibia longer than basitarsus of metatarsus (
<figureCitation id="135899B6FFB5A40CC66EF9F3106B14DB" box="[1040,1120,1741,1766]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="37" pageNumber="38">Fig. 63</figureCitation>
). Tarsi simple; basitarsus of metatarsus with a cluster of dense, long setae (
<figureCitation id="135899B6FFB5A40CC0E2F9CF16E01537" box="[668,747,1777,1802]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="37" pageNumber="38">Fig. 63</figureCitation>
).
<emphasis id="B9175921FFB5A40CC17CF9CF17711537" bold="true" box="[770,890,1777,1802]" pageId="37" pageNumber="38">Genitalia:</emphasis>
Parameres broad, short; apex rounded, distinctly expanded. Interparameral split wide, gradually constricted to apex; base with membrane, slightly divided in medial. Median lobe wide at base, filiform apically; distinctly longer than paramere (
<figureCitation id="135899B6FFB5A40CC631F80710A5156F" box="[1103,1198,1849,1874]" captionStart="FIGURES 111122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="37" pageNumber="38">Fig. 119</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FFB5A40CC2B9F863100B1580" blockId="37.[151,1437,478,1982]" pageId="37" pageNumber="38">
<emphasis id="B9175921FFB5A40CC2B9F863152E154B" bold="true" box="[199,293,1885,1910]" pageId="37" pageNumber="38">Female.</emphasis>
Body length 17.017.5 mm; width 7.78.0 mm. Except for sexual dimorphism characteristics of the genus, body color darker (
<figureCitation id="135899B6FFB5A40CC3B8F8BF162215A7" box="[454,553,1921,1946]" captionStart-0="FIGURES 123" captionStart-1="FIGURES 2447" captionStart-2="FIGURES 4865" captionStart-3="FIGURES 6671" captionStartId-0="9.[151,264,1838,1861]" captionStartId-1="10.[151,264,1767,1790]" captionStartId-2="11.[151,264,1850,1873]" captionStartId-3="12.[151,264,954,977]" captionTargetBox-0="[151,1436,195,1817]" captionTargetBox-1="[151,1436,193,1746]" captionTargetBox-2="[187,1411,183,1839]" captionTargetBox-3="[151,1436,193,933]" captionTargetId-0="figure@9.[151,1436,195,1817]" captionTargetId-1="figure@10.[151,1436,193,1746]" captionTargetId-2="figure@11.[187,1411,183,1839]" captionTargetId-3="figure@12.[151,1436,193,933]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." captionText-1="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." captionText-2="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." captionText-3="FIGURES 6671. Metatibia and basitarsus of metatarsus of the Macronotops ovaliceps species group (male in left, female in right). 6667, M. curvimarginatus; 6869, M. medogensis new species; 7071, M. ovaliceps." httpUri-0="https://zenodo.org/record/2585206/files/figure.png" httpUri-1="https://zenodo.org/record/2585208/files/figure.png" httpUri-2="https://zenodo.org/record/2585210/files/figure.png" httpUri-3="https://zenodo.org/record/2585214/files/figure.png" pageId="37" pageNumber="38">Fig. 1 71</figureCitation>
), dorsal tooth of metatibia larger (
<figureCitation id="135899B6FFB5A40CC1C9F8BF100115A7" box="[951,1034,1921,1946]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="37" pageNumber="38">Fig. 64</figureCitation>
), setae on mesotibia and metatibia shorter (
<figureCitation id="135899B6FFB5A40CC28AF89B159C1583" box="[244,407,1957,1982]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="37" pageNumber="38">Figs. 171172</figureCitation>
), and basitarsus of metatarsus with less setae (
<figureCitation id="135899B6FFB5A40CC1E1F89B17FB1583" box="[927,1008,1957,1982]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="37" pageNumber="38">Fig. 64</figureCitation>
).
</paragraph>
<caption id="DF1CD5BBFFB6A40FC2E9F88C174715DA" httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1970,1993]" targetBox="[171,1407,193,1938]" targetPageId="38">
<paragraph id="8BDC8533FFB6A40FC2E9F88C174715DA" blockId="38.[151,1436,1970,2023]" pageId="38" pageNumber="39">
<emphasis id="B9175921FFB6A40FC2E9F88C156515F5" bold="true" box="[151,366,1970,1993]" pageId="38" pageNumber="39">FIGURES 169172.</emphasis>
Habitus of
<taxonomicName id="4C63FEB0FFB6A40FC396F88A16B015F4" baseAuthorityName="Bourgoin" baseAuthorityYear="1916" box="[488,699,1971,1993]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB6A40FC396F88A16B015F4" box="[488,699,1971,1993]" italics="true" pageId="38" pageNumber="39">Macronotops vuilleti</emphasis>
</taxonomicName>
(dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos).
</paragraph>
</caption>
<paragraph id="8BDC8533FFB7A40EC2B9FFA915A712E8" blockId="39.[151,1436,151,789]" pageId="39" pageNumber="40">
<emphasis id="B9175921FFB7A40EC2B9FFA91546128D" bold="true" box="[199,333,151,176]" pageId="39" pageNumber="40">Variability.</emphasis>
Features are stable, but pygidium of some specimens without yellow macula (
<figureCitation id="135899B6FFB7A40EC6ADFFA91154128D" box="[1235,1375,151,176]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="39" pageNumber="40">Figs. 9596</figureCitation>
), and setae are black (
<figureCitation id="135899B6FFB7A40EC335FF82159412E8" box="[331,415,188,213]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="39" pageNumber="40">Fig. 96</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB7A40EC2B9FFE115A413C8" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BDC8533FFB7A40EC2B9FFE115961391" blockId="39.[151,1436,151,789]" pageId="39" pageNumber="40">
<emphasis id="B9175921FFB7A40EC2B9FFE115DA12C5" bold="true" box="[199,465,223,248]" pageId="39" pageNumber="40">Differential diagnosis.</emphasis>
Length of clypeus almost equal to width. Antenna yellowish-brown to dark brown (
<figureCitation id="135899B6FFB7A40EC2E1FE3A15231321" box="[159,296,260,285]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="39" pageNumber="40">Figs. 1617</figureCitation>
). Male pronotum clad with fulvous long setae. Elytra with posthumeral maculae, median maculae, lateral maculae, and distal maculae (
<figureCitation id="135899B6FFB7A40EC043FE1916EF137D" box="[573,740,295,320]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="39" pageNumber="40">Figs. 169, 171</figureCitation>
). Pygidium with a yellow macula (
<figureCitation id="135899B6FFB7A40EC601FE191101137D" box="[1151,1290,295,320]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="39" pageNumber="40">Figs. 94, 97</figureCitation>
), sometimes absent (
<figureCitation id="135899B6FFB7A40EC28FFE7215701358" box="[241,379,332,357]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="39" pageNumber="40">Figs. 9596</figureCitation>
). Abdominal sides without yellow maculae (
<figureCitation id="135899B6FFB7A40EC1FCFE7217E41359" box="[898,1007,332,357]" captionStart-0="FIGURES 123" captionStart-1="FIGURES 2447" captionStart-2="FIGURES 4865" captionStart-3="FIGURES 6671" captionStartId-0="9.[151,264,1838,1861]" captionStartId-1="10.[151,264,1767,1790]" captionStartId-2="11.[151,264,1850,1873]" captionStartId-3="12.[151,264,954,977]" captionTargetBox-0="[151,1436,195,1817]" captionTargetBox-1="[151,1436,193,1746]" captionTargetBox-2="[187,1411,183,1839]" captionTargetBox-3="[151,1436,193,933]" captionTargetId-0="figure@9.[151,1436,195,1817]" captionTargetId-1="figure@10.[151,1436,193,1746]" captionTargetId-2="figure@11.[187,1411,183,1839]" captionTargetId-3="figure@12.[151,1436,193,933]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." captionText-1="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." captionText-2="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." captionText-3="FIGURES 6671. Metatibia and basitarsus of metatarsus of the Macronotops ovaliceps species group (male in left, female in right). 6667, M. curvimarginatus; 6869, M. medogensis new species; 7071, M. ovaliceps." httpUri-0="https://zenodo.org/record/2585206/files/figure.png" httpUri-1="https://zenodo.org/record/2585208/files/figure.png" httpUri-2="https://zenodo.org/record/2585210/files/figure.png" httpUri-3="https://zenodo.org/record/2585214/files/figure.png" pageId="39" pageNumber="40">Figs. 1 70</figureCitation>
,
<figureCitation id="135899B6FFB7A40EC181FE7210211359" box="[1023,1066,332,356]" captionStart="FIGURES 169172" captionStartId="38.[151,264,1970,1993]" captionTargetBox="[171,1407,193,1938]" captionTargetId="figure@38.[171,1407,193,1938]" captionTargetPageId="38" captionText="FIGURES 169172. Habitus of Macronotops vuilleti (dorsal and ventral view). 169170, male (Mount Phu Pane, Hua Phan, NE Laos); 171172, female (Mount Phu Pane, Hua Phan, NE Laos)." httpUri="https://zenodo.org/record/2585240/files/figure.png" pageId="39" pageNumber="40">172</figureCitation>
); male abdomen with a slightly longitudinal groove. Protibia without tiny tooth (
<figureCitation id="135899B6FFB7A40EC0C8FE51173613B5" box="[694,829,367,392]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="39" pageNumber="40">Figs. 3940</figureCitation>
); dorsal tooth of male metatibia longer than basitarsus of metatarsus (
<figureCitation id="135899B6FFB7A40EC342FEAA15871390" box="[316,396,404,429]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="39" pageNumber="40">Fig. 63</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FFB7A40EC2B9FE8915A413C8" blockId="39.[151,1436,151,789]" pageId="39" pageNumber="40">
This species is similar to
<taxonomicName id="4C63FEB0FFB7A40EC079FE87172913ED" box="[519,802,439,464]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="fulvoguttatus">
<emphasis id="B9175921FFB7A40EC079FE87172913ED" box="[519,802,439,464]" italics="true" pageId="39" pageNumber="40">Macronota fulvoguttatus</emphasis>
</taxonomicName>
, and the differences are provided in the differential diagnosis of that species.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB7A40EC2B9FEC11502109A" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BDC8533FFB7A40EC2B9FEC117C7102A" blockId="39.[151,1436,151,789]" box="[199,972,511,536]" pageId="39" pageNumber="40">
<emphasis id="B9175921FFB7A40EC2B9FEC1156B1025" bold="true" box="[199,352,511,536]" pageId="39" pageNumber="40">Distribution.</emphasis>
<collectingCountry id="F374C5A3FFB7A40EC319FEC115A11025" box="[359,426,511,536]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
(
<emphasis id="B9175921FFB7A40EC3C4FEC116AF1025" bold="true" box="[442,676,511,536]" pageId="39" pageNumber="40">new country record</emphasis>
):
<collectingRegion id="49A74BD1FFB7A40EC0C2FD3E171E102A" box="[700,789,512,535]" country="China" name="Yunnan" pageId="39" pageNumber="40">Yunnan</collectingRegion>
;
<collectingCountry id="F374C5A3FFB7A40EC15FFEC117881025" box="[801,899,511,536]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
;
<collectingCountry id="F374C5A3FFB7A40EC1F0FD3E17C3102A" box="[910,968,512,535]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
.
</paragraph>
<paragraph id="8BDC8533FFB7A40EC2B9FD1A1502109A" blockId="39.[151,1436,151,789]" pageId="39" pageNumber="40">
<emphasis id="B9175921FFB7A40EC2B9FD1A15821000" bold="true" box="[199,393,548,573]" pageId="39" pageNumber="40">Natural history.</emphasis>
Larvae of this species were excavated from several rotten trunks in February and finally emerged in May. The habitus is a slope by a stream in Mount Huanglianshan of southern
<collectingRegion id="49A74BD1FFB7A40EC6E8FD7610E41062" box="[1174,1263,584,607]" country="China" name="Yunnan" pageId="39" pageNumber="40">Yunnan</collectingRegion>
, and the trunks were lying under a forest composed of bamboo and
<taxonomicName id="4C63FEB0FFB7A40EC08BFD52115A10B8" authority="Crevost et Lemarie" authorityName="Crevost et Lemarie" box="[757,1361,620,645]" class="Magnoliopsida" family="Zingiberaceae" genus="Amomum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Zingiberales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="tsaoko">
<emphasis id="B9175921FFB7A40EC08BFD5217BA10B8" box="[757,945,620,645]" italics="true" pageId="39" pageNumber="40">Amomum tsaoko</emphasis>
Crevost
<emphasis id="B9175921FFB7A40EC662FD53103B10B9" box="[1052,1072,621,644]" italics="true" pageId="39" pageNumber="40">et</emphasis>
Lemarie (Zingiberaceae)
</taxonomicName>
(
<figureCitation id="135899B6FFB7A40EC71AFD5214F11095" captionStart="FIGURES 276281" captionStartId="58.[151,264,1812,1835]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@58.[151,1436,193,1792]" captionTargetPageId="58" captionText="FIGURES 276281. Habitat of Macronotops. 276277, habitat of M. sexmaculatus and M. medogensis new species in Valley near Bononggong of Mêdog (Xizang, China); 278, an adult of M. sexmaculatus feeding on sap of bamboo (photograph by Yang He); 279281, habitat of M. vuilleti in Mount Huanglianshan (Yunnan, China), and larva in fallen rotten wood." httpUri="https://zenodo.org/record/2585270/files/figure.png" pageId="39" pageNumber="40">Figs. 279281</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB7A40EC2B9FD8A155A1128" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8BDC8533FFB7A40EC2B9FD8A155A1128" blockId="39.[151,1436,151,789]" pageId="39" pageNumber="40">
<emphasis id="B9175921FFB7A40EC2B9FD8A153010F0" bold="true" box="[199,315,692,717]" pageId="39" pageNumber="40">Remarks.</emphasis>
<taxonomicName id="4C63FEB0FFB7A40EC338FD8B161110F0" authorityName="Bourgoin" authorityYear="1916" box="[326,538,692,717]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vuilleti">
<emphasis id="B9175921FFB7A40EC338FD8B161110F0" box="[326,538,692,717]" italics="true" pageId="39" pageNumber="40">Macronota vuilleti</emphasis>
</taxonomicName>
was rarely reported and previously only known from northeastern
<collectingCountry id="F374C5A3FFB7A40EC74CFD8B116310F1" box="[1330,1384,693,716]" name="Laos" pageId="39" pageNumber="40">Laos</collectingCountry>
and northern
<collectingCountry id="F374C5A3FFB7A40EC282FDE9155010CD" box="[252,347,727,752]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
(
<bibRefCitation id="EFF2F8C2FFB7A40EC317FDE9161E10CD" author="Bourgoin, A." box="[361,533,727,752]" pageId="39" pageNumber="40" pagination="133 - 137" refId="ref34313" refString="Bourgoin, A. (1916) Description de trois Macronota nouveaux (Col. Scarabaeidae). Bulletin de la Societe Entomologique de France Paris, 1916 (7), 133 - 137." type="journal article" year="1916">Bourgoin 1916</bibRefCitation>
;
<bibRefCitation id="EFF2F8C2FFB7A40EC05EFDE916A910CD" author="Sakai, K." box="[544,674,727,752]" pageId="39" pageNumber="40" pagination="11 - 22" refId="ref35646" refString="Sakai, K. (2008) Record of Cetoniid beetles from the central and northern Laos (Coleoptera, Scarabaeidae). Saikaku-Tsushin, 16, 11 - 22. [in Japanese]" type="journal article" year="2008">Sakai 2008</bibRefCitation>
). This species is also occurring in southern
<collectingRegion id="49A74BD1FFB7A40EC6FAFDE610D210D2" box="[1156,1241,728,751]" country="China" name="Yunnan" pageId="39" pageNumber="40">Yunnan</collectingRegion>
of
<collectingCountry id="F374C5A3FFB7A40EC683FDE9114B10CD" box="[1277,1344,727,752]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
close to the
<typeStatus id="54D83B91FFB7A40EC2BEFDC314FB1128" box="[192,240,765,789]" pageId="39" pageNumber="40">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>