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<document id="F09B87FF08538326DD8754CDC2328471" ID-DOI="10.11646/zootaxa.5476.1.29" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681921" ID-ZooBank="DD4EA1FE-A0BE-4BA4-9AD3-E460B131C518" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="felipe" checkinTime="1720375669801" checkinUser="plazi" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2024" docId="03DD87DAFFAC125728EEFCB187691921" docLanguage="en" docName="zootaxa.5476.1.29.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.29" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyphira ngankee Rahayu &amp; Ng, 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="377" masterDocId="FFE4FFA2FFBE12442879FFAF843C1F54" masterDocTitle="On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific" masterLastPageNumber="392" masterPageNumber="358" pageNumber="376" updateTime="1721070200110" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="9A857B855123F0140E9C32411ECAEB2E">On the genus Lyphira Galil, 2009 (Crustacea: Brachyura: Leucosiidae), with descriptions of four new species from the West Pacific</mods:title>
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<mods:namePart id="6AA6D05117620072E5873E8DD407122A">Rahayu, Dwi Listyo</mods:namePart>
<mods:affiliation id="65A0800AF24BA11A491F484853FE944C">Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia.</mods:affiliation>
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<mods:namePart id="6AFB2E2C07AB70E6E1D0F1D152EE823A">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="EA388102494499427BA02C9FBE0874DF">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore.</mods:affiliation>
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<taxonomicName id="4C744D4FFFAC125628EEFCB185591C6D" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[151,357,798,825]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
<emphasis id="B900EADEFFAC125628EEFCB185591C6D" bold="true" box="[151,357,798,825]" italics="true" pageId="18" pageNumber="376">Lyphira ngankee</emphasis>
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<paragraph id="8BCB36CCFFAC125628EEFCEC80411C09" blockId="18.[151,1149,798,861]" box="[151,1149,834,861]" pageId="18" pageNumber="376">
(
<figureCitation id="134F2A49FFAC125628E6FCEC85251C09" box="[159,281,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1E–J</figureCitation>
,
<figureCitation id="134F2A49FFAC1256295CFCED85561C09" box="[293,362,834,861]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAC1256290FFCEC85A61C09" box="[374,410,835,861]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
,
<figureCitation id="134F2A49FFAC125629DFFCEC85F61C09" box="[422,458,835,861]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="18" pageNumber="376">8A</figureCitation>
,
<figureCitation id="134F2A49FFAC125629AFFCEC85C61C09" box="[470,506,835,861]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="18" pageNumber="376">9D</figureCitation>
,
<figureCitation id="134F2A49FFAC12562A7FFCEC860A1C09" box="[518,566,835,861]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="18" pageNumber="376">10E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562A38FCEC864C1C09" box="[577,624,835,861]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">11E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562A02FCEC86F61C09" box="[635,714,835,861]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="18" pageNumber="376">13F, G</figureCitation>
,
<figureCitation id="134F2A49FFAC12562AACFCEC873B1C09" box="[725,775,835,861]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="18" pageNumber="376">14D</figureCitation>
,
<figureCitation id="134F2A49FFAC12562B6AFCED87031C09" box="[787,831,834,861]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="18" pageNumber="376">15F</figureCitation>
,
<figureCitation id="134F2A49FFAC12562B32FCEC87461C09" box="[843,890,835,861]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="18" pageNumber="376">16E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562BFFFCEC87EB1C09" box="[902,983,835,861]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="18" pageNumber="376">18A‒E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562B9AFCEC80041C09" box="[995,1080,835,861]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="18" pageNumber="376">20A‒D</figureCitation>
,
<figureCitation id="134F2A49FFAC12562C3AFCEC80491C09" box="[1091,1141,835,861]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="18" pageNumber="376">21D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAC125628EEFC2880ED1B6D" pageId="18" pageNumber="376" type="reference_group">
<paragraph id="8BCB36CCFFAC125628EEFC2885D31CEA" blockId="18.[151,1436,903,1081]" pageId="18" pageNumber="376">
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC2885D31CEA" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628EEFC2885411CCB" box="[151,381,903,927]" italics="true" pageId="18" pageNumber="376">
<taxonomicName id="4C744D4FFFAC125628EEFC2885631CCB" ID-CoL="4GDVK" authorityName="Stimpson" authorityYear="1858" box="[151,351,903,927]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="tuberculosa">Philyra tuberculosa</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD510DDFFAC12562904FC2886221CCB" author="Shen, C. - J." box="[381,542,903,927]" page="185" pageId="18" pageNumber="376" year="1931">
<bibRefCitation id="EFE54B3DFFAC12562904FC2886221CCB" author="Shen, C. - J." box="[381,542,903,927]" pageId="18" pageNumber="376" pagination="185 - 197" refId="ref21785" refString="Shen, C. - J. (1931) The crabs of Hong Kong. Part 2. Hong Kong Naturalist, 2 (3), 185 - 197." type="journal article" year="1931">Shen 1931: 185</bibRefCitation>
</treatmentCitation>
, text figs. 1, 2, 4;
<bibRefCitation id="EFE54B3DFFAC12562AACFC2887781CCB" author="Shen, C. - J." box="[725,836,903,927]" pageId="18" pageNumber="376" pagination="1 - 300" refId="ref21817" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, 9 (1), 1 - 300, pls. 1 - 10." type="journal article" year="1932">Shen 1932</bibRefCitation>
: text fig. 11e, f. (not
<taxonomicName id="4C744D4FFFAC12562C60FC2881B41CCB" ID-CoL="4GDVK" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[1049,1416,903,927]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="tuberculosa">
<emphasis id="B900EADEFFAC12562C60FC2880DE1CCB" box="[1049,1250,903,927]" italics="true" pageId="18" pageNumber="376">Philyra tuberculosa</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562C91FC2881B41CCB" author="Stimpson, W." box="[1256,1416,903,927]" pageId="18" pageNumber="376" pagination="159 - 163" refId="ref21886" refString="Stimpson, W. (1858) Crustacea Oxystomata. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VI. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 159 - 163. https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C744D4FFFAC125628BEFC0985DF1CEA" ID-CoL="76YLX" authority="Bell, 1855" authorityName="Bell" authorityYear="1855" box="[199,483,934,958]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="B900EADEFFAC125628BEFC09854C1CE9" box="[199,368,934,958]" italics="true" pageId="18" pageNumber="376">Philyra carinata</emphasis>
<bibRefCitation id="EFE54B3DFFAC1256290FFC0985DF1CEA" author="Bell, T." box="[374,483,934,958]" pageId="18" pageNumber="376" pagination="361 - 367" refId="ref19289" refString="Bell, T. (1855) Horae carcinologicae, or Notices of Crustacea. I. A Monograph of the Leucosiidae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species. Annals and Magazine of Natural History, Series 2, 16 (45), 361 - 367. https: // doi. org / 10.5962 / bhl. title. 9957" type="journal article" year="1855">Bell, 1855</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCB36CCFFAC125628EEFC6B80E11C88" blockId="18.[151,1436,903,1081]" box="[151,1245,964,989]" pageId="18" pageNumber="376">
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC6B80E11C88" box="[151,1245,964,989]" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628EEFC6B856B1C88" box="[151,343,964,988]" italics="true" pageId="18" pageNumber="376">
<taxonomicName id="4C744D4FFFAC125628EEFC6B85051C88" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[151,313,964,988]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globosa">Philyra globosa</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD510DDFFAC1256292EFC6B85CA1C88" author="Shen, C. - J." box="[343,502,964,989]" page="216" pageId="18" pageNumber="376" year="1940">
<bibRefCitation id="EFE54B3DFFAC1256292EFC6B85CA1C88" author="Shen, C. - J." box="[343,502,964,989]" pageId="18" pageNumber="376" pagination="211 - 242" refId="ref21852" refString="Shen, C. - J. (1940) The brachyuran fauna of Hongkong. Journal of the Hong Kong Fisheries Research Station, 1 (2), 211 - 242." type="journal article" year="1940">Shen 1940: 216</bibRefCitation>
</treatmentCitation>
(not
<taxonomicName id="4C744D4FFFAC12562A52FC6A87461C89" authority="Fabricius, 1793" authorityName="Fabricius" authorityYear="1793" box="[555,890,964,989]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globosus">
<emphasis id="B900EADEFFAC12562A52FC6A86EA1C88" box="[555,726,964,988]" italics="true" pageId="18" pageNumber="376">Cancer globosus</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562AA5FC6B87461C89" author="Fabricius, J. C." box="[732,890,964,989]" pageId="18" pageNumber="376" refId="ref19754" refString="Fabricius, J. C. (1793) Entomologia systematica emendata et aucta secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vol. 2. impensis Christ. Gottl. Proft, Hafniae, viii + 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius, 1793</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C744D4FFFAC12562BECFC6A80ED1C88" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[917,1233,964,988]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="globus">
<emphasis id="B900EADEFFAC12562BECFC6A80161C88" box="[917,1066,964,988]" italics="true" pageId="18" pageNumber="376">Cancer globus</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562C49FC6B80ED1C88" author="Fabricius, J. C." box="[1072,1233,964,988]" pageId="18" pageNumber="376" refId="ref19698" refString="Fabricius, J. C. (1775) Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, adiectis Synonymis, Locis, Descriptionibus, Observationibus. Flensburgi &amp; Lipsiae, Korte, xxvii + 832 pp. https: // doi. org / 10.5962 / bhl. title. 36510" type="book" year="1775">Fabricius, 1775</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCB36CCFFAC125628EEFC4C81B61CAF" blockId="18.[151,1436,903,1081]" box="[151,1418,995,1019]" pageId="18" pageNumber="376">
<treatmentCitationGroup id="AB6411E2FFAC125628EEFC4C81B61CAF" box="[151,1418,995,1019]" pageId="18" pageNumber="376">
<taxonomicName id="4C744D4FFFAC125628EEFC4C855B1CAF" ID-CoL="76YMQ" authorityName="Doflein" authorityYear="1904" box="[151,359,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Philyra" isUncertain="true" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
<emphasis id="B900EADEFFAC125628EEFC4C84DF1CAF" box="[151,227,995,1019]" italics="true" pageId="18" pageNumber="376">Philyra</emphasis>
cf.
<emphasis id="B900EADEFFAC12562973FC4C855B1CAF" box="[266,359,995,1019]" italics="true" pageId="18" pageNumber="376">laminata</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAC1256291EFC4C85571CAF" box="[359,363,995,1019]" italics="true" pageId="18" pageNumber="376">.</emphasis>
—
<treatmentCitation id="0AD510DDFFAC125629FDFC4C86251CAF" author="Hill, D. S." box="[388,537,995,1019]" page="199" pageId="18" pageNumber="376" year="1982">
<bibRefCitation id="EFE54B3DFFAC125629FDFC4C86251CAF" author="Hill, D. S." box="[388,537,995,1019]" pageId="18" pageNumber="376" pagination="195 - 205" refId="ref20600" refString="Hill, D. S. (1982) The Leucosiidae (Crustacea: Decapoda) of Hong Kong. In: Morton, B. S. &amp; Tseng, C. K. (Eds.), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 195 - 205." type="book chapter" year="1982">Hill 1982: 199</bibRefCitation>
</treatmentCitation>
, pl. 3C. (not
<taxonomicName id="4C744D4FFFAC12562AE7FC4C87E21CAF" ID-CoL="76YMQ" authority="Doflein, 1904" authorityName="Doflein" authorityYear="1904" box="[670,990,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
<emphasis id="B900EADEFFAC12562AE7FC4C87761CAF" box="[670,842,995,1019]" italics="true" pageId="18" pageNumber="376">Philyra laminata</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562B28FC4C87E21CAF" author="Doflein, F." box="[849,990,995,1019]" pageId="18" pageNumber="376" pagination="1 - 58" refId="ref19632" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Vol. 6. Verlag von Gustav Fischer, Jena, pp. 1 - 6 + i - xiv + 1 - 314 pp., figs. I - LVIII, atlas pls. 1 - 58." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C744D4FFFAC12562B81FC4B81BA1CAF" ID-CoL="6LSP4" authority="(Doflein, 1904))" authorityName="" baseAuthorityName="Doflein" baseAuthorityYear="1904" box="[1016,1414,995,1019]" class="Malacostraca" family="Leucosiidae" genus="Heteronucia" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="laminata">
<emphasis id="B900EADEFFAC12562B81FC4B80EA1CAF" box="[1016,1238,995,1019]" italics="true" pageId="18" pageNumber="376">Heteronucia laminata</emphasis>
(
<bibRefCitation id="EFE54B3DFFAC12562C9DFC4C81491CAF" author="Doflein, F." box="[1252,1397,995,1019]" pageId="18" pageNumber="376" pagination="1 - 58" refId="ref19632" refString="Doflein, F. (1904) Brachyura. In: Wissenschaftliche Ergebnisse der Deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899. Vol. 6. Verlag von Gustav Fischer, Jena, pp. 1 - 6 + i - xiv + 1 - 314 pp., figs. I - LVIII, atlas pls. 1 - 58." type="book chapter" year="1904">Doflein, 1904</bibRefCitation>
))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCB36CCFFAC125628EEFBAD80E21B4E" blockId="18.[151,1436,903,1081]" box="[151,1246,1026,1050]" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628EEFBAD85BF1B4E" box="[151,387,1026,1050]" italics="true" pageId="18" pageNumber="376">
<taxonomicName id="4C744D4FFFAC125628EEFBAD85591B4E" ID-CoL="76YMK" authorityName="Ortmann" authorityYear="1892" box="[151,357,1026,1050]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">Philyra heterograna</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD510DDFFAC125629FAFBAD86971B4E" author="Blackmore &amp; Rainbow" box="[387,683,1026,1050]" pageId="18" pageNumber="376" year="2000">
<bibRefCitation id="EFE54B3DFFAC125629FAFBAD86971B4E" author="Blackmore, G. &amp; Rainbow, P. S." box="[387,683,1026,1050]" pageId="18" pageNumber="376" pagination="517 - 529" refId="ref19375" refString="Blackmore, G. &amp; Rainbow, P. S. (2000) Epibenthic crab (Crustacea: Brachyura) assemblages of the southeastern waters of Hong Kong: the 1998 Trawl Programme. In: Morton, B. (Ed.), Proceedings of the Tenth International marine Biological Workshop: The marine Flora and Fauna of Hong Kong and southern China. Hong Kong University Press, Hong Kong, pp. 517 - 529." type="book chapter" year="2000">Blackmore &amp; Rainbow 2000</bibRefCitation>
</treatmentCitation>
: appendix 1. (not
<taxonomicName id="4C744D4FFFAC12562B19FBAD80EE1B4E" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[864,1234,1026,1050]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
<emphasis id="B900EADEFFAC12562B19FBAD80121B4E" box="[864,1070,1026,1050]" italics="true" pageId="18" pageNumber="376">Philyra heterograna</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562C4CFBAD80EE1B4E" author="Ortmann, A." box="[1077,1234,1026,1050]" pageId="18" pageNumber="376" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BCB36CCFFAC125628EEFB8F80ED1B6D" blockId="18.[151,1436,903,1081]" box="[151,1233,1056,1081]" pageId="18" pageNumber="376">
<treatmentCitationGroup id="AB6411E2FFAC125628EEFB8F80ED1B6D" box="[151,1233,1056,1081]" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628EEFB8F85BA1B6C" box="[151,390,1056,1080]" italics="true" pageId="18" pageNumber="376">
<taxonomicName id="4C744D4FFFAC125628EEFB8F85541B6C" ID-CoL="3WTBK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,360,1056,1080]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">Lyphira heterograna</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AD510DDFFAC125629FFFB8E86611B6D" author="Wong, K. J. H. &amp; Tao, S. - R. &amp; Leung, K. M. Y." box="[390,605,1056,1081]" page="20" pageId="18" pageNumber="376" year="2021">
<bibRefCitation id="EFE54B3DFFAC125629FFFB8E86611B6D" author="Wong, K. J. H. &amp; Tao, S. - R. &amp; Leung, K. M. Y." box="[390,605,1056,1081]" pageId="18" pageNumber="376" pagination="1 - 74" refId="ref22379" refString="Wong, K. J. H., Tao, S. - R. &amp; Leung, K. M. Y. (2021) Subtidal crabs of Hong Kong: Brachyura (Crustacea: Decapoda) from benthic trawl surveys conducted by the University of Hong Kong, 2012 to 2018. Regional Studies in Marine Science, 48, 1 - 74. 102013. https: // doi. org / 10.1016 / j. rsma. 2021.102013" type="journal article" year="2021">
Wong
<emphasis id="B900EADEFFAC125629B1FB8E85C01B6C" box="[456,508,1056,1080]" italics="true" pageId="18" pageNumber="376">et al.</emphasis>
2021: 20
</bibRefCitation>
</treatmentCitation>
, fig. 30a–c, pl. 6C. (not
<taxonomicName id="4C744D4FFFAC12562B2AFB8F80FA1B6D" ID-CoL="76YMK" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[851,1222,1056,1081]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="heterograna">
<emphasis id="B900EADEFFAC12562B2AFB8F801D1B6C" box="[851,1057,1056,1080]" italics="true" pageId="18" pageNumber="376">Philyra heterograna</emphasis>
<bibRefCitation id="EFE54B3DFFAC12562C51FB8E80FA1B6D" author="Ortmann, A." box="[1064,1222,1057,1081]" pageId="18" pageNumber="376" pagination="532 - 588" refId="ref21357" refString="Ortmann, A. (1892) Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Die Decapoden-Krebse des Strassburger Museums, mit besonderer Beruck- sichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und z. Z. im Strassburger Museum aufbewahrten Formen. V. Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588, pl. 26. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAC125628EEFBC880F41BB9" pageId="18" pageNumber="376" type="materials_examined">
<paragraph id="8BCB36CCFFAC125628EEFBC880F41BB9" blockId="18.[151,1437,1126,2017]" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628EEFBC885021BD4" bold="true" box="[151,318,1126,1153]" pageId="18" pageNumber="376">Type material</emphasis>
.
<materialsCitation id="3B1C3C91FFAC12562934FBC886B71BF1" collectingDate="1998-11-12" collectionCode="ZRC" collectorName="N. K. Ng &amp; Y. Cai" location="Nanao Island" pageId="18" pageNumber="376" specimenCode="ZRC 1999.0456" specimenCount="1" typeStatus="holotype">
<emphasis id="B900EADEFFAC12562934FBC885881BD5" box="[333,436,1127,1153]" italics="true" pageId="18" pageNumber="376">
<typeStatus id="54CF886EFFAC12562934FBC885881BD5" box="[333,436,1127,1153]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
</emphasis>
: male (20.8 × 21.8 mm),
<specimenCode id="DBD29EB7FFAC12562AACFBC887B11BD5" box="[725,909,1126,1153]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="376">ZRC 1999.0456</specimenCode>
,
<location id="8EAB6017FFAC12562BE0FBC8800C1BD5" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAC12562BE0FBC8800C1BD5" box="[921,1072,1127,1153]" name="Nanao Island" pageId="18" pageNumber="376">Nanao Island</location>
, Gianjing fish port, Guangdong, coll.
<collectorName id="2681531AFFAC125628A9FB24850D1BF1" box="[208,305,1163,1189]" pageId="18" pageNumber="376">N.K. Ng</collectorName>
&amp;
<collectorName id="2681531AFFAC1256292EFB24859D1BF1" box="[343,417,1163,1189]" pageId="18" pageNumber="376">Y. Cai</collectorName>
,
<date id="FFCA100CFFAC125629D4FB2486B71BF1" box="[429,651,1163,1189]" pageId="18" pageNumber="376" value="1998-11-12">
<collectingDate id="EF8EE9E4FFAC125629D4FB2486B71BF1" box="[429,651,1163,1189]" pageId="18" pageNumber="376" value="1998-11-12">12 November 1998</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B1C3C91FFAC12562AEEFB2480AB1BF1" box="[663,1175,1162,1189]" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC12562AEEFB2487351BF0" box="[663,777,1163,1188]" italics="true" pageId="18" pageNumber="376">
<typeStatus id="54CF886EFFAC12562AEEFB2487351BF0" box="[663,777,1163,1188]" pageId="18" pageNumber="376" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D72FD45FFAC12562B60FB25874C1BF1" box="[793,880,1162,1189]" count="5" pageId="18" pageNumber="376" type="male">5 males</specimenCount>
(largest 20.7 × 21.3 mm)
</materialsCitation>
,
<materialsCitation id="3B1C3C91FFAC12562CDAFB2480F81BB9" collectingDate="2012-08-27" collectionCode="ZRC" country="Hong Kong" elevation="12" latitude="22.215433" location="Lantau Island" longLatPrecision="1" longitude="113.9095" municipality="University Trawl" pageId="18" pageNumber="376" specimenCode="ZRC 2022.0628" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D72FD45FFAC12562CDAFB24812C1BF1" box="[1187,1296,1163,1189]" count="2" pageId="18" pageNumber="376" type="female">2 females</specimenCount>
(larger 18.8 × 18.8 mm),
<specimenCode id="DBD29EB7FFAC1256294CFB0085CC1B9D" box="[309,496,1199,1225]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="376">ZRC 2022.0628</specimenCode>
, station W2, southeast of
<location id="8EAB6017FFAC12562B5CFB0087F91B9D" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAC12562B5CFB0087F91B9D" box="[805,965,1199,1225]" country="Hong Kong" latitude="22.215433" longLatPrecision="1" longitude="113.9095" municipality="University Trawl" name="Lantau Island" pageId="18" pageNumber="376">Lantau Island</location>
,
<collectingCountry id="F363765CFFAC12562BADFB0080621B9D" box="[980,1118,1199,1225]" name="Hong Kong" pageId="18" pageNumber="376">Hong Kong</collectingCountry>
, 22°11.264'N 113°51.358'E –
<geoCoordinate id="EE40500BFFAC125628D5FB7C85051BB9" box="[172,313,1235,1261]" degrees="22" direction="north" minutes="12.926" orientation="latitude" pageId="18" pageNumber="376" precision="1" value="22.215433">22°12.926'N</geoCoordinate>
<geoCoordinate id="EE40500BFFAC12562938FB7D85E71BB9" box="[321,475,1234,1261]" degrees="113" direction="east" minutes="54.570" orientation="longitude" pageId="18" pageNumber="376" precision="1" value="113.9095">113°54.570'E</geoCoordinate>
, 10.0–11.6 m, coll. Hong Kong
<collectingMunicipality id="6BAFACB6FFAC12562B3BFB7C80381BB9" box="[834,1028,1235,1261]" pageId="18" pageNumber="376">University Trawl</collectingMunicipality>
,
<date id="FFCA100CFFAC12562C76FB7C80F81BB9" box="[1039,1220,1235,1261]" pageId="18" pageNumber="376" value="2012-08-27">
<collectingDate id="EF8EE9E4FFAC12562C76FB7C80F81BB9" box="[1039,1220,1235,1261]" pageId="18" pageNumber="376" value="2012-08-27">27 August 2012</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAC125728BEFB59864B1EF9" lastPageId="19" lastPageNumber="377" pageId="18" pageNumber="376" type="diagnosis">
<paragraph id="8BCB36CCFFAC125728BEFB5980641EDD" blockId="18.[151,1437,1126,2017]" lastBlockId="19.[151,1437,150,1653]" lastPageId="19" lastPageNumber="377" pageId="18" pageNumber="376">
<emphasis id="B900EADEFFAC125628BEFB5985061A44" bold="true" box="[199,314,1270,1296]" pageId="18" pageNumber="376">Diagnosis</emphasis>
. Carapace approximately as broad as long, rounded (
<figureCitation id="134F2A49FFAC12562BF1FB5887DB1A45" box="[904,999,1271,1297]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562B8BFB59800B1A45" box="[1010,1079,1270,1297]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAC12562C3BFB5880541A45" box="[1090,1128,1271,1297]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
); dorsal surface of carapace covered with closely spaced minute granules, hepatic, branchial and intestinal regions with slightly larger, more prominent granules;. Frontal margin granulated; postfrontal lobe slightly convex with slight concave medially (
<figureCitation id="134F2A49FFAC12562D1DFA9084FA1A29" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">Figs. 11E</figureCitation>
). Endostomial spine strong, produced, visible from dorsal view (
<figureCitation id="134F2A49FFAC12562BC0FACC80241A29" box="[953,1048,1379,1405]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1F</figureCitation>
,
<figureCitation id="134F2A49FFAC12562C51FACC80451A29" box="[1064,1145,1379,1405]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="18" pageNumber="376">13F, G</figureCitation>
,
<figureCitation id="134F2A49FFAC12562CF1FACC80801A29" box="[1160,1212,1379,1405]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="18" pageNumber="376">14D</figureCitation>
). Lateral margin of carapace with row of small granules interspersed by larger granule; posterior margin gently convex (
<figureCitation id="134F2A49FFAC12562C99FA28815C1AF5" box="[1248,1376,1414,1441]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAC12562D13FA2881AC1AF5" box="[1386,1424,1415,1441]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
). Posterior carapace margin with row of granules, slightly larger granule medially and on each posterolateral corner, with margin between them gently convex (
<figureCitation id="134F2A49FFAC12562A0BFA6086DE1ABD" box="[626,738,1487,1513]" captionStart="FIGURE 10" captionStartId="15.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1593]" captionTargetId="figure-20@15.[151,1435,181,1593]" captionTargetPageId="15" captionText="FIGURE 10. Posterior carapace margin. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681946" httpUri="https://zenodo.org/record/12681946/files/figure.png" pageId="18" pageNumber="376">Figs. 10E</figureCitation>
). Third maxilliped exopod 2 times as long as broad, wider than basal part of endopod, forming petaliform structure with strongly convex outer margin, with submarginal row of low to very low granules along inner margin; endopod with completely fused basis-ischium, distinctly longer than merus, with shallow submarginal sulcus near inner margin (
<figureCitation id="134F2A49FFAC12562B54F99487B31901" box="[813,911,1595,1621]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="18" pageNumber="376">Figs. 1G</figureCitation>
,
<figureCitation id="134F2A49FFAC12562BE3F99487F51901" box="[922,969,1595,1621]" captionStart="FIGURE 11" captionStartId="16.[151,250,1757,1781]" captionTargetBox="[290,1298,181,1733]" captionTargetId="figure-17@16.[290,1298,181,1733]" captionTargetPageId="16" captionText="FIGURE 11. Frontal view of cephalothorax and third maxilliped. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681948" httpUri="https://zenodo.org/record/12681948/files/figure.png" pageId="18" pageNumber="376">11E</figureCitation>
,
<figureCitation id="134F2A49FFAC12562BADF99480341901" box="[980,1032,1595,1621]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="18" pageNumber="376">18A</figureCitation>
). Chelipeds subequal; merus, carpus and chela surfaces smooth or covered with low, small rounded granules (
<figureCitation id="134F2A49FFAC12562BA1F9F0805C192D" box="[984,1120,1630,1657]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAC12562C17F9F080AF192D" box="[1134,1171,1631,1657]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
); merus subcylindrical, symmetrical along entire length, surface granulated, large granules on proximal area and along margins, minutes granules distally; carpus smooth, unarmed; chela stout, palm surface smooth, upper and lower margins with row of granules, longitudinal row of relatively large granules on lower outer surface of palm; pollex slightly bent, cutting edge lined with denticles; dactylus 1.4 times as long as palm along upper margin, row of small granules on upper margin, cutting edge with median large tooth lined with denticles, remaining of edge with small denticles (
<figureCitation id="134F2A49FFAC12562D54F8BC81B31879" box="[1325,1423,1810,1837]" captionStart="FIGURE 15" captionStartId="24.[151,250,1660,1684]" captionTargetBox="[151,1435,181,1636]" captionTargetId="figure-17@24.[151,1435,181,1636]" captionTargetPageId="24" captionText="FIGURE 15. Outer view of male chela. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L. heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; J, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681957" httpUri="https://zenodo.org/record/12681957/files/figure.png" pageId="18" pageNumber="376">Fig. 15F</figureCitation>
). P2–P5 slender, short; merus distinctly longer than carpus and propodus, dactylus longer than propodus, lanceolate, terminating in corneous tips (
<figureCitation id="134F2A49FFAC1256299FF8F486511821" box="[486,621,1882,1909]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="18" pageNumber="376">Figs. 5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAC12562A03F8F4869C1821" box="[634,672,1883,1909]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="18" pageNumber="376">6D</figureCitation>
); fourth leg shortest with merus 3.8 times as long as broad, margin lined with small granules, more prominent on ventral margin (
<figureCitation id="134F2A49FFAC12562B1FF8D087F218CD" box="[870,974,1919,1945]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="18" pageNumber="376">Fig. 16E</figureCitation>
). Thoracic sternites transversely narrow, surface finely granulated, larger granules laterally (
<figureCitation id="134F2A49FFAC12562AAFF80C870C18E9" box="[726,816,1955,1981]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="18" pageNumber="376">Fig. 8A</figureCitation>
). Male sternopleonal cavity deep, reaching to proximal distance between fused thoracic sternites 1–3. Male pleon narrow, slender, long; somite 1 longitudinally narrow, wide; somites 2–6 fused, forming elongate trapezoidal plate, shallow suture just visible between somites 5 and 6, surface smooth; somite 6 longitudinally subrectangular, surface with one granule submedially, lateral margins gently convex; telson triangular, longer than wide (
<figureCitation id="134F2A49FFAD12572A07FF7086E31FAD" box="[638,735,223,249]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1H</figureCitation>
,
<figureCitation id="134F2A49FFAD12572A92FF7087331FAD" box="[747,783,223,249]" captionStart="FIGURE 8" captionStartId="13.[151,250,1738,1762]" captionTargetBox="[151,1435,181,1714]" captionTargetId="figure-20@13.[151,1435,181,1714]" captionTargetPageId="13" captionText="FIGURE 8. Male thoracic sternum and pleon. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; C, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; D, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; E, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; F, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681942" httpUri="https://zenodo.org/record/12681942/files/figure.png" pageId="19" pageNumber="377">8A</figureCitation>
,
<figureCitation id="134F2A49FFAD12572B63FF7087771FAD" box="[794,843,223,249]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18B</figureCitation>
). G1 elongate, slender, slightly dilated distally; apical process short, bent at an angle of 60º along the longitudinal axis, with blunt tip (
<figureCitation id="134F2A49FFAD12572C64FEAC804B1E49" box="[1053,1143,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1J</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CFDFEAC80E81E49" box="[1156,1236,259,285]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18C‒E</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C99FEAC810B1E49" box="[1248,1335,259,285]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="19" pageNumber="377">20A‒D</figureCitation>
). Female sternopleonal cavity reaches distal part of sternite 1; female pleon broad, convex, smooth, somites 2‒6 fused, suture between somites 2 and 3; telson shorter than its basal width (
<figureCitation id="134F2A49FFAD12572B31FEE4879C1E31" box="[840,928,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1I</figureCitation>
,
<figureCitation id="134F2A49FFAD12572BD7FEE487EF1E31" box="[942,979,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1822,1846]" captionTargetBox="[256,1332,181,1798]" captionTargetId="figure-17@14.[256,1332,181,1798]" captionTargetPageId="14" captionText="FIGURE 9. Female thoracic sternum and pleon.A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681944" httpUri="https://zenodo.org/record/12681944/files/figure.png" pageId="19" pageNumber="377">9D</figureCitation>
); vulvae spaced apart on sternite 6, each opening weakly crescent-shaped, relatively short, without opercular cover (
<figureCitation id="134F2A49FFAD12572B9AFEC080771EDD" box="[995,1099,367,393]" captionStart="FIGURE 21" captionStartId="30.[151,250,1646,1670]" captionTargetBox="[151,1435,181,1622]" captionTargetId="figure-17@30.[151,1435,181,1622]" captionTargetPageId="30" captionText="FIGURE 21. Vulva. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681977" httpUri="https://zenodo.org/record/12681977/files/figure.png" pageId="19" pageNumber="377">Fig. 21D</figureCitation>
).
</paragraph>
<paragraph id="8BCB36CCFFAD125728BEFE3D864B1EF9" blockId="19.[151,1437,150,1653]" box="[199,631,402,429]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728BEFE3D85271EF8" bold="true" box="[199,283,402,428]" pageId="19" pageNumber="377">Colour</emphasis>
. Light grey on dorsal surfaces.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125728BEFE1885E71D69" pageId="19" pageNumber="377" type="etymology">
<paragraph id="8BCB36CCFFAD125728BEFE1885E71D69" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728BEFE1885741E85" bold="true" box="[199,328,439,465]" pageId="19" pageNumber="377">Etymology</emphasis>
. The species is named after the late Ng Ngan Kee, an esteemed colleague and very good friend, departed all too soon in 2022. She was a leader in varunid systematics and also an avid educator and field scientist; the
<typeStatus id="54CF886EFFAD125728B8FE50851F1D4D" box="[193,291,511,537]" pageId="19" pageNumber="377" type="holotype">holotype</typeStatus>
specimen was actually collected by her during one of her many forays throughout Asia. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125728BEFDE886791979" pageId="19" pageNumber="377" type="discussion">
<paragraph id="8BCB36CCFFAD125728BEFDE8852A1CD5" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728BEFDE885081D35" bold="true" box="[199,308,583,609]" pageId="19" pageNumber="377">Remarks</emphasis>
. The difference in the shape of carapace, the P5 merus and the G1 between
<taxonomicName id="4C744D4FFFAD12572CF5FDE8813B1D35" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[1164,1287,583,609]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572CF5FDE8813B1D35" box="[1164,1287,583,609]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572D76FDE881721D35" bold="true" box="[1295,1358,583,609]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572D76FDE881721D35" box="[1295,1358,583,609]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C744D4FFFAD12572DFFFDE8851C1DD1" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="heterograna">
<emphasis id="B900EADEFFAD12572DFFFDE8851C1DD1" italics="true" pageId="19" pageNumber="377">L. heterograna</emphasis>
</taxonomicName>
are well defined. In
<taxonomicName id="4C744D4FFFAD12572A77FDC486B61DD1" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[526,650,619,645]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572A77FDC486B61DD1" box="[526,650,619,645]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572AEBFDC486EE1DD1" bold="true" box="[658,722,619,645]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572AEBFDC486EE1DD1" box="[658,722,619,645]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the carapace is relatively more rounded and more convex, the endostomial spine is strongly produced, clearly visible from dorsal view, the P5 merus leg is short, only 3.8 times as long as broad, the exopod of the third maxilliped is broad, strongly convex, and the apical process of G1 short, broad and bent at an angle of 60° along the longitudinal axis (
<figureCitation id="134F2A49FFAD12572B2EFD7887C91DA5" box="[855,1013,727,753]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1E–G, J</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C7BFD7980741DA5" box="[1026,1096,726,753]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C2CFD7880451DA5" box="[1109,1145,727,753]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="19" pageNumber="377">6D</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CFFFD7880E11DA5" box="[1158,1245,727,753]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">13 F, G</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C90FD7881201DA5" box="[1257,1308,727,753]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="19" pageNumber="377">14D</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D50FD7881641DA5" box="[1321,1368,727,753]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">16E</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D1CFD7884F21C41" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">18A, C‒E</figureCitation>
).
<taxonomicName id="4C744D4FFFAD1257289BFD5485F01C41" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[226,460,763,789]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" sensu="stricto" species="heterograna">
<emphasis id="B900EADEFFAD1257289BFD5485F01C41" box="[226,460,763,789]" italics="true" pageId="19" pageNumber="377">Lyphira heterograna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357A5FFAD125729ACFD54862E1C41" box="[469,530,763,789]" pageId="19" pageNumber="377" sensu="stricto">s. str.</taxonomicNameLabel>
has the carapace usually slightly longer than broad, the endostomial spines are short, the P5 merus is relatively longer 4.3 times as long as broad, the exopod of the third maxilliped is narrower, not as convex, and the G1 with apical process long, very slightly bent (
<figureCitation id="134F2A49FFAD12572BBEFCEC80711C09" box="[967,1101,835,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1A–C</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C23FCEC809D1C09" box="[1114,1185,835,861]" captionStart="FIGURE 2" captionStartId="4.[151,250,1924,1948]" captionTargetBox="[151,1435,813,1901]" captionTargetId="figure-285@4.[151,1435,813,1901]" captionTargetPageId="4" captionText="FIGURE 2. A, Lyphira heterograna (Ortmann, 1892), lectotype male (12.4 × 12.2 mm) (MZS Cru 3191) (specimen number Ortmann 658), Japan.A, overall dorsal habitus; B, overall dorsal habitus (carapace slightly tilted posteriorly); C, overall ventral habitus; D, anterior thoracic sternum, buccal cavity and third maxillipeds." figureDoi="http://doi.org/10.5281/zenodo.12681930" httpUri="https://zenodo.org/record/12681930/files/figure.png" pageId="19" pageNumber="377">2A, B</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CD6FCEC80C91C09" box="[1199,1269,835,861]" captionStart="FIGURE 3" captionStartId="7.[151,250,1818,1842]" captionTargetBox="[151,1435,273,1794]" captionTargetId="figure-45@7.[151,1435,273,1794]" captionTargetPageId="7" captionText="FIGURE 3. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. heterograna (Ortmann, 1892), male (8.8 × 9.3 mm) (NHM 1930.11.14.5) [paratype of Philyra peitaihoensis Shen, 1932], China; C, L.heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; D, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; E, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; F, L. chomel n. sp., paratype male (8.8 × 9.0 mm) (ZRC 1985.0091), Singapore; G, L. chomel n. sp., paratype male (12.6 × 12.8 mm) (ZRC 1985.0092), Singapore." figureDoi="http://doi.org/10.5281/zenodo.12681932" httpUri="https://zenodo.org/record/12681932/files/figure.png" pageId="19" pageNumber="377">3A, B</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D7AFCEC81091C09" box="[1283,1333,835,861]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">13A</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D3AFCEC81A41C09" box="[1347,1432,835,861]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">16A, B</figureCitation>
,
<figureCitation id="134F2A49FFAD125728EEFCC885361CD5" box="[151,266,871,897]" captionStart="FIGURE 17" captionStartId="26.[151,250,1698,1722]" captionTargetBox="[164,1422,191,1665]" captionTargetId="figure-17@26.[151,1435,181,1674]" captionTargetPageId="26" captionText="FIGURE 17. Gonopods.A‒G, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; H‒L, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; M–R, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia. A, H, M right third maxilliped (denuded); B, I, N, pleon; C, O, right G1 (ventral view); D, J, P, right G1 (dorsal view); E, Q, distal part of right G1 (ventral view); F, K, L, R, distal part of right G1 (dorsal view); G, right G2. Scales: A–J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681967" httpUri="https://zenodo.org/record/12681967/files/figure.png" pageId="19" pageNumber="377">17A, C‒F</figureCitation>
).
</paragraph>
<paragraph id="8BCB36CCFFAD125728BEFC2485BD1CB9" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD125728BEFC2485BF1CF1" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[199,387,907,933]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
<emphasis id="B900EADEFFAD125728BEFC2485BF1CF1" box="[199,387,907,933]" italics="true" pageId="19" pageNumber="377">Lyphira ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD125729F0FC2485F41CF1" bold="true" box="[393,456,907,933]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD125729F0FC2485F41CF1" box="[393,456,907,933]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is perhaps most similar to
<taxonomicName id="4C744D4FFFAD12572A8AFC2487761CF1" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[755,842,907,933]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAD12572A8AFC2487761CF1" box="[755,842,907,933]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572B29FC2487B21CF1" bold="true" box="[848,910,907,933]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572B29FC2487B21CF1" box="[848,910,907,933]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, sharing the characters of a strong endostomial spine and a wide exopod of the third maxilliped. The differences between the two are discussed under the remarks for the latter species.
</paragraph>
<paragraph id="8BCB36CCFFAD125728BEFC5886BD1A59" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD125728BEFC5885BB1B45" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[199,391,1015,1041]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="sp. nov.">
<emphasis id="B900EADEFFAD125728BEFC5885BB1B45" box="[199,391,1015,1041]" italics="true" pageId="19" pageNumber="377">Lyphira ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD125729EAFC5885EA1B45" bold="true" box="[403,470,1015,1041]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD125729EAFC5885EA1B45" box="[403,470,1015,1041]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C744D4FFFAD12572A6CFC58864C1B45" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[533,624,1015,1041]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAD12572A6CFC58864C1B45" box="[533,624,1015,1041]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572A02FC5886821B45" bold="true" box="[635,702,1015,1041]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572A02FC5886821B45" box="[635,702,1015,1041]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are superficially similar to
<taxonomicName id="4C744D4FFFAD12572C71FC5880541B44" authorityName="Galil" authorityYear="2009" box="[1032,1128,1015,1040]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B900EADEFFAD12572C71FC5880541B44" box="[1032,1128,1015,1040]" italics="true" pageId="19" pageNumber="377">L. ovata</emphasis>
</taxonomicName>
, described from
<specimenCount id="9D72FD45FFAD12572D4BFC5881A01B45" box="[1330,1436,1015,1041]" count="1" pageId="19" pageNumber="377" type="male">one male</specimenCount>
specimen from
<collectingCountry id="F363765CFFAD12572930FBB485811B61" box="[329,445,1051,1077]" name="Sri Lanka" pageId="19" pageNumber="377">Sri Lanka</collectingCountry>
.
<taxonomicName id="4C744D4FFFAD125729B0FBB486541B60" authorityName="Galil" authorityYear="2009" box="[457,616,1051,1077]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="B900EADEFFAD125729B0FBB486541B60" box="[457,616,1051,1077]" italics="true" pageId="19" pageNumber="377">Lyphira ovata</emphasis>
</taxonomicName>
, however, can easily be separated from these two species in having the dorsal carapace surface more densely covered with granules and the anterolateral margin is almost straight in dorsal view (cf.
<bibRefCitation id="EFE54B3DFFAD1257297DFBCC85BC1B29" author="Galil, B. S." box="[260,384,1123,1149]" pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: fig. 16A). In
<taxonomicName id="4C744D4FFFAD12572A58FBCC86A11B29" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[545,669,1123,1149]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572A58FBCC86A11B29" box="[545,669,1123,1149]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572ADCFBCC86D91B29" bold="true" box="[677,741,1123,1149]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572ADCFBCC86D91B29" box="[677,741,1123,1149]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C744D4FFFAD12572B67FBCC874A1B29" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[798,886,1123,1149]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAD12572B67FBCC874A1B29" box="[798,886,1123,1149]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572B07FBCC87821B29" bold="true" box="[894,958,1123,1149]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572B07FBCC87821B29" box="[894,958,1123,1149]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, there are distinctly fewer granules on the dorsal carapace surface and the anterolateral margin is distinctly convex in dorsal view (
<figureCitation id="134F2A49FFAD12572C1DFB28808F1BF5" box="[1124,1203,1159,1185]" captionStart="FIGURE 4" captionStartId="8.[151,250,1809,1833]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-17@8.[151,1435,181,1785]" captionTargetPageId="8" captionText="FIGURE 4. Overall dorsal habitus, Lyphira linda n. sp. A, holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; B, male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; C, male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; D, male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; E, male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; F, male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand." figureDoi="http://doi.org/10.5281/zenodo.12681934" httpUri="https://zenodo.org/record/12681934/files/figure.png" pageId="19" pageNumber="377">Figs. 4</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CC4FB29813D1BF5" box="[1213,1281,1158,1185]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
).
<bibRefCitation id="EFE54B3DFFAD12572D6CFB2884F71B91" author="Galil, B. S." pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil (2009: 303)</bibRefCitation>
did not figure the G1, only describing it as “with apical process digitate, squat”, suggesting it is similar to the condition observed for
<taxonomicName id="4C744D4FFFAD125729E3FB6086281BBD" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[410,532,1231,1257]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD125729E3FB6086281BBD" box="[410,532,1231,1257]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572A63FB6086641BBD" bold="true" box="[538,600,1231,1257]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572A63FB6086641BBD" box="[538,600,1231,1257]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C744D4FFFAD12572AF7FB6086D91BBD" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[654,741,1231,1257]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="linda" status="n. sp.">
<emphasis id="B900EADEFFAD12572AF7FB6086D91BBD" box="[654,741,1231,1257]" italics="true" pageId="19" pageNumber="377">L. linda</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572A92FB6087151BBD" bold="true" box="[747,809,1231,1257]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572A92FB6087151BBD" box="[747,809,1231,1257]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, in which the distal process is short and not prominently elongate (
<figureCitation id="134F2A49FFAD1257297CFB5C85F01A59" box="[261,460,1267,1293]" captionStart="FIGURE 18" captionStartId="27.[151,250,1824,1848]" captionTargetBox="[215,1373,194,1773]" captionTargetId="figure-20@27.[202,1386,181,1800]" captionTargetPageId="27" captionText="FIGURE 18. A–E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; F–K, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; L–P, L. linda n. sp., paratype male (14.2 × 13.9 mm) (ZRC 2022.1004), Singapore; Q–V, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand.A, F, Q, right third maxilliped (denuded); B, G, R, pleon; C, H, L, S, right G1 (dorsal view); D, I, M, T, U, distal part of right G1 (dorsal view); E, J, N, V, distal part of right G1 (ventral view); K, O, P distal part of right G1 (submesial view). Scales: A–C, F–H, L, Q–S = 1.0 mm; D, E, I–K, M–P, T–V = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681969" httpUri="https://zenodo.org/record/12681969/files/figure.png" pageId="19" pageNumber="377">Figs.18D, E, O, P</figureCitation>
,
<figureCitation id="134F2A49FFAD125729A1FB5C86491A59" box="[472,629,1267,1293]" captionStart="FIGURE 20" captionStartId="29.[151,250,1639,1663]" captionTargetBox="[252,1327,191,1613]" captionTargetId="figure-20@29.[239,1349,181,1615]" captionTargetPageId="29" captionText="FIGURE 20. G1. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; C, L. ngankee n. sp., paratype male (19.2 × 19.9 mm) (ZRC 2022.0628), Hong Kong; D, L. ngankee n. sp., paratype male (15.5 × 15.1 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (16.7 × 17.0 mm) (ZRC 2015.0439); Thailand; G, L. linda n. sp., male (18.4 × 19.0 mm) (ZRC 1985.1632), Singapore; H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. linda n. sp., male (18.4 × 18.6 mm) (ZRC 2003.0107), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (21.2 × 21.7 mm) (ZRC 2014.0191), Vietnam; M, L. linda n. sp., male (22.9 × 21.4 mm) (ZRC 2014.0191), Vietnam; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia; O, L. bellagrana n. sp., male (13.2 × 12.6 mm) (ZRC 2022.0745), Papua, Indonesia; P, L. bellagrana n. sp., male (10.6 × 9.9 mm) (ZRC 2022.0750), Papua, Indonesia. A, C, E–H, K–P, right G1; B, D, I, J, left G1 (laterally inverted)." figureDoi="http://doi.org/10.5281/zenodo.12681975" httpUri="https://zenodo.org/record/12681975/files/figure.png" pageId="19" pageNumber="377">20A‒D, E‒M</figureCitation>
).
</paragraph>
<paragraph id="8BCB36CCFFAD125728BEFAB886791979" blockId="19.[151,1437,150,1653]" pageId="19" pageNumber="377">
The wide exopod of the third maxilliped of
<taxonomicName id="4C744D4FFFAD12572AD7FAB887141A65" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[686,808,1303,1329]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572AD7FAB887141A65" box="[686,808,1303,1329]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572B56FAB887511A65" bold="true" box="[815,877,1303,1329]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572B56FAB887511A65" box="[815,877,1303,1329]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
similar to those of
<taxonomicName id="4C744D4FFFAD12572C3DFAB8808B1A64" authorityName="Trivedi, Soni &amp; Vachhrajani" authorityYear="2016" box="[1092,1207,1303,1328]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="B900EADEFFAD12572C3DFAB8808B1A64" box="[1092,1207,1303,1328]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D4FFFAD12572C95FAB881571A65" authorityName="Galil" authorityYear="2009" box="[1260,1387,1303,1329]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B900EADEFFAD12572C95FAB881571A65" box="[1260,1387,1303,1329]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
</taxonomicName>
, but differences can be seen in the more rounded carapace in
<taxonomicName id="4C744D4FFFAD12572B76FA9487B61A01" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[783,906,1339,1365]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572B76FA9487B61A01" box="[783,906,1339,1365]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572BE8FA9487F31A01" bold="true" box="[913,975,1339,1365]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572BE8FA9487F31A01" box="[913,975,1339,1365]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="134F2A49FFAD12572BA6FA9480031A01" box="[991,1087,1339,1365]" captionStart="FIGURE 1" captionStartId="3.[151,250,1368,1392]" captionTargetBox="[165,1421,188,1331]" captionTargetId="figure-20@3.[151,1435,181,1345]" captionTargetPageId="3" captionText="FIGURE 1. A–D, overall dorsal habitus Lyphira heterograna (Ortmann, 1892) (paratypes of Philyra peitaihoensis Shen, 1932; no size indicated); E–J, L. ngankee n. sp., (as Philyra tuberculosa Stimpson, 1858) from Hong Kong. A–D, F, after Shen (1932: text fig. 11); E, H–J, after Shen (1931: figs. 1, 4). A, E, overall dorsal habitus; B, F, dorsal view of frontal margin showing endostomial spines; C, right third maxilliped; D, lateral view of cephalothorax; G, third maxillipeds; H, male pleon; I, female pleon; J, distal part of G1." figureDoi="http://doi.org/10.5281/zenodo.12681928" httpUri="https://zenodo.org/record/12681928/files/figure.png" pageId="19" pageNumber="377">Figs. 1E</figureCitation>
,
<figureCitation id="134F2A49FFAD12572C32FA9580AC1A01" box="[1099,1168,1338,1365]" captionStart="FIGURE 5" captionStartId="9.[151,250,1246,1270]" captionTargetBox="[151,1435,181,1222]" captionTargetId="figure-20@9.[151,1435,181,1222]" captionTargetPageId="9" captionText="FIGURE 5. Overall dorsal habitus. A, Lyphira ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao, China; B, L. ngankee n. sp., paratype male (20.7 × 21.7 mm) (ZRC 2022.0628), Hong Kong; C, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; D, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681936" httpUri="https://zenodo.org/record/12681936/files/figure.png" pageId="19" pageNumber="377">5A, B</figureCitation>
,
<figureCitation id="134F2A49FFAD12572CE2FA9480821A01" box="[1179,1214,1339,1365]" captionStart="FIGURE 6" captionStartId="11.[151,250,1822,1846]" captionTargetBox="[242,1346,181,1798]" captionTargetId="figure-20@11.[242,1346,181,1798]" captionTargetPageId="11" captionText="FIGURE 6. Overall dorsal habitus. A, Lyphira heterograna (Ortmann, 1892), female (17.5 × 17.6 mm) (KPM NH 0103043), Japan; B, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; C, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; D, L. ngankee n. sp., Hong Kong, paratype female (18.8 × 18.8 mm) (ZRC 2022.0628), Hong Kong; E, L. linda n. sp., paratype female (15.5 × 16.1 mm) (ZRC 1981.9.2.10), Singapore; F, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.0317), Thailand; G, L. bellagrana n. sp., paratype female (13.7 × 12.7 mm) (ZRC 2000.2077), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681938" httpUri="https://zenodo.org/record/12681938/files/figure.png" pageId="19" pageNumber="377">6D</figureCitation>
) (longer than broad in
<taxonomicName id="4C744D4FFFAD125728CDFAF0851A1A2C" authorityName="Trivedi, Soni &amp; Vachhrajani" authorityYear="2016" box="[180,294,1375,1400]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="B900EADEFFAD125728CDFAF0851A1A2C" box="[180,294,1375,1400]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D4FFFAD12572925FAF085E71A2D" authorityName="Galil" authorityYear="2009" box="[348,475,1375,1401]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B900EADEFFAD12572925FAF085E71A2D" box="[348,475,1375,1401]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
</taxonomicName>
), the presence of strong endostomial spines in
<taxonomicName id="4C744D4FFFAD12572B90FAF0805F1A2D" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[1001,1123,1375,1401]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee">
<emphasis id="B900EADEFFAD12572B90FAF0805F1A2D" box="[1001,1123,1375,1401]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
(
<figureCitation id="134F2A49FFAD12572C0AFAF0813C1A2D" box="[1139,1280,1375,1401]" captionStart="FIGURE 13" captionStartId="22.[151,250,1685,1709]" captionTargetBox="[216,1372,181,1661]" captionTargetId="figure-17@22.[216,1372,181,1661]" captionTargetPageId="22" captionText="FIGURE 13. Dorsal view of frontal margin. A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), Yellow Sea, male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; C, L. acutidens (Chen, 1987), female (13.3 × 13.5 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. chomel n. sp., paratype female (11.4 × 11.1 mm) (ZRC 1985.98), Malaysia; F, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; G, L. ngankee n. sp., paratype male (20.7 × 21.3 mm) (ZRC 2022.0628), Hong Kong; H, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; I, L. linda n. sp., male (12.6 × 12.7 mm) (ZRC 1985.0097), Malaysia; J, L. linda n. sp., female (18.4 × 18.3 mm) (ZRC 1999.317), Thailand; K, L. linda n. sp., male (18.9 × 18.7 mm) (ZRC 1999.0507), Singapore; L, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; M, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; N, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681952" httpUri="https://zenodo.org/record/12681952/files/figure.png" pageId="19" pageNumber="377">Figs. 13F, G</figureCitation>
,
<figureCitation id="134F2A49FFAD12572D72FAF081011A2D" box="[1291,1341,1375,1401]" captionStart="FIGURE 14" captionStartId="23.[151,250,1808,1832]" captionTargetBox="[177,1410,181,1784]" captionTargetId="figure-20@23.[177,1410,181,1784]" captionTargetPageId="23" captionText="FIGURE 14. Endostomial spine.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPM NH 0130526), Japan; B, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea, female; C, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; D, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; E, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; F, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; G, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; H, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.12681955" httpUri="https://zenodo.org/record/12681955/files/figure.png" pageId="19" pageNumber="377">14D</figureCitation>
) (absent in
<taxonomicName id="4C744D4FFFAD125728CAFA2C851A1AC8" authorityName="Trivedi, Soni &amp; Vachhrajani" authorityYear="2016" box="[179,294,1411,1436]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="B900EADEFFAD125728CAFA2C851A1AC8" box="[179,294,1411,1436]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D4FFFAD12572922FA2C85E61AC9" authorityName="Galil" authorityYear="2009" box="[347,474,1411,1437]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B900EADEFFAD12572922FA2C85E61AC9" box="[347,474,1411,1437]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
</taxonomicName>
), and the P5 merus is proportionately shorter in
<taxonomicName id="4C744D4FFFAD12572B8DFA2C80521AC9" authorityName="Rahayu &amp; Ng" authorityYear="2024" box="[1012,1134,1411,1437]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee">
<emphasis id="B900EADEFFAD12572B8DFA2C80521AC9" box="[1012,1134,1411,1437]" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
, 3.8 times as long as broad (
<figureCitation id="134F2A49FFAD125728E6FA08853E1A95" box="[159,258,1447,1473]" captionStart="FIGURE 16" captionStartId="25.[151,250,1722,1746]" captionTargetBox="[151,1435,181,1698]" captionTargetId="figure-20@25.[151,1435,181,1698]" captionTargetPageId="25" captionText="FIGURE 16. P5.A, Lyphira heterograna (Ortmann, 1892), male (16.5 × 16.9 mm) (KPMNH 0130526), Japan; B,L. heterograna (Ortmann, 1892), male (14.2 × 14.4 mm) (NHM 2006.558), Korea; C, L. acutidens (Chen, 1987), male (9.3 × 9.1 mm) (ZRC 2002.0504), Yellow Sea; D, L. chomel n. sp., holotype male (10.1 × 10.3 mm) (ZRC 1999.9798), Malaysia; E, L. ngankee n. sp., holotype male (20.8 × 21.8 mm) (ZRC 1999.0456), Nanao; F, L. linda n. sp., holotype male (18.4 × 18.2 mm) (ZRC 1981.9.2.7), Singapore; G, H, L. linda n. sp., male (18.1 × 18.5 mm) (ZRC 2011.0646), Thailand; I, L. linda n. sp., male (17.4 × 17.1 mm) (ZRC 2003.0107), Thailand; J, L. bellagrana n. sp., holotype male (15.7 × 15.0 mm) (MZB Cru 5627), Papua, Indonesia; K, L. bellagrana n. sp., male (12.5 × 11.8 mm) (ZRC 2022.1031), Papua, Indonesia. A–G, I–K, right legs; H, left leg." figureDoi="http://doi.org/10.5281/zenodo.12681963" httpUri="https://zenodo.org/record/12681963/files/figure.png" pageId="19" pageNumber="377">Fig. 16E</figureCitation>
) (proportionately longer, being 4 times in
<taxonomicName id="4C744D4FFFAD12572AA2FA0887661A95" authorityName="Galil" authorityYear="2009" box="[731,858,1447,1473]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B900EADEFFAD12572AA2FA0887661A95" box="[731,858,1447,1473]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
</taxonomicName>
but much shorter in
<emphasis id="B900EADEFFAD12572C3BFA0880861A94" box="[1090,1210,1447,1472]" italics="true" pageId="19" pageNumber="377">
<taxonomicName id="4C744D4FFFAD12572C3BFA08808A1A94" authorityName="Trivedi, Soni &amp; Vachhrajani" authorityYear="2016" box="[1090,1206,1447,1472]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">L. georgei</taxonomicName>
,
</emphasis>
3.2 times as long as broad) (cf.
<bibRefCitation id="EFE54B3DFFAD1257296FFA6485AE1AB1" author="Galil, B. S." box="[278,402,1483,1509]" pageId="19" pageNumber="377" pagination="279 - 320" refId="ref19817" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: fig. 17A, B;
<bibRefCitation id="EFE54B3DFFAD12572A55FA64873E1AB1" author="Trivedi, J. N. &amp; Soni, G. M. &amp; Vachhrajani, K. D." box="[556,770,1483,1509]" pageId="19" pageNumber="377" pagination="148 - 154" refId="ref22214" refString="Trivedi, J. N., Soni, G. M. &amp; Vachhrajani, K. D. (2016) A new species of Lyphira Galil, 2009 (Crustacea, Decapoda, Leucosiidae) from Gujarat, India. Tropical Zoology, 29 (3), 148 - 154. https: // doi. org / 10.1080 / 03946975.2016.1181954" type="journal article" year="2016">
Trivedi
<emphasis id="B900EADEFFAD12572AFCFA6486821AB1" box="[645,702,1483,1509]" italics="true" pageId="19" pageNumber="377">et al.</emphasis>
2016
</bibRefCitation>
: figs. 1a, b). Their colours in life are also different, with
<taxonomicName id="4C744D4FFFAD12572DFFFA6484C8195D" authorityName="Rahayu &amp; Ng" authorityYear="2024" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="ngankee" status="n. sp.">
<emphasis id="B900EADEFFAD12572DFFFA6484C8195D" italics="true" pageId="19" pageNumber="377">L. ngankee</emphasis>
</taxonomicName>
<emphasis id="B900EADEFFAD12572882FA408505195D" bold="true" box="[251,313,1519,1545]" pageId="19" pageNumber="377">
<taxonomicNameLabel id="A23357A5FFAD12572882FA408505195D" box="[251,313,1519,1545]" pageId="19" pageNumber="377" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
being greyish white overall while the carapace of
<taxonomicName id="4C744D4FFFAD12572B1EFA4087E5195C" authorityName="Trivedi, Soni &amp; Vachhrajani" authorityYear="2016" box="[871,985,1519,1544]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="B900EADEFFAD12572B1EFA4087E5195C" box="[871,985,1519,1544]" italics="true" pageId="19" pageNumber="377">L. georgei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C744D4FFFAD12572C74FA4080B0195D" authorityName="Galil" authorityYear="2009" box="[1037,1164,1519,1545]" class="Malacostraca" family="Leucosiidae" genus="Lyphira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="377" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="B900EADEFFAD12572C74FA4080B0195D" box="[1037,1164,1519,1545]" italics="true" pageId="19" pageNumber="377">L. perplexa</emphasis>
</taxonomicName>
are orange to purple (cf.
<bibRefCitation id="EFE54B3DFFAD125728EEF9BC85571979" author="Trivedi, J. N. &amp; Soni, G. M. &amp; Vachhrajani, K. D." box="[151,363,1555,1581]" pageId="19" pageNumber="377" pagination="148 - 154" refId="ref22214" refString="Trivedi, J. N., Soni, G. M. &amp; Vachhrajani, K. D. (2016) A new species of Lyphira Galil, 2009 (Crustacea, Decapoda, Leucosiidae) from Gujarat, India. Tropical Zoology, 29 (3), 148 - 154. https: // doi. org / 10.1080 / 03946975.2016.1181954" type="journal article" year="2016">
Trivedi
<emphasis id="B900EADEFFAD12572889F9BC85141979" box="[240,296,1555,1581]" italics="true" pageId="19" pageNumber="377">et al.</emphasis>
2016
</bibRefCitation>
; unpublished data).
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125728BEF99886D01905" box="[199,748,1590,1617]" pageId="19" pageNumber="377" type="materials_examined">
<paragraph id="8BCB36CCFFAD125728BEF99886D01905" blockId="19.[151,1437,150,1653]" box="[199,748,1590,1617]" pageId="19" pageNumber="377">
<materialsCitation id="3B1C3C91FFAD125728BEF99886D41905" box="[199,744,1590,1617]" country="China" location="Nanao Island" pageId="19" pageNumber="377" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="B900EADEFFAD125728BEF998855C1904" bold="true" box="[199,352,1590,1617]" pageId="19" pageNumber="377">
<typeStatus id="54CF886EFFAD125728BEF998853E1905" box="[199,258,1591,1617]" pageId="19" pageNumber="377">Type</typeStatus>
locality
</emphasis>
.
<location id="8EAB6017FFAD12572917F99886391905" LSID="urn:lsid:plazi:treatment:03DD87DAFFAC125728EEFCB187691921:8EAB6017FFAD12572917F99886391905" box="[366,517,1591,1617]" country="China" name="Nanao Island" pageId="19" pageNumber="377" stateProvince="Guangdong">Nanao Island</location>
,
<collectingRegion id="49B0F82EFFAD12572A69F99886AA1905" box="[528,662,1591,1617]" country="China" name="Guangdong" pageId="19" pageNumber="377">Guangdong</collectingRegion>
,
<collectingCountry id="F363765CFFAD12572AD8F99886D41905" box="[673,744,1591,1617]" name="China" pageId="19" pageNumber="377">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36E6547FFAD125728BEF9F587691921" box="[199,853,1626,1653]" pageId="19" pageNumber="377" type="distribution">
<paragraph id="8BCB36CCFFAD125728BEF9F587691921" blockId="19.[151,1437,150,1653]" box="[199,853,1626,1653]" pageId="19" pageNumber="377">
<emphasis id="B900EADEFFAD125728BEF9F585651920" bold="true" box="[199,345,1626,1652]" pageId="19" pageNumber="377">Distribution</emphasis>
. Nanao Island, Guangdong, and
<collectingCountry id="F363765CFFAD12572AB3F9F4876D1921" box="[714,849,1627,1653]" name="Hong Kong" pageId="19" pageNumber="377">Hong Kong</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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