treatments-xml/data/1B/B7/4B/1BB74B62D9525B7A88A89E7FCBC129FE.xml
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<document id="9C3C5588F4393E0C7F10E9BA43BA380F" ID-DOI="10.3897/zookeys.1202.121633" ID-ZooBank="F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" ID-publisher-id="121633" URI-arpha="CB9CB628-9D1E-53A4-84B5-3B9C013566B4" URI-zoobank="http://zoobank.org/F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1716826741008" checkinUser="pensoft" docAuthor="Zhang, Ludan, Wu, Zheng, Li, Shuqiang &amp; Yao, Zhiyuan" docDate="2024" docId="1BB74B62D9525B7A88A89E7FCBC129FE" docLanguage="en" docName="ZooKeys 1202: 255-286" docOrigin="ZooKeys 1202" docSource="https://zookeys.pensoft.net/article/121633/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana yunnan Zhang, Li &amp; Yao 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="286" masterDocId="CB9CB6289D1E53A484B53B9C013566B4" masterDocTitle="Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China" masterLastPageNumber="286" masterPageNumber="255" pageNumber="255" updateTime="1716918634710" updateUser="admin">
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<mods:title id="20967839E2D2992EA074BA52D0FCD4DB">Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China</mods:title>
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<mods:namePart id="6C9A4DCB9F2A838709F1601D04CA2A04">Zhang, Ludan</mods:namePart>
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<mods:namePart id="978928890BE582D3677915BA9F1F3AD2">Wu, Zheng</mods:namePart>
<mods:affiliation id="531524247630DB509992D5110D4AFAE3">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="81BB350C79D8FD1366AB4FB98AA4A5BE">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="EA224D647764C0AE408BDBCC6E52BF41">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:namePart id="8239F85B173BB4EB3495C0A4DA09E02C">Yao, Zhiyuan</mods:namePart>
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<mods:affiliation id="3C08DE08E28F42A510BCCD10234A329F">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:date id="01EA7B0407AAAA42634236CC65BBC6DB">2024</mods:date>
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<treatment id="1BB74B62D9525B7A88A89E7FCBC129FE" ID-DOI="http://doi.org/10.5281/zenodo.11356058" ID-Zenodo-Dep="11356058" ID-arpha="1BB74B62-D952-5B7A-88A8-9E7FCBC129FE" ID-zoobank="https://zoobank.org/757F74E0-B68C-47E6-9260-5061A26D54A4" LSID="urn:lsid:plazi:treatment:1BB74B62D9525B7A88A89E7FCBC129FE" LSID-ZBK="urn:lsid:zoobank.org:act:757F74E0-B68C-47E6-9260-5061A26D54A4" httpUri="http://treatment.plazi.org/id/1BB74B62D9525B7A88A89E7FCBC129FE">
<subSubSection id="16D2078C82263F353ACEAAA02D0FB2A9" type="nomenclature">
<paragraph id="E272C03D6CD2184BB3F1F79DF2A2D728">
<taxonomicName id="8F919DB2BB9D62D6484A6020F0A3153A" authority="Zhang, Li &amp; Yao" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yunnan" status="sp. nov.">
<emphasis id="B051D27B50332BBA0517745032584C6B" italics="true">Belisana yunnan</emphasis>
Zhang, Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="068421A8DEEBAD84876E7AD01531266C" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4536306038EC2A19F36AD7B1797ABE55" type="description">
<paragraph id="A180A3F45EC4C1D881679C86BA3E6612">
<figureCitation id="2540049BA006FE0C862698972A627581" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">Figs 16</figureCitation>
,
<figureCitation id="6BB1CCA8553367A65B196558B3ED885A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17</figureCitation>
,
<figureCitation id="9BED52A9677073712986F286C6FFEDF8" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 C, D</figureCitation>
,
<figureCitation id="69E665D2D2BC952C8BE14D013D1883C9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJYAI" type="type material">
<paragraph id="84C9B5DA1ADF89FB0C5C6578DE2658E6">
<heading id="DF3DA4BA33A3274DADE70AEB48845180" reason="title">Type material.</heading>
</paragraph>
<paragraph id="E21AECF108FFDB710CB3ED778627A902">
<materialsCitation id="C6592BF5CAC2BD9D7E9DDE1B067D10B2" collectingDate="2010-08-24" collectionCode="IZCAS" collectorName="C Wang &amp; Q Zhao &amp; L Lin" country="China" county="Dehong" elevation="1636" latitude="24.23215" location="Xianren Cave" longLatPrecision="1" longitude="98.42605" municipality="Luxi" specimenCode="Ar 44981, Ar 44982-83, Ar 44984-86" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="A6793780D7283ADE760107E652147A99" bold="true" italics="true">
<typeStatus id="B6E8E9903A14E6A975FAE77780095AFC" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="5D7E0EE81E178093D1BF55541E9F5DF1" count="1" type="male"></specimenCount>
(
<named-content id="740069364378EF356F956FFB795557D9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="83373B970DEA39D4E2EBCE540E04DA7F" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="55ED1DBC02A2D6C78DEF420DE6CC03D6">Ar 44981</specimenCode>
) and
<emphasis id="1308200AB7C119D7A5BEFF49573DB0BD" bold="true">
<emphasis id="7676012240C03788FD80B03577338A0E" bold="true" italics="true">
<typeStatus id="52C1F8DC3CB4A68089E4BFF09BE5B6F7" type="paratype">paratype</typeStatus>
</emphasis>
s
</emphasis>
<specimenCount id="AB1A9777722C3F022207F1F8848ABBF6" count="2" type="male">2 ♂</specimenCount>
(
<named-content id="5B0D82785A1DC294D5FEC77AD2F50D72" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="646DD70B8059EDA54127DE5FDDE5682A" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="66162D189F13CBB486ED948ED4105E4B">Ar 4498283</specimenCode>
)
<specimenCount id="D8398A6822563B2FB6B91F0B1A8E065D" count="3" type="female">3 ♀</specimenCount>
(
<named-content id="1C2D8D5E6185F618E6FF324CBCE8E3CE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="650546C4B2053AAAA4040B09337680FC" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="0A44ECC5DD8C256323EFB88885F8A6DF">Ar 4498486</specimenCode>
),
<location id="CA542470E7E864F24573A68A1F7F1F19" LSID="urn:lsid:plazi:treatment:1BB74B62D9525B7A88A89E7FCBC129FE:CA542470E7E864F24573A68A1F7F1F19" country="China" county="Dehong" latitude="24.23215" longLatPrecision="1" longitude="98.42605" municipality="Luxi" name="Xianren Cave" stateProvince="Yunnan">Xianren Cave</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHZAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[98.426050,24.232150]}">
<geoCoordinate id="EFD03A03418BFF5FE2C249AB1ECA0E79" degrees="24" direction="north" minutes="13.929" orientation="latitude" precision="1" value="24.23215">24 ° 13.929 ' N</geoCoordinate>
,
<geoCoordinate id="6D1A5B3C9C2779F50D86E2816661570B" degrees="98" direction="east" minutes="25.563" orientation="longitude" precision="1" value="98.42605">98 ° 25.563 ' E</geoCoordinate>
</named-content>
,
<quantity id="93A0D6F8EDCE2007B1D80EE3C9EA050E" metricMagnitude="3" metricUnit="m" metricValue="1.636" unit="m" value="1636.0">
<elevation id="FC7EF5416D0178A8470B3E3E5CCB7BF5" metricMagnitude="3" metricUnit="m" metricValue="1.636" unit="m" value="1636.0">1636 m</elevation>
</quantity>
),
<collectingMunicipality id="9DC5ACD94CF623957B2AD30B6E5CC229">Luxi</collectingMunicipality>
,
<collectingCounty id="389283AF6D6E30832177C5AFCA7C888B">Dehong</collectingCounty>
,
<emphasis id="C73F9E8CDC3216AFC8BB4A61E5CECD37" bold="true">
<collectingRegion id="9350BF0CDAA080C98FB7D5A6E8EE2CD5" country="China" name="Yunnan">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="5961B75AEF09065ADDE2C85D8FB1D229" bold="true">
<collectingCountry id="262928DEEAAF4F29CF630CD3C4945DF1" name="China">China</collectingCountry>
</emphasis>
,
<date id="A9409D812E71D68A4C03BB75E27A60B3" value="2010-08-24">
<collectingDate id="863536BC3B93924A8B935AC2A6CDBDEF" value="2010-08-24">24 / 08 / 2010</collectingDate>
</date>
,
<collectorName id="1AE45E55BF5BA94E5473F2058FBD86C8">C Wang</collectorName>
,
<collectorName id="AA0BE61016877FB286E1551404911818">Q Zhao</collectorName>
&amp;
<collectorName id="99D3B137A304BB4FB0A7F720B748313F">L Lin</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQZAI" type="etymology">
<paragraph id="D5DC9B951A4F466D48AC66145C69C170">
<heading id="0434ACDACB242D5C66F20D8AE138608F" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="72A42787B76CC9A793A00BF0F45939D8">
The specific name refers to the
<typeStatus id="4C4947C2BE4D3CEB26CEE2503A3EA908">type</typeStatus>
locality, which is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVZAI" type="diagnosis">
<paragraph id="D54B99D4FCCBFAB0D8E9BF443C1967D7">
<heading id="39506746B49409823A220E9D3920E2D8" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="4925317A52087A58906E0CC3AA766BD4">
The new species resembles
<taxonomicName id="04034A1E5898C7B601BD7F245AF5D838" authority="Yang &amp; Yao, 2023" authorityName="Yang &amp; Yao" authorityYear="2023" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yuhaoi">
<emphasis id="A6B48929C134911E335F746CE67782E1" italics="true">B. yuhaoi</emphasis>
<bibRefCitation id="3D339DBB272257EEE3D82E263BB5CCF0" DOI="10.3897/zookeys.1186.105736" author="Yang L &amp; Zhao F &amp; He Q &amp; Yao Z" journalOrPublisher="ZooKeys" pagination="175-184" refId="B4" refString="Yang L, Zhao F, He Q, Yao Z (2023) A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China. ZooKeys 1186: 175 - 184. https: // doi. org / 10.3897 / zookeys. 1186.105736" title="A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China." volume="1186" year="2023">Yang &amp; Yao, 2023</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="09EEC21861F5265AC170D48680D73667" DOI="10.3897/zookeys.1186.105736" author="Yang L &amp; Zhao F &amp; He Q &amp; Yao Z" etAl="et al." firstAuthor="Yang" journalOrPublisher="ZooKeys" page="178" pagination="175-184" refId="B4" refString="Yang L, Zhao F, He Q, Yao Z (2023) A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China. ZooKeys 1186: 175184. https://doi.org/10.3897/zookeys.1186.105736" title="A survey of pholcid spiders (Araneae, Pholcidae) from Guiyang, Guizhou Province, China." volume="1186" year="2023">Yang et al. 2023: 178</bibRefCitation>
, figs 2 A, B, 3 A D, 4 A H) by having similar bulbal apophyses and epigyne (Figs
<figureCitation id="E3AA2C9D1853F977BCF80946D0DDAD71" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 A, C</figureCitation>
,
<figureCitation id="30117B015526A46A3FBF212F473D47AE" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 E</figureCitation>
), but can be distinguished by differences in males: tips of cheliceral distal apophyses closer to each other (Fig.
<figureCitation id="4619027197A489F8FF1A5CCF50895B72" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 D</figureCitation>
vs widely separated), procursus without sclerotized dorso-subdistal apophysis (Figs
<figureCitation id="F34DBE012FD4558ECE1306E6813EA428" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 C</figureCitation>
,
<figureCitation id="C50760A0B9508E247661E88BB443C3A0" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 C</figureCitation>
vs present) and with nearly half-round retrolateral flap (Figs
<figureCitation id="3DAA35516B8D5E1EA607DF688CAEFD22" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 D</figureCitation>
,
<figureCitation id="02C92FD92E1F50D1C192F0024DDDBEF4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 D</figureCitation>
vs angular); differences in females: epigynal pockets closer to each other (Figs
<figureCitation id="4B2936C8102FE7969D18E249EB3236BC" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 A, B</figureCitation>
,
<figureCitation id="502537CC4C3E4DE444FD967727EE6E4A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 E, F</figureCitation>
vs widely separated), pore plates nearly isosceles triangle-shaped (Figs
<figureCitation id="0C2B0699850A63700146C0E2D95CBEE5" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 B</figureCitation>
,
<figureCitation id="CA9DB956BEBFFD6CC9FF2A23C270C958" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 F</figureCitation>
vs scalene triangle-shaped).
</paragraph>
<caption id="CA908DF7F56EC39553EEA842D3652458" ID-DOI="10.3897/zookeys.1202.121633.figure16" ID-arpha="83CEE6D7-A307-5DB9-B938-214E1B4EC0CD" httpUri="https://binary.pensoft.net/fig/1055881" startId="F16">
<paragraph id="6DE35D9C919588E72B0CB8709CBCDEFD">
<label id="F9870673652F00C5326FC111B11875FD">Figure 16.</label>
</paragraph>
<paragraph id="73564ADAAC26D7BAA36F76E2A9D915D3">
<taxonomicName id="3870874DD487E3532A92F76103E86F45" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yunnan" status="sp. nov.">
<emphasis id="D88D7148009150C03FB7B2F42022A14E" italics="true">Belisana yunnan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B9B8CBA1134B00DB55166341447658E" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="0775277490CCDF154056D9729FA67420" bold="true">A, B</emphasis>
palp (
<emphasis id="9658583A529C6B23E3F8BFEC9BC985E9" bold="true">A</emphasis>
prolateral view
<emphasis id="2F11B22FA2D9EF0C7A901D4C9AD3146B" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="7A0823F04D75411BA8640C7181AB5814" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="3D059C6D44B6CED1A51C85B80906B83A" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine
<emphasis id="FEFAA632B688EB0F5EE8DA2AEAA0E36D" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap,
<abbrev id="ABBRID0EV3AI" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="4F0C037E0C783BFCFA4A7D2A3B490FC8" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="9DB658710DFF952606F9A51746A08421" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="E01572CE04D3C4499CEEB375AA81665A" ID-DOI="10.3897/zookeys.1202.121633.figure17" ID-arpha="DBB28810-B821-563A-8A48-1378939A53BE" httpUri="https://binary.pensoft.net/fig/1055887" startId="F17">
<paragraph id="F105292DEA360DABF9CD43395430DA3F">
<label id="DDE208BD8CBD2B9EF91AADCAB6B5CBD9">Figure 17.</label>
</paragraph>
<paragraph id="672CB4B9BD5650684A78F33AA3B9ED6A">
<taxonomicName id="769092C4453520E047A97DCBCA35C4BB" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yunnan" status="sp. nov.">
<emphasis id="8C581D0055732A036E90919AC11D7318" italics="true">Belisana yunnan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="05192AE2CE954B63392FCDB384A38CC2" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="0C91BEF73CAB9EE90A6912331DA0A232" bold="true">C F</emphasis>
) and paratype female (
<emphasis id="B93910777C7E673A35B736A9DE01EE4F" bold="true">A, B, G, H</emphasis>
)
<emphasis id="1E8BCD5B5F04566E737FBDF8D3C72128" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="64584926D23F8BF136F34755CBBD68BE" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="4633C23D0237D6CDF4E1263DF4853F89" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="2ABBA41A7185B5D82653FD0FF8BF23FC" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="79887CBEC3F98690433E309E6D434EF6" bold="true">E H</emphasis>
habitus (
<emphasis id="4AA9ABE2B2374D56E863EEEB18FD6ECC" bold="true">E, G</emphasis>
dorsal view
<emphasis id="26DE6BD104DD1C35922550B3E25794B4" bold="true">F</emphasis>
lateral view
<emphasis id="B7B2A26D262A5A213BFD3F5B92FA3A7A" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EJ5AI" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EN5AI" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ER5AI" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EV5AI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EZ5AI" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E45AI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="F46C168A8777B0BB6A9D5DE361558BBA" bold="true">A D</emphasis>
); 0.30 (
<emphasis id="F992035265DB0FECA1C9AF1FE640EB90" bold="true">E H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EK6AI" type="description">
<paragraph id="7680E89EA5B20157FD3F0042F24AD517">
<heading id="7D191564E31C5BDC5FC4F4F7AF2003E7" reason="title">Description.</heading>
</paragraph>
<paragraph id="1D3AA6BE46704F76AAAFA15522490B38">
<emphasis id="009FD0383BF98FA06171D059B8796553" bold="true">Male</emphasis>
(
<emphasis id="FCD5E2ACAD4CAFD38F6E65B3737ADD51" bold="true" italics="true">
<typeStatus id="EE36BE954ECB930691510EA0A2E17531" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.90 (1.98 with clypeus), carapace 0.79 long, 0.71 wide, opisthosoma 1.11 long, 0.87 wide. Leg I: 18.71 (4.85, 0.56, 4.75, 7.05, 1.50), leg III: 9.35 (2.66, 0.30, 2.25, 3.25, 0.89), legs II and IV missing; tibia I
<abbrev id="ABBRID0EV6AI" xlink_title="length/diameter">L / d</abbrev>
: 59. Eye inter-distances and diameters:
<abbrev id="ABBRID0EZ6AI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E46AI" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0EBABI" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EFABI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EJABI" xlink_title="anterior lateral eye">ALE</abbrev>
0.03,
<abbrev id="ABBRID0ENABI" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.63 / 0.60. Habitus as in Fig.
<figureCitation id="8FD59DC5DC5ECADAB282ADC2C723766A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 E, F</figureCitation>
. Carapace yellowish, with brown radiating marks; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae (Fig.
<figureCitation id="7AB8E4728394A991AAD6908B9CE88EF4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 D</figureCitation>
) with a pair of proximo-lateral apophyses and a pair of curved distal apophyses (distance between tips: 0.19). Palp as in Fig.
<figureCitation id="680BAD14C53948372334F475778ECBB4" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 A, B</figureCitation>
; trochanter with ventral apophysis (as long as wide, arrow
<quantity id="0CAAA926359012D956B9E778C191E145" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="5DC3987D804DEA747460805B816A3414" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 B</figureCitation>
); femur with tiny retrolatero-proximal protrusion (arrow
<quantity id="FF37B2516E2A462589D67D7E208FA75D" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="F8749D6C853F4C74D42772A5CF6CA1BB" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-ventral membranous lamella (arrow
<quantity id="06DD7321FCE66F98C8219CCDAE90F97F" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="D84EDC80DB1BA1F4014330405D78ECA0" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 C</figureCitation>
,
<figureCitation id="17222C001DF01BC59639709C6908411C" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 C</figureCitation>
) bearing median sclerotized part, distal membranous lamella (arrow
<quantity id="45495EE40DFBFC76D30B1B33C863B624" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="9489CDD909BCF4A9331E719C43687B06" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 C</figureCitation>
,
<figureCitation id="B899A520B7D72FA73DC922B6BBCDD812" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 C</figureCitation>
) bearing median sclerotized part, distal spine (arrow
<quantity id="F1C5374A6042AB2CDDFA90A130BC1E86" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="DB6E76EFE6240EA4BAB01ECE9F022A63" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 C</figureCitation>
,
<figureCitation id="ACE5CFDD1ACD65626A2C075F200ABF1F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 C</figureCitation>
), and nearly half-round retrolateral flap (Figs
<figureCitation id="AA1C96DFF13022E31F6909A87F97822C" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Belisana yunnan sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-ventral membranous lamella, arrow 2 points at distal membranous lamella, arrow 3 points at distal spine D retrolateral view). Abbreviations: b = bulb, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure16" httpUri="https://binary.pensoft.net/fig/1055881">16 D</figureCitation>
,
<figureCitation id="A2952FC42EF78CB51315CE0B3B95A5C4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana yongsheng sp. nov. C, D B. yunnan sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure19" httpUri="https://binary.pensoft.net/fig/1055889">19 D</figureCitation>
); bulb (Fig.
<figureCitation id="063D9571612EC317F329DF55D94EC2FD" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 C</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria on tibia I at 14 % proximally; legs with short vertical setae on metatarsi; tarsus I with 21 distinct pseudosegments.
</paragraph>
<caption id="E82F6C96CF6F21F5130C8B07A18094D8" ID-DOI="10.3897/zookeys.1202.121633.figure18" ID-arpha="DC703242-9F3A-584E-985E-43B68E5BF24F" httpUri="https://binary.pensoft.net/fig/1055888" startId="F18">
<paragraph id="1D588C765851EB335BF7A6DE8B02DC6D">
<label id="FE63ED6A17EC5094FDA99D590B009958">Figure 18.</label>
</paragraph>
<paragraph id="A0E2BE70D801F4EBDBE9721F99A663D7">
Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species)
<emphasis id="9183CBE6BCDAE4ED7C3EFE95614DF8C1" bold="true">A, B</emphasis>
<taxonomicName id="89CB0D55FAF3626A1E104763952C8D80" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="honghe" status="sp. nov.">
<emphasis id="A8DE21E35831D265DE52C369288738CB" italics="true">Belisana honghe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="410320F38566EDF6892ED9D6DC0A6AD8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="D34DBAC8004BFEFED4434C47E09DC3B1" bold="true">C, D</emphasis>
<taxonomicName id="460373E0D49B476AB0B97702D7419A6A" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="jiuxiang" status="sp. nov.">
<emphasis id="2B50899CDEA614C27207AA1A4590E7A9" italics="true">B. jiuxiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BC75AA948053C96A301CD5DDF925FE05" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="88C05014C907EE784CAB91B60ACAD6EA" bold="true">E, F</emphasis>
<taxonomicName id="4BA383A1FDEE7D4A5CD545501D032369" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lincang" status="sp. nov.">
<emphasis id="C6C2B5FCA0E8DDAD688122EC1DE595C5" italics="true">B. lincang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1F9F4B62FEA72EECC27D8DC6AFE2DBB1" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F719B79ADAA54D56CBE79929A3C74635" bold="true">G, H</emphasis>
<taxonomicName id="B9464B1BDF7125E3661799E11213F2DD" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="luxi" status="sp. nov.">
<emphasis id="8A890FEBE1351D0D2AAA29A577FD8B74" italics="true">B. luxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="81097C84E08119BC855B7F9067042DE5" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="19C853697CC8F43B7C605BA53026A60F" bold="true">I, J</emphasis>
<taxonomicName id="CE15C7851D42ADAE5E935EA34950097A" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tengchong" status="sp. nov.">
<emphasis id="5B00308282E6304A58E6E4E1246188C6" italics="true">B. tengchong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4FAFCA0AEE69BA3926B91D3696A36F9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="1FD06454F7E02B5271351718BA411E75" bold="true">K, L</emphasis>
<taxonomicName id="E2E8CC25CF54AEC08233E697CEF8391B" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="E4F4D39B295F4A928999DFBE9E873235" italics="true">B. tongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A07540FCB9FC35B06652519DF6CFC56B" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviation: f = flap. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="0BD209D354D9DA5DA4B03D86491095F1" ID-DOI="10.3897/zookeys.1202.121633.figure19" ID-arpha="0D4BD583-2957-582B-B7B9-ADF987450230" httpUri="https://binary.pensoft.net/fig/1055889" startId="F19">
<paragraph id="E2ABD0AFA96DEEEE5EFA04297B714B07">
<label id="666A4110A171A45E1F8E2CDBE75CFE2B">Figure 19.</label>
</paragraph>
<paragraph id="AFA2BA55E49C3E37A07E8064EF654508">
Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species)
<emphasis id="C43E07FECC074E0C204ECB78610F5E36" bold="true">A, B</emphasis>
<taxonomicName id="6A144A7AECCA9DABB2CAF58174AC2722" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yongsheng" status="sp. nov.">
<emphasis id="B9C473E77E25B8E0D6E2222BBD306983" italics="true">Belisana yongsheng</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EE4715909BD5DB1EFD44B5C12CB35776" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="82D9ACCE1CCAE1B801603F6C127FB09E" bold="true">C, D</emphasis>
<taxonomicName id="431C465692EE0A6923E7BD15DE828AA7" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yunnan" status="sp. nov.">
<emphasis id="FD3E9227D1E5B3D01A09FBFCB63C4B70" italics="true">B. yunnan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D72FCF409F1EA49BBDA72C5A3FCEC68A" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviation: f = flap. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="FF79E80BE742BCC2E3FA38F6BBBE115C" ID-DOI="10.3897/zookeys.1202.121633.figure20" ID-arpha="1FBF128D-E9B2-53D0-8062-554497E6500C" httpUri="https://binary.pensoft.net/fig/1055890" startId="F20">
<paragraph id="4F250325381904F5E57BBC3280E72D9B">
<label id="7EA53FA2AF81C23C14ACB6614FDEF84C">Figure 20.</label>
</paragraph>
<paragraph id="405081F739687045AFAF826181F2E26F">
Female genitalia in ventral and dorsal views
<emphasis id="631B7C12C9CB4F502C19FE8680AA6F80" bold="true">A, B</emphasis>
<taxonomicName id="307303B204A31AAC4C62C919D8FDEFD1" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="honghe" status="sp. nov.">
<emphasis id="B4DB734F76220B0118F3AEE662D76BEC" italics="true">Belisana honghe</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11DF07ED473E0A4ECD7D1C536247A6CA" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B36BAA398C27922ACF32ACCB3C4F8161" bold="true">C, D</emphasis>
<taxonomicName id="A6CB4B53B88789249F64558AE2AD9CE4" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="jiuxiang" status="sp. nov.">
<emphasis id="AD10C155B4E6A492A11D9A35845E2277" italics="true">B. jiuxiang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C001F8FA05D1F76089B1FDFC191712E8" rank="species">sp. nov.</taxonomicNameLabel>
, arrow points at lateral sclerite
<emphasis id="D3DA693365A9C538690D46BAD836D968" bold="true">E, F</emphasis>
<taxonomicName id="692B5E7F3479D41A935B0D0EF76738BB" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lincang" status="sp. nov.">
<emphasis id="649B4A5303F0C1FB76788A1648088FD7" italics="true">B. lincang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2FF615303BFA4C65081CE617EF41301C" rank="species">sp. nov.</taxonomicNameLabel>
, arrow points at sac-like structure
<emphasis id="21C1CA56C8F2CEDB2ADB951E756E6474" bold="true">G, H</emphasis>
<taxonomicName id="B9EFEDBBC1C7474C27E5A1A5F5893311" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="luxi" status="sp. nov.">
<emphasis id="B84D6092D253FD82F0E6D26C1CD6B368" italics="true">B. luxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0A39F28FF767D4CB503321DD5FF51054" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations:
<abbrev id="ABBRID0EHIBI" xlink_title="anterior arch">aa</abbrev>
= anterior arch,
<abbrev id="ABBRID0ELIBI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EPIBI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10.
</paragraph>
</caption>
<paragraph id="3E0D29846499E2DA2523257632C4ED93">
<emphasis id="4F6D5B9211853DDA221E8E1470E92243" bold="true">Female</emphasis>
(
<emphasis id="B44BEBD558BA9B6733DC957D4B220F22" bold="true" italics="true">
<typeStatus id="F67895679FD730D74D24C4B9B6CCBD64" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="8BD6473486F784BDA026EB7A5A6E9CDD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="92C8AEC52EE5F76F5868A7F4BF7A08F1" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="1FFAA7929909CFA1F464BCB208AB5727">Ar 44984</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="C8A78B237438DCCE5CFA74608CFD5B0A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 G, H</figureCitation>
. Total length 2.14 (2.20 with clypeus), carapace 0.78 long, 0.77 wide, opisthosoma 1.36 long, 1.00 wide; tibia I: 3.88; tibia I
<abbrev id="ABBRID0EHJBI" xlink_title="length/diameter">L / d</abbrev>
: 52. Eye inter-distances and diameters:
<abbrev id="ABBRID0ELJBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0EPJBI" xlink_title="posterior median eye">PME</abbrev>
0.13,
<abbrev id="ABBRID0ETJBI" xlink_title="posterior median eye">PME</abbrev>
0.07,
<abbrev id="ABBRID0EXJBI" xlink_title="posterior median eye">PME</abbrev>
<abbrev id="ABBRID0E2JBI" xlink_title="anterior lateral eye">ALE</abbrev>
0.02. Sternum width / length: 0.62 / 0.56. Epigyne (Figs
<figureCitation id="E20B09F9885BED7274F6C2FA60DFE5E5" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 A</figureCitation>
,
<figureCitation id="FDCA3587EBF1DEDF250CD95F37FBE906" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 E</figureCitation>
) simple and flat, anteriorly slightly sclerotized, with a pair of postero-median pockets 0.12 apart. Vulva (Figs
<figureCitation id="B98D2BC114E29ECA0289ED58DE172CF0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Belisana yunnan sp. nov., holotype male (C F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A D); 0.30 (E H)." figureDoi="10.3897/zookeys.1202.121633.figure17" httpUri="https://binary.pensoft.net/fig/1055887">17 B</figureCitation>
,
<figureCitation id="0E482AB294FEC12CDF1F92B7109B77D5" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Female genitalia in ventral and dorsal views A, B Belisana tengchong sp. nov. C, D B. tongi sp. nov., arrows point at four distinct median teeth E, F B. yunnan sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure21" httpUri="https://binary.pensoft.net/fig/1055891">21 F</figureCitation>
) with ridge-shaped anterior arch and a pair of nearly triangular pore plates.
</paragraph>
<caption id="A81F1F5DC7DB5F9313B2BB61636004D1" ID-DOI="10.3897/zookeys.1202.121633.figure21" ID-arpha="DA4496D5-3EFE-5CB0-887E-952B9C84AD58" httpUri="https://binary.pensoft.net/fig/1055891" startId="F21">
<paragraph id="87CC75F4103AD68C4B2DD86912042A37">
<label id="DC67551BDA982330CF8BDF3DC96205DD">Figure 21.</label>
</paragraph>
<paragraph id="9EC2A65BD14F27758E8634C8FC48F255">
Female genitalia in ventral and dorsal views
<emphasis id="A60C82E356E1974F7D7438AF922E75D0" bold="true">A, B</emphasis>
<taxonomicName id="9C428EFC48011C21FDC2E0A95CEE365E" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tengchong" status="sp. nov.">
<emphasis id="9A10CAC4725CD472E263EDD6881D4882" italics="true">Belisana tengchong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A805A30E23B7892EBF6924C121BA0DC9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="3CC9AAD5612F96425B65E69F1B1E1318" bold="true">C, D</emphasis>
<taxonomicName id="FB2501345E460C47A4F2097C575FE39C" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongi" status="sp. nov.">
<emphasis id="FFCE3FA7D443B1F4AC4EF4A62C2C7ECF" italics="true">B. tongi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AD8B2E7F32033CD8A8936CF135A1DE60" rank="species">sp. nov.</taxonomicNameLabel>
, arrows point at four distinct median teeth
<emphasis id="87C86D7505F99A44CA7B8922EB06F097" bold="true">E, F</emphasis>
<taxonomicName id="E70115ABC119097F68FA0FFF7D4FB595" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="yunnan" status="sp. nov.">
<emphasis id="943A2E303C7AF3D4F0964880DCD68C8F" italics="true">B. yunnan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="421D106237E97DAD9F3C4F5FB1C9F2C2" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations:
<abbrev id="ABBRID0E5LBI" xlink_title="anterior arch">aa</abbrev>
= anterior arch,
<abbrev id="ABBRID0ECMBI" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EGMBI" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EPMBI" type="variation">
<paragraph id="3F80C98DA9650298BDF5FCACBBA8EC4A">
<heading id="8010A0C2054FD364CF11870774C58500" reason="title">Variation.</heading>
</paragraph>
<paragraph id="4E99542501E6C363D2493CE5DBCC46D0">
Tibia I in
<specimenCount id="268E869B8F079CAE87F0E48434E4ACA2" count="2" type="male">two male</specimenCount>
<typeStatus id="31857AB7C5D01F137E602AAA633EAF04" type="paratype">paratypes</typeStatus>
(
<named-content id="1688358A16CD7033237D8B912BAF63BA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="E22F5456878617C9DF96B3284A64FCFF" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="BE31B6606229B665DDE1EEBD2A9AD579">
<specimenCode id="760F2348298A0555F31D150C6724FD05">Ar 44982</specimenCode>
83
</specimenCode>
): 5.38, 6.73. Tibia I in the other
<specimenCount id="B1576F8E209B544A920F97CAFFEE062D" count="2" type="female">two female</specimenCount>
<typeStatus id="0403499BF225C61910DB6FCA15628941" type="paratype">paratypes</typeStatus>
(
<named-content id="FC94A12309E1064E4C5C18739E5A48E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="C6A65B9B006119081F0B10BA199766F6" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="C1D6540CD8DC749FAA0B765053E78BAF">
<specimenCode id="D584C7F86084DDE3ED6916A09E26E60E">Ar 44985</specimenCode>
86
</specimenCode>
): 3.72, 3.85.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4MBI" type="habitat">
<paragraph id="AFB55D396D0125F0296737D17ED870B3">
<heading id="E63425F47B28B2FDA64E67B4C75459A5" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="76C0EC06741F288BE7D6B9B2BF34B60E">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECNBI" type="distribution">
<paragraph id="D711F582C052BFF1C8BD5D2C0FC9BE44">
<heading id="B7B773EB4780D2E97069927640620A26" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="331C02D4B7EAFCACAED95E6307459CCA">
<collectingCountry id="189F7BD535599B65131365B7F42EF79F" name="China">China</collectingCountry>
(
<collectingRegion id="53E2B5409D49DC3AF7BE46F9427A7B93" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<typeStatus id="230F390C6D747EF2C7D95C3337C615CA">type</typeStatus>
locality; Fig.
<figureCitation id="6E670D698408033E985048AB07A26836" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the new Belisana species from Yunnan, China 1 Belisana honghe sp. nov. 2 B. jiuxiang sp. nov. 3 B. lincang sp. nov. 4 B. luxi sp. nov. 5 B. tengchong sp. nov. 6 B. tongi sp. nov. 7 B. yongsheng sp. nov. 8 B. yunnan sp. nov." figureDoi="10.3897/zookeys.1202.121633.figure1" httpUri="https://binary.pensoft.net/fig/1055866">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>