292 lines
39 KiB
XML
292 lines
39 KiB
XML
<document ID-DOI="10.26107/RBZ-2021-0072" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174571" ID-ZooBank="5BE7279D-C928-4BA7-8445-4C4044AA1AEA" approvalRequired="4" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="3" checkinTime="1665291265405" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Chanidaporn & Tumpeesuwan, Sakboworn" docDate="2021" docId="3C0EE70A334CFFCCCEFBF833A2A46E70" docLanguage="en" docName="RafflesBZool.69.555–569.pdf" docOrigin="Raffles Bulletin of Zoology 69" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.4:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="4" docTitle="Anceyoconcha carinata Nahok & Tumpeesuwan & Tumpeesuwan 2021, new species" docType="treatment" docVersion="2" lastPageNumber="561" masterDocId="C0379F723348FFCACD4EFA4FA0776B45" masterDocTitle="Two new species of genus Anceyoconcha S. Tumpeesuwan & C. Tumpeesuwan, in Nahok et al., 2020 (Gastropoda: Pulmonata: Camaenidae), from northeastern Thailand" masterLastPageNumber="569" masterPageNumber="555" pageNumber="559" updateTime="1665307449397" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of genus Anceyoconcha S. Tumpeesuwan & C. Tumpeesuwan, in Nahok et al., 2020 (Gastropoda: Pulmonata: Camaenidae), from northeastern Thailand</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nahok, Benchawan</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tumpeesuwan, Chanidaporn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>555</mods:start>
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<mods:end>569</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.26107/RBZ-2021-0072</mods:identifier>
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<mods:identifier type="ISSN">2345-7600</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7174571</mods:identifier>
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<mods:identifier type="ZooBank">5BE7279D-C928-4BA7-8445-4C4044AA1AEA</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:3C0EE70A334CFFCCCEFBF833A2A46E70" httpUri="http://treatment.plazi.org/id/3C0EE70A334CFFCCCEFBF833A2A46E70" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559">
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<subSubSection box="[949,1360,636,661]" pageId="4" pageNumber="559" type="nomenclature">
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<paragraph blockId="4.[949,1360,636,693]" box="[949,1360,636,661]" pageId="4" pageNumber="559">
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<heading bold="true" box="[949,1360,636,661]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="559" reason="2">
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<emphasis bold="true" box="[949,1360,636,661]" pageId="4" pageNumber="559">
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<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[949,1212,636,661]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="559" phylum="Mollusca" rank="species" species="carinata" status="new species">
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<emphasis bold="true" box="[949,1212,636,661]" italics="true" pageId="4" pageNumber="559">Anceyoconcha carinata</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[1227,1360,637,661]" pageId="4" pageNumber="559" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="559" type="description">
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<paragraph blockId="4.[949,1360,636,693]" box="[1000,1309,669,693]" pageId="4" pageNumber="559">
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(
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<figureCitation box="[1008,1138,669,693]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="4" pageNumber="559">Figs. 3A, B</figureCitation>
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,
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||
<figureCitation box="[1151,1167,669,693]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="4" pageNumber="559">4</figureCitation>
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||
;
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||
<tableCitation box="[1181,1300,669,693]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="2.[135,190,188,210]" captionStartId-1="3.[113,168,189,211]" captionStartId-2="7.[111,166,599,622]" captionText-0="Table 1. Specimens included for molecular analysis, with their GenBank accession numbers. Species identities of sequences obtained from GenBank follow the respective sources." captionText-1="Table 2. Pairwise p-distance values for different species in this study based on mitochondrial COI sequences." captionText-2="Table 3. Comparison of shells, radulae, and genital systems of Anceyoconcha spp. from northeastern Thailand." pageId="4" pageNumber="559">Tables 1–3</tableCitation>
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)
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</paragraph>
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<paragraph blockId="4.[835,1474,731,782]" pageId="4" pageNumber="559">
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<taxonomicName box="[835,1023,731,753]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="559" phylum="Mollusca" rank="species" species="undefined-1">
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<emphasis box="[835,975,731,752]" italics="true" pageId="4" pageNumber="559">Anceyoconcha</emphasis>
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sp.1
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</taxonomicName>
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– Nahok, 2020: 72–73, figs. 25C, D, 39, 46, 47, 50, 53, tables 3, 5, 7–10.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="559" type="materials_examined">
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||
<paragraph blockId="4.[835,1474,822,1135]" pageId="4" pageNumber="559">
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||
<emphasis bold="true" box="[835,1059,822,846]" pageId="4" pageNumber="559">Material examined.</emphasis>
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||
<materialsCitation collectingDate="2017-05-30" country="Thailand" elevation="211" latitude="14.482044" location="Khao Plai Bat" longLatPrecision="1" longitude="102.96972" municipality="Prakhon Chai District" pageId="4" pageNumber="559" specimenCount="1" stateProvince="Buriram" typeStatus="holotype">
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||
<typeStatus box="[1067,1170,822,846]" pageId="4" pageNumber="559">Holotype</typeStatus>
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: NHMSU-00036,
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<location LSID="urn:lsid:plazi:treatment:3C0EE70A334CFFCCCEFBF833A2A46E70:B17800C7334CFFCEC819F979A31F682B" country="Thailand" latitude="14.482044" longLatPrecision="1" longitude="102.96972" municipality="Prakhon Chai District" name="Khao Plai Bat" pageId="4" pageNumber="559" stateProvince="Buriram">Khao Plai Bat</location>
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||
(basalt hill),
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||
<collectingMunicipality box="[1015,1251,854,878]" pageId="4" pageNumber="559">Prakhon Chai District</collectingMunicipality>
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||
,
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||
<collectingRegion box="[1261,1471,854,878]" country="Thailand" name="Buriram" pageId="4" pageNumber="559">Buri Ram Province</collectingRegion>
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||
,
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||
<collectingCountry box="[835,930,886,910]" name="Thailand" pageId="4" pageNumber="559">Thailand</collectingCountry>
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||
(
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||
<geoCoordinate box="[947,1111,885,910]" degrees="14" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="559" precision="1" seconds="55.36" value="14.482044">14°28′55.36″N</geoCoordinate>
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,
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<geoCoordinate box="[1124,1298,885,910]" degrees="102" direction="east" minutes="58" orientation="longitude" pageId="4" pageNumber="559" precision="1" seconds="11.00" value="102.96972">102°58′11.00″E</geoCoordinate>
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), alt.
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<quantity box="[1362,1434,887,911]" metricMagnitude="2" metricUnit="m" metricValue="2.11" pageId="4" pageNumber="559" unit="m" value="211.0">
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<elevation box="[1362,1434,887,911]" metricMagnitude="2" metricUnit="m" metricValue="2.11" pageId="4" pageNumber="559" unit="m" value="211.0">211 m</elevation>
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||
</quantity>
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||
,
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||
<date pageId="4" pageNumber="559" value="2017-05-30">
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<collectingDate pageId="4" pageNumber="559" value="2017-05-30">30 May 2017</collectingDate>
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||
</date>
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, SH = 19.0 mm, SW =
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<quantity box="[1212,1298,918,942]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="4" pageNumber="559" unit="mm" value="7.6">7.6 mm</quantity>
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||
, AH =
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||
<quantity box="[1382,1468,918,942]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="559" unit="mm" value="5.4">5.4 mm</quantity>
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||
, AW =
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||
<quantity box="[918,1007,951,975]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="559" unit="mm" value="3.3">3.3 mm</quantity>
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(
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||
<figureCitation box="[1027,1122,950,975]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="4" pageNumber="559">Fig. 3A</figureCitation>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation collectingDate="2017-05-31" country="Thailand" elevation="251" latitude="14.540934" location="Khao Angkhan" longLatPrecision="1" longitude="102.83711" municipality="Chaloem Phra Kiat District" pageId="4" pageNumber="559" specimenCount="1" stateProvince="Buriram" typeStatus="paratype">
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||
<typeStatus box="[1146,1260,951,975]" pageId="4" pageNumber="559">Paratypes</typeStatus>
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||
: 6 shells, 4 living specimens preserved in ethanol (NHMSU-00037), same data as for holotype. 1 shell (NHMSU-00038),
|
||
<location LSID="urn:lsid:plazi:treatment:3C0EE70A334CFFCCCEFBF833A2A46E70:B17800C7334CFFCEC852F9B9A5B56F4B" box="[1308,1474,1014,1038]" country="Thailand" latitude="14.540934" longLatPrecision="1" longitude="102.83711" municipality="Chaloem Phra Kiat District" name="Khao Angkhan" pageId="4" pageNumber="559" stateProvince="Buriram">Khao Angkhan</location>
|
||
(basalt hill),
|
||
<collectingMunicipality box="[967,1256,1046,1070]" pageId="4" pageNumber="559">Chaloem Phra Kiat District</collectingMunicipality>
|
||
,
|
||
<collectingRegion box="[1265,1470,1046,1070]" country="Thailand" name="Buriram" pageId="4" pageNumber="559">Buri Ram Province</collectingRegion>
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||
,
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||
<collectingCountry box="[835,930,1078,1102]" name="Thailand" pageId="4" pageNumber="559">Thailand</collectingCountry>
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||
(
|
||
<geoCoordinate box="[947,1110,1077,1102]" degrees="14" direction="north" minutes="32" orientation="latitude" pageId="4" pageNumber="559" precision="1" seconds="27.36" value="14.540934">14°32′27.36″N</geoCoordinate>
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||
,
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||
<geoCoordinate box="[1124,1298,1077,1102]" degrees="102" direction="east" minutes="50" orientation="longitude" pageId="4" pageNumber="559" precision="1" seconds="13.60" value="102.83711">102°50′13.60″E</geoCoordinate>
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||
), alt.
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||
<quantity box="[1362,1434,1078,1102]" metricMagnitude="2" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="559" unit="m" value="251.0">
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||
<elevation box="[1362,1434,1078,1102]" metricMagnitude="2" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="559" unit="m" value="251.0">251 m</elevation>
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||
</quantity>
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||
,
|
||
<date pageId="4" pageNumber="559" value="2017-05-31">
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||
<collectingDate pageId="4" pageNumber="559" value="2017-05-31">31 May 2017</collectingDate>
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||
</date>
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||
(
|
||
<figureCitation box="[963,1048,1110,1135]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="4" pageNumber="559">Fig. 3B</figureCitation>
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)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="559" type="diagnosis">
|
||
<paragraph blockId="4.[835,1474,1174,1295]" pageId="4" pageNumber="559">
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<emphasis bold="true" box="[835,952,1174,1198]" pageId="4" pageNumber="559">Diagnosis.</emphasis>
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Shell slightly ovate conic, last whorl with strong ridge around umbilicus (
|
||
<figureCitation box="[1090,1202,1206,1231]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="4" pageNumber="559">Fig. 3A, B</figureCitation>
|
||
). Penis and vagina shorter than flagellum. Flagellum long cylindrical, divided into two portions (
|
||
<figureCitation box="[939,1059,1270,1295]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="4" pageNumber="559">Fig. 4A, B</figureCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="559" type="etymology">
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||
<paragraph blockId="4.[835,1474,1334,1422]" pageId="4" pageNumber="559">
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||
<emphasis bold="true" box="[835,965,1335,1359]" pageId="4" pageNumber="559">Etymology.</emphasis>
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Specific epithet ‘carinata’ derived from Latin word ‘carina’, meaning ‘keel’ and referring to the shell of this new species having a periumbilical keel.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559" type="description">
|
||
<paragraph blockId="4.[835,1475,1462,1774]" pageId="4" pageNumber="559">
|
||
<emphasis bold="true" box="[835,972,1462,1486]" pageId="4" pageNumber="559">Description.</emphasis>
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Shell (
|
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<figureCitation box="[1044,1155,1462,1487]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="4" pageNumber="559">Fig. 3A, B</figureCitation>
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||
;
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<tableCitation box="[1165,1245,1462,1487]" captionStart="Table 3" captionStartId="7.[111,166,599,622]" captionText="Table 3. Comparison of shells, radulae, and genital systems of Anceyoconcha spp. from northeastern Thailand." pageId="4" pageNumber="559">Table 3</tableCitation>
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): slightly ovate conic, whorls 6½, shell height 18.5–22.0 mm, shell width
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.8" metricValueMin="7.2" pageId="4" pageNumber="559" unit="mm" value="8.0" valueMax="8.8" valueMin="7.2">7.2–8.8 mm</quantity>
|
||
, aperture height
|
||
<quantity box="[1053,1182,1526,1550]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.1" metricValueMin="5.4" pageId="4" pageNumber="559" unit="mm" value="6.25" valueMax="7.1" valueMin="5.4">5.4–7.1 mm</quantity>
|
||
, and aperture width
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="6.1" metricValueMin="3.3" pageId="4" pageNumber="559" unit="mm" value="4.699999999999999" valueMax="6.1" valueMin="3.3">3.3–6.1 mm</quantity>
|
||
. Brownish colour; semi-transparent and lustrous. Apex obtuse, acute with light colour; embryonic shell smooth. Last whorl straight, rounded; with a periumbilical keel near aperture. Peripheral keel and peripheral band absent. Aperture large, irregularly rounded or ovoid, quite oblique, with thin, reflexed margins. Peristome expanded and reflexed. Umbilicus, a minute perforation, narrow and deep.
|
||
</paragraph>
|
||
<paragraph blockId="4.[834,1475,1814,2094]" lastBlockId="6.[136,778,189,437]" lastPageId="6" lastPageNumber="561" pageId="4" pageNumber="559">
|
||
Genital system (n = 3) (
|
||
<figureCitation box="[1084,1199,1814,1839]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="4" pageNumber="559">Fig. 4A–F</figureCitation>
|
||
): atrium short. Penis short cylindrical. Epiphallus divided into two portions, proximal part (ep1) stout and swollen, distal part (ep2) long cylindrical. Penial retractor muscle present. Flagellum approximately almost the same length as distal part of epiphallus (ep2); with proximal part twisted and distal part S-shaped. Vas deferens long. Vagina short cylindrical but longer than penis. Gametolytic duct thickened at base and gradually slender to small tube distally. Gametolytic sac small swollen gland at distal end of gametolytic duct. Free oviduct short. Uterus and prostate gland long and stout. Inner wall of penis with short, stout, and smooth longitudinal pilaster (
|
||
<figureCitation box="[674,765,253,277]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4C</figureCitation>
|
||
). Inner wall of ep1 with four corrugated longitudinal pilasters (
|
||
<figureCitation box="[144,230,317,341]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4C</figureCitation>
|
||
); inner wall of ep2 with three smooth longitudinal pilasters (
|
||
<figureCitation box="[241,329,349,373]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4D</figureCitation>
|
||
). Inner wall of flagellum with three stout and smooth longitudinal pilasters (
|
||
<figureCitation box="[521,609,381,405]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4E</figureCitation>
|
||
). Inner wall of vagina with two longitudinal pilasters (
|
||
<figureCitation box="[563,647,413,437]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7174577" ID-Zenodo-Dep="7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="5" pageNumber="560" startId="5.[113,148,1898,1920]" targetBox="[113,1451,268,1883]" targetPageId="5">
|
||
<paragraph blockId="5.[113,1452,1898,2035]" pageId="5" pageNumber="560">
|
||
Fig. 3. Shell morphology of
|
||
<taxonomicName box="[395,583,1898,1920]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="560" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis box="[395,535,1898,1919]" italics="true" pageId="5" pageNumber="560">Anceyoconcha</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
in northeastern Thailand. A, B,
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[903,1012,1898,1920]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="560" phylum="Mollusca" rank="species" species="carinata" status="new species">
|
||
<emphasis box="[903,1012,1898,1920]" italics="true" pageId="5" pageNumber="560">A. carinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[1026,1146,1898,1920]" pageId="5" pageNumber="560" rank="species">new species</taxonomicNameLabel>
|
||
; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E,
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[1061,1178,1927,1948]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="560" phylum="Mollusca" rank="species" species="elongata" status="new species">
|
||
<emphasis box="[1061,1076,1927,1948]" italics="true" pageId="5" pageNumber="560">A</emphasis>
|
||
.
|
||
<emphasis box="[1091,1178,1927,1948]" italics="true" pageId="5" pageNumber="560">elongata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[1194,1319,1927,1949]" pageId="5" pageNumber="560" rank="species">new species</taxonomicNameLabel>
|
||
; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F,
|
||
<emphasis box="[205,220,1985,2006]" italics="true" pageId="5" pageNumber="560">A</emphasis>
|
||
. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G,
|
||
<taxonomicName authorityName="Redfield" authorityYear="1853" box="[765,886,1985,2006]" class="Gastropoda" family="Camaenidae" genus="Bulimus" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="560" phylum="Mollusca" rank="species" species="siamensis">
|
||
<emphasis box="[765,780,1985,2006]" italics="true" pageId="5" pageNumber="560">A</emphasis>
|
||
.
|
||
<emphasis box="[793,886,1985,2006]" italics="true" pageId="5" pageNumber="560">siamensis</emphasis>
|
||
</taxonomicName>
|
||
(ZRCBUU-0325). H,
|
||
<taxonomicName box="[1107,1267,1985,2006]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="560" phylum="Mollusca" rank="species" species="rhombostoma">
|
||
<emphasis box="[1107,1122,1985,2006]" italics="true" pageId="5" pageNumber="560">A</emphasis>
|
||
.
|
||
<emphasis box="[1135,1267,1985,2006]" italics="true" pageId="5" pageNumber="560">rhombostoma</emphasis>
|
||
</taxonomicName>
|
||
(NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[136,778,477,725]" pageId="6" pageNumber="561">
|
||
Radula (n = 3) (
|
||
<figureCitation box="[315,427,477,501]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4G–I</figureCitation>
|
||
): comprises 126–128 transverse rows of teeth, each row containing 65–73 teeth, (22–24) + (10–12) + 1 + (10–12) + (22–24). Central teeth symmetric and unicuspid, narrowly rounded. Lateral teeth oblique rounded, bicuspid with indistinct or distinct blunt ectocone (
|
||
<figureCitation box="[145,230,637,661]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4H</figureCitation>
|
||
), larger than central teeth. Marginal teeth gradually changing from broadly tricuspid (13th–28th) to tetracuspid (29th–36th) with two small ectocones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="561" type="discussion">
|
||
<paragraph blockId="6.[135,777,765,1205]" pageId="6" pageNumber="561">
|
||
<emphasis bold="true" box="[136,254,765,789]" pageId="6" pageNumber="561">Remarks.</emphasis>
|
||
The shell morphology of
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[575,710,766,789]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="561" phylum="Mollusca" rank="species" species="carinata" status="new species">
|
||
<emphasis box="[575,591,766,789]" italics="true" pageId="6" pageNumber="561">A</emphasis>
|
||
.
|
||
<emphasis box="[612,710,766,789]" italics="true" pageId="6" pageNumber="561">carinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel pageId="6" pageNumber="561" rank="species">new species</taxonomicNameLabel>
|
||
, differs from that of many other congeners in the presence of a strong keel around the umbilicus (
|
||
<figureCitation box="[647,762,829,853]" captionStart="Fig" captionStartId="5.[113,148,1898,1920]" captionTargetBox="[113,1451,268,1883]" captionTargetId="figure-15@5.[111,1453,268,1883]" captionTargetPageId="5" captionText="Fig. 3. Shell morphology of Anceyoconcha spp. in northeastern Thailand. A, B, A. carinata, new species; A, holotype (NHMSU-00036) from Khao Plai Bat, Buri Ram; B (NHMSU-00038) from Khao Angkhan, Buri Ram. C–E, A. elongata, new species; C, holotype (NHMSU-00039) from Phu Po, Kalasin; D (NHMSU-00041) from Phu No, Kalasin; E (NHMSU-00045) from Phanom Sawai Forest Park, Surin. F, A. sp.1 (NHMSU-00043) from Pha Nam Yoi, Roi Et. G, A. siamensis (ZRCBUU-0325). H, A. rhombostoma (NHMSU-00022). Fig. 3G photograph courtesy of Pongrat Dumrongrojwattana." figureDoi="http://doi.org/10.5281/zenodo.7174577" httpUri="https://zenodo.org/record/7174577/files/figure.png" pageId="6" pageNumber="561">Fig. 3A, B</figureCitation>
|
||
).
|
||
<taxonomicName box="[136,445,861,885]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="561" phylum="Mollusca" rank="species" species="rhombostoma">
|
||
<emphasis box="[136,445,861,885]" italics="true" pageId="6" pageNumber="561">Anceyoconcha rhombostoma</emphasis>
|
||
</taxonomicName>
|
||
also possesses the keel, but its shell is ovate trochoid, shorter than that of
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[597,718,894,917]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="561" phylum="Mollusca" rank="species" species="carinata" status="new species">
|
||
<emphasis box="[597,718,894,917]" italics="true" pageId="6" pageNumber="561">A. carinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel pageId="6" pageNumber="561" rank="species">new species</taxonomicNameLabel>
|
||
, and the presence of a keel on its periphery runs to an angle of the rhomboid-shaped aperture (see
|
||
<bibRefCitation author="Sutcharit C & Backeljau T & Panha S" pageId="6" pageNumber="561" pagination="51 - 76" refId="ref10265" refString="Sutcharit C, Backeljau T & Panha S (2019) Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae). ZooKeys, 870: 51 - 76. https: // doi. org / 10.3897 / zookeys. 870.36970." type="journal article" year="2019">Sutcharit et al., 2019</bibRefCitation>
|
||
: fig. 3C–I;
|
||
<bibRefCitation author="Nahok B & Tumpeesuwan S & Tumpeesuwan C" box="[364,573,989,1013]" pageId="6" pageNumber="561" pagination="80 - 90" refId="ref9557" refString="Nahok B, Tumpeesuwan S & Tumpeesuwan C (2020) Anceyoconcha, a replacement name for the preoccupied tree snail genus Giardia Ancey, 1907 (Pulmonata: Helicoidea: Camaenidae). Raffles Bulletin of Zoology, 68: 80 - 90. https: // doi. org / 10.26107 / RBZ- 2020 - 0009." type="journal article" year="2020">Nahok et al., 2020</bibRefCitation>
|
||
: fig. 4B;
|
||
<bibRefCitation author="Sutcharit C & Thach P & Chhuoy S & Ngor PB & Jeratthitikul E & Siriwut W & Srisonchai R & Ng TH & Pholyotha A & Jirapatrasilp P & Panha S" pageId="6" pageNumber="561" pagination="1 - 46" refId="ref10365" refString="Sutcharit C, Thach P, Chhuoy S, Ngor PB, Jeratthitikul E, Siriwut W, Srisonchai R, Ng TH, Pholyotha A, Jirapatrasilp P & Panha S (2020) Annotated checklist of the land snail fauna from southern Cambodia (Mollusca, Gastropoda). ZooKeys, 948: 1 - 46. https: // doi. org / 10.3897 / zookeys. 948.51671." type="journal article" year="2020">Sutcharit et al., 2020</bibRefCitation>
|
||
: fig. 12C, D). The genitalia of
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[596,716,1022,1045]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="561" phylum="Mollusca" rank="species" species="carinata" status="new species">
|
||
<emphasis box="[596,716,1022,1045]" italics="true" pageId="6" pageNumber="561">A. carinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel pageId="6" pageNumber="561" rank="species">new species</taxonomicNameLabel>
|
||
, possess three characters that are distinct from those of all other congeners in that the vagina and penis are shorter than the flagellum, and the flagellum is divided into two sections with a twining proximal part and S-shaped distal part (
|
||
<figureCitation box="[194,314,1181,1205]" captionStart="Fig" captionStartId="8.[135,170,2006,2028]" captionTargetBox="[213,1396,188,1991]" captionTargetId="figure-8@8.[213,1396,188,1994]" captionTargetPageId="8" captionText="Fig. 4. Anceyoconcha carinata, new species. A, B, genital system, paratype (NHMSU-00037). C–F, inner wall of genital system; C, penis (p) and proximal portion of epiphallus (ep1); D, distal portion of epiphallus (ep2); E, flagellum (fl); F, vagina (v). G, view of transverse rows of radula, with number of lateral and marginal teeth indicated (c = central tooth); H, right side of central tooth and lateral teeth; I, right side of marginal teeth. J, natural habitat and living adult." figureDoi="http://doi.org/10.5281/zenodo.7174579" httpUri="https://zenodo.org/record/7174579/files/figure.png" pageId="6" pageNumber="561">Fig. 4A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="561" type="distribution">
|
||
<paragraph blockId="6.[136,775,1245,1333]" pageId="6" pageNumber="561">
|
||
<emphasis bold="true" box="[136,293,1245,1269]" pageId="6" pageNumber="561">Distribution.</emphasis>
|
||
<taxonomicName authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[304,578,1245,1269]" class="Gastropoda" family="Camaenidae" genus="Anceyoconcha" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="561" phylum="Mollusca" rank="species" species="carinata" status="new species">
|
||
<emphasis box="[304,578,1245,1269]" italics="true" pageId="6" pageNumber="561">Anceyoconcha carinata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[597,741,1245,1269]" pageId="6" pageNumber="561" rank="species">new species</taxonomicNameLabel>
|
||
, is currently known from the basalt hill areas in Prakhon Chai and Chaloem Phra Kiat Districts,
|
||
<collectingRegion box="[506,719,1309,1333]" country="Thailand" name="Buriram" pageId="6" pageNumber="561">Buri Ram Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |