treatments-xml/data/7D/2E/87/7D2E87A4D139FF9DFC9EFABDFEDC41B7.xml

468 lines
60 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E3FA162C320365C8656D48B272068377" ID-DOI="10.5281/zenodo.13345878" ID-ISSN="1732-2421" ID-Zenodo-Dep="13345878" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723061107117" checkinUser="felipe" docAuthor="Machalski, Marcin, Jagt, John W. M., Dortangs, Rudi W., Mulder, Eric W. A. &amp; Radwański, Andrzej" docDate="2003" docId="7D2E87A4D139FF9DFC9EFABDFEDC41B7" docLanguage="en" docName="ActaPalaeontolPol.48.3.397-408.pdf" docOrigin="Acta Palaeontologica Polonica 48 (3)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Mosasaurus hoffmanni Mantell 1829" docType="treatment" docVersion="3" lastPageNumber="403" masterDocId="8117FFDCD138FF9BFFB2FFF9FFBC4049" masterDocTitle="Campanian and Maastrichtian mosasaurid reptiles from central Poland" masterLastPageNumber="408" masterPageNumber="397" pageNumber="398" updateTime="1724110895222" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D38C34A67DBFE36510087E5DE4DF797E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2DD33800235D98500707F3213BFE1D03">
<mods:title id="B1029F256D88233A16BD2975CC5CB7CD">Campanian and Maastrichtian mosasaurid reptiles from central Poland</mods:title>
</mods:titleInfo>
<mods:name id="F86F6D20FF3AD9067432512E6637E9A4" type="personal">
<mods:role id="E1C8DF231C8BA17C0BFE311BC3116186">
<mods:roleTerm id="C8E2DFABE88B99BE3F28CED77006F380">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F0F0DD1234C139AC01ED284F1D9B9F2">Machalski, Marcin</mods:namePart>
</mods:name>
<mods:name id="4F2F470559BB9217B222EF576A634E27" type="personal">
<mods:role id="53F760D666F854CB65B00D189938E988">
<mods:roleTerm id="6210276E9D38DCE80CB58A10ABB06DDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C09C4EBEA6A980189ACAFFAF2B2E860">Jagt, John W. M.</mods:namePart>
</mods:name>
<mods:name id="D2536CD0D33085FE0CEF61F43EE0207F" type="personal">
<mods:role id="25AA861B2CD840B68BCC1C8A560728B9">
<mods:roleTerm id="FEB095C7156F452F3D3A9C57AF01B4E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C198DAC7C71DBF82302FE2D263502E92">Dortangs, Rudi W.</mods:namePart>
</mods:name>
<mods:name id="5127E3CE8A5A63C3C001DC3F10341DDF" type="personal">
<mods:role id="286A665138AF0F9323449CED33048157">
<mods:roleTerm id="0EE345B378170929921D072166FD1DF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F1408831532B1EF88F6AE62CCDEA4A6">Mulder, Eric W. A.</mods:namePart>
</mods:name>
<mods:name id="239EF12EA8D46C0D7D04F813CEFCC865" type="personal">
<mods:role id="E36C8EA503B34388F01CCC6F66A80522">
<mods:roleTerm id="D18BE0B8B131A4B531B7A675020EE4F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9B91B4C40880B33432D744A2BEC065C">Radwański, Andrzej</mods:namePart>
</mods:name>
<mods:typeOfResource id="6EC4AA8FE9D51DBB457E850F20C90354">text</mods:typeOfResource>
<mods:relatedItem id="344059D992CE1C5F8F3CB4578F849ED5" type="host">
<mods:titleInfo id="993965EE2D1BE01609601E99DD664AE2">
<mods:title id="3DD9D2AEE694EBAFBA2EED5468132F0A">Acta Palaeontologica Polonica</mods:title>
</mods:titleInfo>
<mods:part id="8A2C9383A9F1377C756134C76A05544D">
<mods:date id="3086F514FCA0726181437072586180A1">2003</mods:date>
<mods:detail id="72233E075CD06C10A085E6B9CB60998D" type="pubDate">
<mods:number id="810EABDAE0E33D8E58D41F6844971A9A">2003-09-30</mods:number>
</mods:detail>
<mods:detail id="066AC81FFFDD8E6D7E98A3EB70D5498D" type="volume">
<mods:number id="C89E91A618830636065C737AD56457A6">48</mods:number>
</mods:detail>
<mods:detail id="A9131C5D832E92FFDCB7F09232E18C65" type="issue">
<mods:number id="B56C19F4B89AAF270DC990704A04EB83">3</mods:number>
</mods:detail>
<mods:extent id="D47D34845E3039B1D59F407018F88523" unit="page">
<mods:start id="AB5D43D5D68AC5B74B0948841A2CC471">397</mods:start>
<mods:end id="B35B2CF94B367EF236C06D1D87BD9E2A">408</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1DB2AB5702FCE24F50DF364428D775C1">journal article</mods:classification>
<mods:identifier id="0E4459673E25930A8F128CBDB8ACED2A" type="DOI">10.5281/zenodo.13345878</mods:identifier>
<mods:identifier id="EAB90EA7C08485CD65FC516688F9986F" type="ISSN">1732-2421</mods:identifier>
<mods:identifier id="F7C50F9EA74C94C6E0B32E379C17BBA3" type="Zenodo-Dep">13345878</mods:identifier>
</mods:mods>
<treatment id="7D2E87A4D139FF9DFC9EFABDFEDC41B7" LSID="urn:lsid:plazi:treatment:7D2E87A4D139FF9DFC9EFABDFEDC41B7" httpUri="http://treatment.plazi.org/id/7D2E87A4D139FF9DFC9EFABDFEDC41B7" lastPageId="6" lastPageNumber="403" pageId="1" pageNumber="398">
<subSubSection id="BD9D6539D139FF9AFC9EFABDFA8A4529" box="[812,1334,1347,1376]" pageId="1" pageNumber="398" type="nomenclature">
<paragraph id="F53836B2D139FF9AFC9EFABDFA8A4529" blockId="1.[812,1334,1347,1409]" box="[812,1334,1347,1376]" pageId="1" pageNumber="398">
<heading id="AE7081DED139FF9AFC9EFABDFA8A4529" box="[812,1334,1347,1376]" fontSize="12" level="2" pageId="1" pageNumber="398" reason="9">
<taxonomicName id="32874D31D139FF9AFC9EFABDFA8A4529" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[812,1334,1347,1376]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="398" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D139FF9AFC9EFABDFC744529" box="[812,968,1348,1376]" italics="true" pageId="1" pageNumber="398">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D139FF9AFC4BFABAFBC74529" box="[1017,1147,1347,1376]" italics="true" pageId="1" pageNumber="398">hoffmanni</emphasis>
<bibRefCitation id="91164B43D139FF9AFB31FABAFA8A4529" author="Mantell, G. A." box="[1155,1334,1347,1376]" pageId="1" pageNumber="398" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell, 1829</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BD9D6539D139FF9DFC9EFA92FEDC41B7" lastPageId="6" lastPageNumber="403" pageId="1" pageNumber="398" type="description">
<paragraph id="F53836B2D139FF9AFC9EFA92FC3045C8" blockId="1.[812,1334,1347,1409]" box="[812,908,1387,1409]" pageId="1" pageNumber="398">
<figureCitation id="6DBC2A37D139FF9AFC9EFA92FC3445C8" box="[812,904,1387,1409]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[102,140,1722,1741]" captionStartId-1="3.[102,141,1591,1610]" captionStartId-2="4.[102,139,1355,1374]" captionStartId-3="5.[966,1004,970,989]" captionTargetBox-0="[105,1481,225,1704]" captionTargetBox-1="[104,1482,225,1564]" captionTargetBox-2="[103,1484,225,1327]" captionTargetBox-3="[104,932,799,1839]" captionTargetId-0="figure-88@2.[105,1482,225,1704]" captionTargetId-1="figure-138@3.[104,1483,225,1564]" captionTargetId-2="figure-298@4.[103,1486,225,1327]" captionTargetId-3="figure-221@5.[104,932,799,1839]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Fig.2. Mosasaurus cf. hoffmanni Mantell,1829,fromtheDaniangreensandatNasiłów(redepositedlateLateMaastrichtianmaterial). A.IGPUWAR1in buccal(A1),lingual(A2),andanterior(A3)views. B.IGPUWAR2inposterior(B 1)andlingual(B2)views. C.IGPUWAR3inposterior(C 1),buccal(C2), and anterior (C3) views." captionText-1="Fig.3. Mosasaurus cf. hoffmanni Mantell,1829,fromtheupperUpperMaastrichtianopokaatNasiłów. A.IGPUWAR4inposterior(A 1),buccal(A2),anterior (A3), and lingual (A4) views. B. OGP 1254 in lingual (B1), posterior (B2), buccal (B3), and anterior (B4) views." captionText-2="Fig.4. Mosasaurus cf. hoffmanni Mantell,1829,fromtheUpperCampanianopokaatMaruszów,ZPALR.I/1,partialupperjawinbuccal(A),occlusal(B), and lingual (C) views." captionText-3="Fig.6. Mosasaurus cf. hoffmanni Mantell,1829,fromthe Upper Campanian opoka at Maruszów. A. ZPAL R.I/1, lefthandsidetoothinBin anterior(A1),buccal(A2),and posterior (A3) views. B. ZPAL R.I/1, three tooth crowns preserved in original jaw position (although jaw bone is not preserved)." figureDoi-0="http://doi.org/10.5281/zenodo.13345882" figureDoi-1="http://doi.org/10.5281/zenodo.13345884" figureDoi-2="http://doi.org/10.5281/zenodo.13345886" figureDoi-3="http://doi.org/10.5281/zenodo.13345888" httpUri-0="https://zenodo.org/record/13345882/files/figure.png" httpUri-1="https://zenodo.org/record/13345884/files/figure.png" httpUri-2="https://zenodo.org/record/13345886/files/figure.png" httpUri-3="https://zenodo.org/record/13345888/files/figure.png" pageId="1" pageNumber="398">Figs. 26</figureCitation>
.
</paragraph>
<paragraph id="F53836B2D139FF9AFC9EFA6AFA99476C" blockId="1.[812,1486,1427,2026]" pageId="1" pageNumber="398">
<emphasis id="C7F3EAA0D139FF9AFC9EFA6AFC3045E2" box="[812,908,1427,1451]" italics="true" pageId="1" pageNumber="398">Material</emphasis>
.—Twoteeth(IGPUWAR1,IGPUWAR2),found togetherandpossiblybelongingtoasingleindividual,andone tooth crown (IGPUWAR3), from the Danian greensand at Nasiłów (redeposited late Late Maastrichtian material). Two toothcrowns(IGPUWAR4,OGP1254)fromtheupperUp per Maastrichtian opoka at Nasiłów. A fragmentary jaw and isolated tooth crowns (ZPAL R. I/15), representing a single individual, from the Upper Campanian of Maruszów, previously recorded as
<taxonomicName id="32874D31D139FF9AFC46F969FB1D46EE" box="[1012,1185,1680,1703]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="398" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C7F3EAA0D139FF9AFC46F969FBC746EE" box="[1012,1147,1680,1703]" italics="true" pageId="1" pageNumber="398">Mosasaurus</emphasis>
sp.
</taxonomicName>
(sp. A) by
<bibRefCitation id="91164B43D139FF9AFAAFF976FCE7468F" author="Sulimski, A." pageId="1" pageNumber="398" pagination="243 - 250" refId="ref7863" refString="Sulimski, A. 1968. Remains of Upper Cretaceous Mosasauridae (Reptilia) of central Poland. Acta Palaeontologica Polonica 13: 243 - 250." type="journal article" year="1968">Sulimski (1968: 245</bibRefCitation>
,pl.1;pl.2:14,refiguredhereinasFigs.46).Comparative material from the
<typeStatus id="2A3C8810D139FF9AFB9EF937FBE746AF" box="[1068,1115,1742,1766]" pageId="1" pageNumber="398">type</typeStatus>
Maastrichtian area (
<typeStatus id="2A3C8810D139FF9AFAF6F937FAC946AF" box="[1348,1397,1742,1766]" pageId="1" pageNumber="398">type</typeStatus>
area of
<taxonomicName id="32874D31D139FF9AFC9EF917FB95474C" authorityName="Mantell" authorityYear="1829" box="[812,1065,1773,1797]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="398" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D139FF9AFC9EF917FB95474C" box="[812,1065,1773,1797]" italics="true" pageId="1" pageNumber="398">Mosasaurus hoffmanni</emphasis>
</taxonomicName>
) includes NHMM 0049842, 007129, 0066843b, 1997262, 1997266, and 1997267.
</paragraph>
<paragraph id="F53836B2D139FF99FC9EF8CCFDF647A3" blockId="1.[812,1486,1427,2026]" lastBlockId="2.[102,774,1843,2026]" lastPageId="2" lastPageNumber="399" pageId="1" pageNumber="398">
<emphasis id="C7F3EAA0D139FF9AFC9EF8CCFC0C4705" box="[812,944,1845,1868]" italics="true" pageId="1" pageNumber="398">Description</emphasis>
.—The bestpreserved specimen is an anterior (?premaxillary)tooth,IGPUWAR2(
<figureCitation id="6DBC2A37D139FF9AFB56F8ADFA8C4725" box="[1252,1328,1876,1900]" captionStart="Fig" captionStartId="2.[102,140,1722,1741]" captionTargetBox="[105,1481,225,1704]" captionTargetId="figure-88@2.[105,1482,225,1704]" captionTargetPageId="2" captionText="Fig.2. Mosasaurus cf. hoffmanni Mantell,1829,fromtheDaniangreensandatNasiłów(redepositedlateLateMaastrichtianmaterial). A.IGPUWAR1in buccal(A1),lingual(A2),andanterior(A3)views. B.IGPUWAR2inposterior(B 1)andlingual(B2)views. C.IGPUWAR3inposterior(C 1),buccal(C2), and anterior (C3) views." figureDoi="http://doi.org/10.5281/zenodo.13345882" httpUri="https://zenodo.org/record/13345882/files/figure.png" pageId="1" pageNumber="398">Fig.2B</figureCitation>
),100.3mmin height (including root, with small resorption pit). It is asymmetrically bicarinate, with a deeply Ushaped cross section, and with markedly unequal lingual and buccal surfaces. The buccal surface ismore orlessflat,with23 facets (bestvisibleproximally);onthelingualsurfacethefacetsarelesswell developed,andtheirnumbercannotbedeterminedprecisely. Both the anterior and posterior carinae are well developed, with minute serrations over their entire length. The crown has a marked posterior and lingual recurvature. The enamel shows beading, best developed proximally.
</paragraph>
<caption id="A1F8663AD13AFF99FFD4F943FEFC474C" ID-DOI="http://doi.org/10.5281/zenodo.13345882" ID-Zenodo-Dep="13345882" httpUri="https://zenodo.org/record/13345882/files/figure.png" pageId="2" pageNumber="399" startId="2.[102,140,1722,1741]" targetBox="[105,1481,225,1704]" targetPageId="2" targetType="figure">
<paragraph id="F53836B2D13AFF99FFD4F943FEFC474C" blockId="2.[102,1485,1722,1801]" pageId="2" pageNumber="399">
Fig.2.
<taxonomicName id="32874D31D13AFF99FF16F942FDAC4684" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[164,528,1722,1741]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="399" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13AFF99FF16F942FEAF4684" box="[164,275,1723,1741]" italics="true" pageId="2" pageNumber="399">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D13AFF99FE81F943FE334684" box="[307,399,1722,1741]" italics="true" pageId="2" pageNumber="399">hoffmanni</emphasis>
<bibRefCitation id="91164B43D13AFF99FE21F943FDAC4684" author="Mantell, G. A." box="[403,528,1722,1741]" pageId="2" pageNumber="399" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell,1829</bibRefCitation>
</taxonomicName>
,fromtheDaniangreensandatNasiłów(redepositedlateLateMaastrichtianmaterial).
<emphasis id="C7F3EAA0D13AFF99FAA9F943FA974684" bold="true" box="[1307,1323,1722,1741]" pageId="2" pageNumber="399">A</emphasis>
.IGPUWAR1in buccal(A
<subScript id="690334F7D13AFF99FF0FF926FF7946A4" attach="left" box="[189,197,1759,1773]" fontSize="6" pageId="2" pageNumber="399">1</subScript>
),lingual(A
<subScript id="690334F7D13AFF99FE83F926FE8546A4" attach="left" box="[305,313,1759,1773]" fontSize="6" pageId="2" pageNumber="399">2</subScript>
),andanterior(A
<subScript id="690334F7D13AFF99FE60F926FE6646A4" attach="left" box="[466,474,1759,1773]" fontSize="6" pageId="2" pageNumber="399">3</subScript>
)views.
<emphasis id="C7F3EAA0D13AFF99FD96F92EFD8F46A3" bold="true" box="[548,563,1751,1770]" pageId="2" pageNumber="399">B</emphasis>
.IGPUWAR2inposterior(B
<subScript id="690334F7D13AFF99FCF1F926FCF746A4" attach="right" box="[835,843,1759,1773]" fontSize="6" pageId="2" pageNumber="399">1</subScript>
)andlingual(B
<subScript id="690334F7D13AFF99FC67F926FC6146A4" attach="left" box="[981,989,1759,1773]" fontSize="6" pageId="2" pageNumber="399">2</subScript>
)views.
<emphasis id="C7F3EAA0D13AFF99FB9AF92FFB8446A0" bold="true" box="[1064,1080,1750,1769]" pageId="2" pageNumber="399">C</emphasis>
.IGPUWAR3inposterior(C
<subScript id="690334F7D13AFF99FAFAF926FAEC46A4" attach="right" box="[1352,1360,1759,1773]" fontSize="6" pageId="2" pageNumber="399">1</subScript>
),buccal(C
<subScript id="690334F7D13AFF99FA0AF926FA7C46A4" attach="right" box="[1464,1472,1759,1773]" fontSize="6" pageId="2" pageNumber="399">2</subScript>
), and anterior (C
<subScript id="690334F7D13AFF99FF5DF902FF4B4740" attach="left" box="[239,247,1787,1801]" fontSize="6" pageId="2" pageNumber="399">3</subScript>
) views.
</paragraph>
</caption>
<paragraph id="F53836B2D13AFF99FCE0F8CDFA7C47A3" blockId="2.[812,1485,1843,2026]" pageId="2" pageNumber="399">
IGPUWAR1 (
<figureCitation id="6DBC2A37D13AFF99FC4FF8CAFBEB4705" box="[1021,1111,1843,1868]" captionStart="Fig" captionStartId="2.[102,140,1722,1741]" captionTargetBox="[105,1481,225,1704]" captionTargetId="figure-88@2.[105,1482,225,1704]" captionTargetPageId="2" captionText="Fig.2. Mosasaurus cf. hoffmanni Mantell,1829,fromtheDaniangreensandatNasiłów(redepositedlateLateMaastrichtianmaterial). A.IGPUWAR1in buccal(A1),lingual(A2),andanterior(A3)views. B.IGPUWAR2inposterior(B 1)andlingual(B2)views. C.IGPUWAR3inposterior(C 1),buccal(C2), and anterior (C3) views." figureDoi="http://doi.org/10.5281/zenodo.13345882" httpUri="https://zenodo.org/record/13345882/files/figure.png" pageId="2" pageNumber="399">Fig. 2A</figureCitation>
) is a massive lateral tooth,
<quantity id="327F9B57D13AFF99FA21F8CAFCE44722" metricMagnitude="-2" metricUnit="m" metricValue="8.959999999999999" pageId="2" pageNumber="399" unit="mm" value="89.6">89.6 mm</quantity>
in total height (as preserved), including the incomplete root. It has an elliptical cross section, with both buccal and lingual surfaces showing barely visible facets and beading. Both posterior and anterior carinae show minute serrations. Thecrownismoderatelyposteriorlyandlinguallyrecurved.
</paragraph>
<caption id="A1F8663AD13BFF98FFD4F9CEFB36462F" ID-DOI="http://doi.org/10.5281/zenodo.13345884" ID-Zenodo-Dep="13345884" httpUri="https://zenodo.org/record/13345884/files/figure.png" pageId="3" pageNumber="400" startId="3.[102,141,1591,1610]" targetBox="[104,1482,225,1564]" targetPageId="3" targetType="figure">
<paragraph id="F53836B2D13BFF98FFD4F9CEFB36462F" blockId="3.[102,1485,1591,1642]" pageId="3" pageNumber="400">
Fig.3.
<taxonomicName id="32874D31D13BFF98FF11F9C1FDB24603" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[163,526,1591,1610]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="400" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13BFF98FF11F9C1FEAE4603" box="[163,274,1592,1610]" italics="true" pageId="3" pageNumber="400">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D13BFF98FE83F9CEFE314603" box="[305,397,1591,1610]" italics="true" pageId="3" pageNumber="400">hoffmanni</emphasis>
<bibRefCitation id="91164B43D13BFF98FE20F9CEFDB24603" author="Mantell, G. A." box="[402,526,1591,1610]" pageId="3" pageNumber="400" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell,1829</bibRefCitation>
</taxonomicName>
,fromtheupperUpperMaastrichtianopokaatNasiłów.
<emphasis id="C7F3EAA0D13BFF98FBB6F9CEFBA84603" bold="true" box="[1028,1044,1591,1610]" pageId="3" pageNumber="400">A</emphasis>
.IGPUWAR4inposterior(A
<subScript id="690334F7D13BFF98FA94F9B9FA924607" attach="right" box="[1318,1326,1600,1614]" fontSize="6" pageId="3" pageNumber="400">1</subScript>
),buccal(A
<emphasis id="C7F3EAA0D13BFF98FA25F9B9FA234607" bold="true" box="[1431,1439,1600,1614]" pageId="3" pageNumber="400">
<subScript id="690334F7D13BFF98FA25F9B9FA234607" attach="left" box="[1431,1439,1600,1614]" fontSize="6" pageId="3" pageNumber="400">2</subScript>
</emphasis>
),anterior (A
<subScript id="690334F7D13BFF98FF06F9A5FF004623" attach="left" box="[180,188,1628,1642]" fontSize="6" pageId="3" pageNumber="400">3</subScript>
), and lingual (A
<subScript id="690334F7D13BFF98FEE0F9A5FEE64623" attach="right" box="[338,346,1628,1642]" fontSize="6" pageId="3" pageNumber="400">4</subScript>
) views.
<emphasis id="C7F3EAA0D13BFF98FE1AF9ADFE0B462E" bold="true" box="[424,439,1620,1639]" pageId="3" pageNumber="400">B</emphasis>
. OGP 1254 in lingual (B
<subScript id="690334F7D13BFF98FD29F9A5FD1F4623" attach="left" box="[667,675,1628,1642]" fontSize="6" pageId="3" pageNumber="400">1</subScript>
), posterior (B
<subScript id="690334F7D13BFF98FC91F9A5FC974623" attach="left" box="[803,811,1628,1642]" fontSize="6" pageId="3" pageNumber="400">2</subScript>
), buccal (B
<subScript id="690334F7D13BFF98FC27F9A5FC214623" attach="left" box="[917,925,1628,1642]" fontSize="6" pageId="3" pageNumber="400">3</subScript>
), and anterior (B
<subScript id="690334F7D13BFF98FB8BF9A5FBFD4623" attach="left" box="[1081,1089,1628,1642]" fontSize="6" pageId="3" pageNumber="400">4</subScript>
) views.
</paragraph>
</caption>
<paragraph id="F53836B2D13BFF98FF39F96EFED94722" blockId="3.[102,775,1686,2026]" pageId="3" pageNumber="400">
IGPUWAR3 (as preserved 57.0 mm in height; see
<figureCitation id="6DBC2A37D13BFF98FD61F96EFF3B4687" captionStart="Fig" captionStartId="2.[102,140,1722,1741]" captionTargetBox="[105,1481,225,1704]" captionTargetId="figure-88@2.[105,1482,225,1704]" captionTargetPageId="2" captionText="Fig.2. Mosasaurus cf. hoffmanni Mantell,1829,fromtheDaniangreensandatNasiłów(redepositedlateLateMaastrichtianmaterial). A.IGPUWAR1in buccal(A1),lingual(A2),andanterior(A3)views. B.IGPUWAR2inposterior(B 1)andlingual(B2)views. C.IGPUWAR3inposterior(C 1),buccal(C2), and anterior (C3) views." figureDoi="http://doi.org/10.5281/zenodo.13345882" httpUri="https://zenodo.org/record/13345882/files/figure.png" pageId="3" pageNumber="400">Fig. 2C</figureCitation>
) and IGPUWAR4 (as preserved 51.6 mminheight; see
<figureCitation id="6DBC2A37D13BFF98FFD4F92CFF0346A7" box="[102,191,1749,1774]" captionStart="Fig" captionStartId="3.[102,141,1591,1610]" captionTargetBox="[104,1482,225,1564]" captionTargetId="figure-138@3.[104,1483,225,1564]" captionTargetPageId="3" captionText="Fig.3. Mosasaurus cf. hoffmanni Mantell,1829,fromtheupperUpperMaastrichtianopokaatNasiłów. A.IGPUWAR4inposterior(A 1),buccal(A2),anterior (A3), and lingual (A4) views. B. OGP 1254 in lingual (B1), posterior (B2), buccal (B3), and anterior (B4) views." figureDoi="http://doi.org/10.5281/zenodo.13345884" httpUri="https://zenodo.org/record/13345884/files/figure.png" pageId="3" pageNumber="400">Fig. 3A</figureCitation>
) show elliptical cross sections, and appear to have occupied a comparable position in the dental or maxillary ramus.Thesespecimenshavemoreclearlydevelopedfacets, on both the buccal and lingual surfaces, and show distinct beading of the enamel.
</paragraph>
<paragraph id="F53836B2D13BFF98FF39F88AFC5B47C2" blockId="3.[102,775,1686,2026]" lastBlockId="3.[812,1486,1687,2026]" pageId="3" pageNumber="400">
OGP 1254 (
<figureCitation id="6DBC2A37D13BFF98FEA8F88AFEC847C2" box="[282,372,1907,1931]" captionStart="Fig" captionStartId="3.[102,141,1591,1610]" captionTargetBox="[104,1482,225,1564]" captionTargetId="figure-138@3.[104,1483,225,1564]" captionTargetPageId="3" captionText="Fig.3. Mosasaurus cf. hoffmanni Mantell,1829,fromtheupperUpperMaastrichtianopokaatNasiłów. A.IGPUWAR4inposterior(A 1),buccal(A2),anterior (A3), and lingual (A4) views. B. OGP 1254 in lingual (B1), posterior (B2), buccal (B3), and anterior (B4) views." figureDoi="http://doi.org/10.5281/zenodo.13345884" httpUri="https://zenodo.org/record/13345884/files/figure.png" pageId="3" pageNumber="400">Fig. 3B</figureCitation>
) is a comparatively small (from a subadult individual?), bicarinate tooth crown (
<quantity id="327F9B57D13BFF98FDCFF86AFD5B47E3" box="[637,743,1939,1963]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="3" pageNumber="400" unit="mm" value="31.5">31.5 mm</quantity>
in height,aspreserved),14.2mminwidth(atthebase).Incross sectionitismoreorlesselliptical,withthelingualandbuccal surfacesmarkedlyunequal.Thebuccalsurfaceisfaintlyconvex,with3unequalfacets;onthelingualsurfacethereareup to9suchfacets,narrowerandespeciallymarkedproximally, reaching to just above midheight; beading is well visible. Bothcarinaearewelldeveloped,theanterioronebeingmore pronounced, withminuteserrationsovertheirentirelengths. The crown is moderately lingually, but more clearly posteriorly recurved.
</paragraph>
<paragraph id="F53836B2D13BFF9FFCE0F86AFDD1462A" blockId="3.[812,1486,1687,2026]" lastBlockId="4.[102,775,1454,1635]" lastPageId="4" lastPageNumber="401" pageId="3" pageNumber="400">
Teeth and tooth crowns in the Upper Campanian specimen ZPAL R. I/15 (
<figureCitation id="6DBC2A37D13BFF98FBAEF84BFB3B4783" box="[1052,1159,1970,1994]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[102,139,1355,1374]" captionStartId-1="5.[966,1004,970,989]" captionTargetBox-0="[103,1484,225,1327]" captionTargetBox-1="[104,932,799,1839]" captionTargetId-0="figure-298@4.[103,1486,225,1327]" captionTargetId-1="figure-221@5.[104,932,799,1839]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig.4. Mosasaurus cf. hoffmanni Mantell,1829,fromtheUpperCampanianopokaatMaruszów,ZPALR.I/1,partialupperjawinbuccal(A),occlusal(B), and lingual (C) views." captionText-1="Fig.6. Mosasaurus cf. hoffmanni Mantell,1829,fromthe Upper Campanian opoka at Maruszów. A. ZPAL R.I/1, lefthandsidetoothinBin anterior(A1),buccal(A2),and posterior (A3) views. B. ZPAL R.I/1, three tooth crowns preserved in original jaw position (although jaw bone is not preserved)." figureDoi-0="http://doi.org/10.5281/zenodo.13345886" figureDoi-1="http://doi.org/10.5281/zenodo.13345888" httpUri-0="https://zenodo.org/record/13345886/files/figure.png" httpUri-1="https://zenodo.org/record/13345888/files/figure.png" pageId="3" pageNumber="400">Figs. 46</figureCitation>
) show a Ushaped cross section, withboth the buccal and lingual surfaces facetted, with buccal facets wider and more distinct than those on the lingualside.Theyalsoshowwelldevelopedanteriorandposterior carinae, a slight posterior recurvature and a more pronounced lingual recurvature. Enamel beading is less pronounced than in the Late Maastrichtian specimens, but does occur in patches, both proximally and distally.
</paragraph>
<caption id="A1F8663AD13CFF9FFFD4FAB2FE8D4533" ID-DOI="http://doi.org/10.5281/zenodo.13345886" ID-Zenodo-Dep="13345886" httpUri="https://zenodo.org/record/13345886/files/figure.png" pageId="4" pageNumber="401" startId="4.[102,139,1355,1374]" targetBox="[103,1484,225,1327]" targetPageId="4" targetType="figure">
<paragraph id="F53836B2D13CFF9FFFD4FAB2FE8D4533" blockId="4.[102,1485,1354,1402]" pageId="4" pageNumber="401">
Fig.4.
<taxonomicName id="32874D31D13CFF9FFF11FAB2FDAD4517" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[163,529,1355,1374]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFF11FAB2FEAE4514" box="[163,274,1355,1373]" italics="true" pageId="4" pageNumber="401">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D13CFF9FFE80FAB2FE324517" box="[306,398,1355,1374]" italics="true" pageId="4" pageNumber="401">hoffmanni</emphasis>
<bibRefCitation id="91164B43D13CFF9FFE20FAB2FDAD4517" author="Mantell, G. A." box="[402,529,1355,1374]" pageId="4" pageNumber="401" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell,1829</bibRefCitation>
</taxonomicName>
,fromtheUpperCampanianopokaatMaruszów,ZPALR.I/1,partialupperjawinbuccal(
<emphasis id="C7F3EAA0D13CFF9FFA8BFAB3FAF54514" bold="true" box="[1337,1353,1354,1373]" pageId="4" pageNumber="401">A</emphasis>
),occlusal(
<emphasis id="C7F3EAA0D13CFF9FFA03FAB2FA7C4517" bold="true" box="[1457,1472,1355,1374]" pageId="4" pageNumber="401">B</emphasis>
), and lingual (
<emphasis id="C7F3EAA0D13CFF9FFF6AFA9FFF544530" bold="true" box="[216,232,1382,1401]" pageId="4" pageNumber="401">C</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="F53836B2D13CFF9FFFD4F981FB1345AC" blockId="4.[102,775,1656,2026]" lastBlockId="4.[812,1486,1454,1952]" pageId="4" pageNumber="401">
<emphasis id="C7F3EAA0D13CFF9FFFD4F981FF5D46C6" box="[102,225,1656,1679]" italics="true" pageId="4" pageNumber="401">Discussion</emphasis>
.—Typical anteriorand medianposterior teethof adultspecimens of
<taxonomicName id="32874D31D13CFF9FFE8FF961FE6F46E6" authorityName="Mantell" authorityYear="1829" box="[317,467,1687,1711]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFE8FF961FE6F46E6" box="[317,467,1687,1711]" italics="true" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
arerobust,bicarinate,have asymmetrical crowns (lingualsidemore inflated)withafew wide “prismatic” facets on the buccal side of the crown, and more numerous facets on the lingual side; they also reveal beading of the enamel (
<bibRefCitation id="91164B43D13CFF9FFE31F8ECFDC34764" author="Lingham-Soliar, T." box="[387,639,1813,1837]" pageId="4" pageNumber="401" pagination="155 - 180" refId="ref6951" refString="Lingham-Soliar, T. 1995. Anatomy and functional morphology of the largest marine reptile known, Mosasaurus hoffmanni (Mosasauridae, Reptilia) from the Upper Cretaceous, Upper Maastrichtian of The Netherlands. PhilosophicalTransactions ofthe RoyalSociety ofLondon B 347: 155 - 180." type="journal article" year="1995">LinghamSoliar 1995</bibRefCitation>
;
<bibRefCitation id="91164B43D13CFF9FFD22F8ECFF1D4705" author="Jagt, J. W. M. &amp; Kuypers, M. M. M. &amp; andPeeters, H. H. G." pageId="4" pageNumber="401" pagination="47 - 59" refId="ref6521" refString="Jagt, J. W. M., Kuypers, M. M. M., andPeeters, H. H. G. 1995. Mosasauriersin het Natuurhistorisch Museum Maastricht opnieuw onder de loep genomen. Natuurhistorisch Maandblad 84: 47 - 59." type="journal article" year="1995">Jagt et al. 1995</bibRefCitation>
;
<bibRefCitation id="91164B43D13CFF9FFF03F8CCFE224705" author="Kuypers, M. M. M. &amp; Jagt, J. W. M. &amp; Peeters, H. H. G. &amp; De Graaf, D. Th. &amp; Dortangs, R. W. &amp; Deckers, M. J. M. &amp; Eysermans, D. &amp; Janssen, M. J. &amp; Arpot, L." box="[177,414,1844,1869]" pageId="4" pageNumber="401" pagination="4 - 47" refId="ref6661" refString="Kuypers, M. M. M., Jagt, J. W. M., Peeters, H. H. G., De Graaf, D. Th., Dortangs, R. W., Deckers, M. J. M., Eysermans, D., Janssen, M. J., and Arpot, L. 1998. Laat-kretaceische mosasauriers uit Luik-Limburg. Nieuwe vondsten leiden tot nieuwe inzichten. Publicaties van het Natuurhistorisch Genootschap in Limburg 41: 4 - 47." type="journal article" year="1998">Kuypers et al. 1998</bibRefCitation>
). The posterior teeth are more symmetrical, but are still characterised by beaded enamel and distinct facets (op. cit.). The beading of the enamel was held tobe diagnostic of teeth of
<taxonomicName id="32874D31D13CFF9FFE7EF86DFDDF47E2" authorityName="Mantell" authorityYear="1829" box="[460,611,1939,1963]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFE7EF86DFDDF47E2" box="[460,611,1939,1963]" italics="true" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
by
<bibRefCitation id="91164B43D13CFF9FFD3EF86AFF644783" author="Kuypers, M. M. M. &amp; Jagt, J. W. M. &amp; Peeters, H. H. G. &amp; De Graaf, D. Th. &amp; Dortangs, R. W. &amp; Deckers, M. J. M. &amp; Eysermans, D. &amp; Janssen, M. J. &amp; Arpot, L." pageId="4" pageNumber="401" pagination="4 - 47" refId="ref6661" refString="Kuypers, M. M. M., Jagt, J. W. M., Peeters, H. H. G., De Graaf, D. Th., Dortangs, R. W., Deckers, M. J. M., Eysermans, D., Janssen, M. J., and Arpot, L. 1998. Laat-kretaceische mosasauriers uit Luik-Limburg. Nieuwe vondsten leiden tot nieuwe inzichten. Publicaties van het Natuurhistorisch Genootschap in Limburg 41: 4 - 47." type="journal article" year="1998">Kuypers et al. (1998)</bibRefCitation>
. However, it occurs in African and Polish specimens herein referred to as “
<taxonomicName id="32874D31D13CFF9FFE7BF82BFD6C47A3" box="[457,720,2002,2026]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="subGenus" subGenus="Leiodon">
<emphasis id="C7F3EAA0D13CFF9FFE7BF82BFDEE47A0" box="[457,594,2002,2025]" italics="true" pageId="4" pageNumber="401">Mosasaurus</emphasis>
(
<emphasis id="C7F3EAA0D13CFF9FFDD9F82BFD7B47A3" box="[619,711,2002,2026]" italics="true" pageId="4" pageNumber="401">Leiodon</emphasis>
)
</taxonomicName>
cfr.
<taxonomicName id="32874D31D13CFF9FFC9EFA57FCC6458C" authorityName="Owen" authorityYear="1845" box="[812,890,1454,1477]" class="Elasmobranchii" family="Tetraodontidae" genus="Leiodon" kingdom="Animalia" order="Tetraodontiformes" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="anceps">
<emphasis id="C7F3EAA0D13CFF9FFC9EFA57FCC6458C" box="[812,890,1454,1477]" italics="true" pageId="4" pageNumber="401">anceps</emphasis>
</taxonomicName>
”and inone ofthePolishspecimens referred toas
<taxonomicName id="32874D31D13CFF9FFA1DFA57FC7545AC" authorityName="Dollo" authorityYear="1889" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="lemonnieri">
<emphasis id="C7F3EAA0D13CFF9FFA1DFA57FA71458C" box="[1455,1485,1454,1477]" italics="true" pageId="4" pageNumber="401">M.</emphasis>
cf.
<emphasis id="C7F3EAA0D13CFF9FFCE2FA34FC7545AC" box="[848,969,1485,1509]" italics="true" pageId="4" pageNumber="401">lemonnieri</emphasis>
</taxonomicName>
as well (see below).
</paragraph>
<paragraph id="F53836B2D13CFF9FFCE0FA17FC55468B" blockId="4.[812,1486,1454,1952]" pageId="4" pageNumber="401">
TheLateMaastrichtianspecimensIGPUWAR14from Nasiłów fall within the range of dental variation of adult specimens of
<taxonomicName id="32874D31D13CFF9FFC74F9D4FBE1460C" authorityName="Mantell" authorityYear="1829" box="[966,1117,1581,1605]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFC74F9D4FBE1460C" box="[966,1117,1581,1605]" italics="true" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
from the environs of Maastricht. The single tooth from the upper Upper Maastrichtian opoka of Nasiłów (MZ VIII/Vr66), referred toby
<bibRefCitation id="91164B43D13CFF9FFAAAF995FA7146CD" author="Sulimski, A." box="[1304,1485,1644,1668]" pageId="4" pageNumber="401" pagination="243 - 250" refId="ref7863" refString="Sulimski, A. 1968. Remains of Upper Cretaceous Mosasauridae (Reptilia) of central Poland. Acta Palaeontologica Polonica 13: 243 - 250." type="journal article" year="1968">Sulimski(1968)</bibRefCitation>
as
<taxonomicName id="32874D31D13CFF9FFCFBF975FC4A46EA" box="[841,1014,1675,1699]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="C7F3EAA0D13CFF9FFCFBF975FC6E46EA" box="[841,978,1676,1699]" italics="true" pageId="4" pageNumber="401">Mosasaurus</emphasis>
sp.
</taxonomicName>
(sp.B),alsofallswithintheadultcategory of
<emphasis id="C7F3EAA0D13CFF9FFCF9F952FC55468B" box="[843,1001,1706,1730]" italics="true" pageId="4" pageNumber="401">
<taxonomicName id="32874D31D13CFF9FFCF9F952FC59468B" authorityName="Mantell" authorityYear="1829" box="[843,997,1706,1730]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">M. hoffmanni</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="F53836B2D13CFF9FFCE0F932FAD247E9" blockId="4.[812,1486,1454,1952]" pageId="4" pageNumber="401">
Teeth of subadult
<taxonomicName id="32874D31D13CFF9FFBACF935FB0946AA" authorityName="Mantell" authorityYear="1829" box="[1054,1205,1739,1763]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFBACF935FB0946AA" box="[1054,1205,1739,1763]" italics="true" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
are more slender. Specimen OGP 1254 from Nasiłów may belong here. The size of the tooth crowns of the Upper Campanian specimen ZPAL R. I/15 from Maruszów suggests that these remains originatefromasubadultindividualaswell.Itiswellmatchedby smaller sized material of
<taxonomicName id="32874D31D13CFF9FFBF9F890FB5F47C9" authorityName="Mantell" authorityYear="1829" box="[1099,1251,1896,1920]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9FFBF9F890FB5F47C9" box="[1099,1251,1896,1920]" italics="true" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
from the Maastrichtian
<typeStatus id="2A3C8810D13CFF9FFCEFF871FC3147E9" box="[861,909,1928,1952]" pageId="4" pageNumber="401">type</typeStatus>
area (e.g.,
<bibRefCitation id="91164B43D13CFF9FFBBBF871FB5547E9" author="Kuypers, M. M. M. &amp; Jagt, J. W. M. &amp; Peeters, H. H. G. &amp; De Graaf, D. Th. &amp; Dortangs, R. W. &amp; Deckers, M. J. M. &amp; Eysermans, D. &amp; Janssen, M. J. &amp; Arpot, L." box="[1033,1257,1928,1952]" pageId="4" pageNumber="401" pagination="4 - 47" refId="ref6661" refString="Kuypers, M. M. M., Jagt, J. W. M., Peeters, H. H. G., De Graaf, D. Th., Dortangs, R. W., Deckers, M. J. M., Eysermans, D., Janssen, M. J., and Arpot, L. 1998. Laat-kretaceische mosasauriers uit Luik-Limburg. Nieuwe vondsten leiden tot nieuwe inzichten. Publicaties van het Natuurhistorisch Genootschap in Limburg 41: 4 - 47." type="journal article" year="1998">Kuypers et al. 1998</bibRefCitation>
: pl. 1: 5, 6).
</paragraph>
<paragraph id="F53836B2D13CFF9EFC9EF84BFCBB47A3" blockId="4.[812,1486,1970,2026]" lastBlockId="5.[102,775,1872,2026]" lastPageId="5" lastPageNumber="402" pageId="4" pageNumber="401">
<emphasis id="C7F3EAA0D13CFF9FFC9EF84BFB7A4783" box="[812,1222,1970,1994]" italics="true" pageId="4" pageNumber="401">Stratigraphic and geographic range</emphasis>
.—In the Maastrichtian
<typeStatus id="2A3C8810D13CFF9FFC9EF82BFCE747A3" box="[812,859,2002,2026]" pageId="4" pageNumber="401">type</typeStatus>
area, the lowermost welldocumented occurrence of
<taxonomicName id="32874D31D13CFF9EFA1DF82BFF694721" authorityName="Mantell" authorityYear="1829" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="5" lastPageNumber="402" order="Squamata" pageId="4" pageNumber="401" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13CFF9EFA1DF82BFF694721" italics="true" lastPageId="5" lastPageNumber="402" pageId="4" pageNumber="401">M. hoffmanni</emphasis>
</taxonomicName>
,basedonthediagnosticskeletalmaterial,iswithin the upper half of the Lanaye Member (Gulpen Formation, lower Upper Maastrichtian
<taxonomicName id="32874D31D13DFF9EFE2BF868FD0F47E0" authority="Zone" authorityName="Zone" box="[409,691,1937,1961]" class="Cephalopoda" family="Dimitobelidae" genus="Belemnitella" kingdom="Animalia" order="Belemnitida" pageId="5" pageNumber="402" phylum="Mollusca" rank="species" species="junior">
<emphasis id="C7F3EAA0D13DFF9EFE2BF868FDD247E1" box="[409,622,1937,1961]" italics="true" pageId="5" pageNumber="402">Belemnitella junior</emphasis>
Zone
</taxonomicName>
). However, finds of isolated teeth and tooth crowns from the Vijlen Member (Gulpen Formation, Lower Maastrichtian) in the
</paragraph>
<paragraph id="F53836B2D13DFF9EFC62FD0FFC62435B" blockId="5.[932,1485,758,950]" box="[976,990,758,786]" pageId="5" pageNumber="402">
<emphasis id="C7F3EAA0D13DFF9EFC62FD0FFC62435B" box="[976,990,758,786]" italics="true" pageId="5" pageNumber="402">Ą</emphasis>
</paragraph>
<paragraph id="F53836B2D13DFF9EFC74FCE7FBBE43E8" blockId="5.[932,1485,758,950]" pageId="5" pageNumber="402">
Fig.5.
<taxonomicName id="32874D31D13DFF9EFBB7FCE6FAC44378" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[1029,1400,798,817]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFBB7FCE6FBC84378" box="[1029,1140,799,817]" italics="true" pageId="5" pageNumber="402">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D13DFF9EFB24FCE7FB4E4378" box="[1174,1266,798,817]" italics="true" pageId="5" pageNumber="402">hoffmanni</emphasis>
<bibRefCitation id="91164B43D13DFF9EFB45FCE7FAC44378" author="Mantell, G. A." box="[1271,1400,798,817]" pageId="5" pageNumber="402" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell,1829</bibRefCitation>
</taxonomicName>
,fromthe Upper Campanian opoka at Maruszów.
<emphasis id="C7F3EAA0D13DFF9EFA9CFCC3FA824304" bold="true" box="[1326,1342,826,845]" pageId="5" pageNumber="402">A</emphasis>
. ZPAL R.I/
<quantity id="327F9B57D13DFF9EFA18FCC3FA714304" box="[1450,1485,826,845]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="402" unit="in" value="4.0">4 in</quantity>
lingual(A
<subScript id="690334F7D13DFF9EFB93FCA6FB954324" attach="right" box="[1057,1065,863,877]" fontSize="6" pageId="5" pageNumber="402">1</subScript>
)andanterior(A
<subScript id="690334F7D13DFF9EFB0EFCA6FB784324" attach="left" box="[1212,1220,863,877]" fontSize="6" pageId="5" pageNumber="402">2</subScript>
)views.
<emphasis id="C7F3EAA0D13DFF9EFABDFCAFFAA24320" bold="true" box="[1295,1310,854,873]" pageId="5" pageNumber="402">B</emphasis>
.ZPALR.I/2inlingual (B
<subScript id="690334F7D13DFF9EFBBEFC82FBA843C0" attach="left" box="[1036,1044,891,905]" fontSize="6" pageId="5" pageNumber="402">1</subScript>
), anterior
<emphasis id="C7F3EAA0D13DFF9EFBCAFC8BFB3C43CC" bold="true" box="[1144,1152,882,901]" pageId="5" pageNumber="402">(</emphasis>
B
<subScript id="690334F7D13DFF9EFB3DFC82FB2B43C0" attach="left" box="[1167,1175,891,905]" fontSize="6" pageId="5" pageNumber="402">2</subScript>
), posterior (B
<subScript id="690334F7D13DFF9EFAAEFC82FA9843C0" attach="left" box="[1308,1316,891,905]" fontSize="6" pageId="5" pageNumber="402">3</subScript>
), and buccal (B
<subScript id="690334F7D13DFF9EFA0FFC82FA7943C0" attach="left" box="[1469,1477,891,905]" fontSize="6" pageId="5" pageNumber="402">4</subScript>
) views.
</paragraph>
<paragraph id="F53836B2D13DFF9EFC16FC64FC7C43FF" blockId="5.[932,1485,758,950]" box="[932,960,925,950]" pageId="5" pageNumber="402">
<emphasis id="C7F3EAA0D13DFF9EFC16FC64FC7C43FF" box="[932,960,925,950]" italics="true" pageId="5" pageNumber="402">Ą</emphasis>
</paragraph>
<caption id="A1F8663AD13DFF9EFC74FC33FBF34420" ID-DOI="http://doi.org/10.5281/zenodo.13345888" ID-Zenodo-Dep="13345888" httpUri="https://zenodo.org/record/13345888/files/figure.png" pageId="5" pageNumber="402" startId="5.[966,1004,970,989]" targetBox="[104,932,799,1839]" targetPageId="5" targetType="figure">
<paragraph id="F53836B2D13DFF9EFC74FC33FBF34420" blockId="5.[966,1485,970,1130]" pageId="5" pageNumber="402">
Fig.6.
<taxonomicName id="32874D31D13DFF9EFBB7FC32FAC44394" authority="Mantell, 1829" authorityName="Mantell" authorityYear="1829" box="[1029,1400,970,989]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFBB7FC32FBC84394" box="[1029,1140,971,989]" italics="true" pageId="5" pageNumber="402">Mosasaurus</emphasis>
cf.
<emphasis id="C7F3EAA0D13DFF9EFB24FC33FB4E4394" box="[1174,1266,970,989]" italics="true" pageId="5" pageNumber="402">hoffmanni</emphasis>
<bibRefCitation id="91164B43D13DFF9EFB45FC33FAC44394" author="Mantell, G. A." box="[1271,1400,970,989]" pageId="5" pageNumber="402" pagination="201 - 216" refId="ref7241" refString="Mantell, G. A. 1829. A tabular arrangement of the organic remains of the countyofSussex. Transactions of the Geological Society of London (2) 3: 201 - 216." type="journal article" year="1829">Mantell,1829</bibRefCitation>
</taxonomicName>
,fromthe Upper Campanian opoka at Maruszów.
<emphasis id="C7F3EAA0D13DFF9EFA88FC1FFAF643B0" bold="true" box="[1338,1354,998,1017]" pageId="5" pageNumber="402">A</emphasis>
. ZPAL R.I/1, lefthandsidetoothinBin anterior(A
<subScript id="690334F7D13DFF9EFA93FBF2FA954450" attach="left" box="[1313,1321,1035,1049]" fontSize="6" pageId="5" pageNumber="402">1</subScript>
),buccal(A
<subScript id="690334F7D13DFF9EFA20FBF2FA264450" attach="left" box="[1426,1434,1035,1049]" fontSize="6" pageId="5" pageNumber="402">2</subScript>
),and posterior (A
<subScript id="690334F7D13DFF9EFB87FBDEFB81447C" attach="right" box="[1077,1085,1063,1077]" fontSize="6" pageId="5" pageNumber="402">3</subScript>
) views.
<emphasis id="C7F3EAA0D13DFF9EFB3FFBE6FB20447B" bold="true" box="[1165,1180,1055,1074]" pageId="5" pageNumber="402">B</emphasis>
. ZPAL R.I/1, three tooth crowns preserved in original jaw position (although jaw bone is not preserved).
</paragraph>
</caption>
<paragraph id="F53836B2D13DFF9EFC77FB5FFB09461E" blockId="5.[846,1486,1190,1623]" pageId="5" pageNumber="402">
HaccourtLixhearea(Liège,NEBelgium),suggest that
<taxonomicName id="32874D31D13DFF9EFB9BFB3FFB024494" authorityName="Mantell" authorityYear="1829" box="[1065,1214,1221,1245]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFB9BFB3FFB024494" box="[1065,1214,1221,1245]" italics="true" pageId="5" pageNumber="402">M. hoffmanni</emphasis>
</taxonomicName>
or a closely related speciesmadeitsfirstappearanceintheareaearlier. The species is fairly common in the Emael and Nekummembers(MaastrichtFormation,Upper Maastrichtian), in particular in the Eben Emael (Bassenge) area (Liège). The
<typeStatus id="2A3C8810D13DFF9EFAA5FA9AFAFA4532" box="[1303,1350,1379,1403]" pageId="5" pageNumber="402">type</typeStatus>
of
<taxonomicName id="32874D31D13DFF9EFAC3FA9AFBB645D3" authorityName="Mantell" authorityYear="1829" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFAC3FA9AFBB645D3" italics="true" pageId="5" pageNumber="402">M. hoffmanni</emphasis>
</taxonomicName>
(MNHN AC 9648) is from the upper third of the Nekum Member (Bardet and Jagt 1996). The species ranges up to the K/Pg boundary. Reworked tooth crowns have been recorded from the basal Geulhem Member (Houthem Formation) of Early Paleocene age (
<bibRefCitation id="91164B43D13DFF9EFC7EF9C6FB15461E" author="Kuypers, M. M. M. &amp; Jagt, J. W. M. &amp; Peeters, H. H. G. &amp; De Graaf, D. Th. &amp; Dortangs, R. W. &amp; Deckers, M. J. M. &amp; Eysermans, D. &amp; Janssen, M. J. &amp; Arpot, L." box="[972,1193,1599,1623]" pageId="5" pageNumber="402" pagination="4 - 47" refId="ref6661" refString="Kuypers, M. M. M., Jagt, J. W. M., Peeters, H. H. G., De Graaf, D. Th., Dortangs, R. W., Deckers, M. J. M., Eysermans, D., Janssen, M. J., and Arpot, L. 1998. Laat-kretaceische mosasauriers uit Luik-Limburg. Nieuwe vondsten leiden tot nieuwe inzichten. Publicaties van het Natuurhistorisch Genootschap in Limburg 41: 4 - 47." type="journal article" year="1998">Kuypers et al. 1998</bibRefCitation>
).
</paragraph>
<paragraph id="F53836B2D13DFF9EFC58F98DFA714701" blockId="5.[812,1486,1652,2026]" pageId="5" pageNumber="402">
Other records of
<taxonomicName id="32874D31D13DFF9EFB1DF98DFAFB46C5" authorityName="Mantell" authorityYear="1829" box="[1199,1351,1652,1676]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFB1DF98DFAFB46C5" box="[1199,1351,1652,1676]" italics="true" pageId="5" pageNumber="402">M. hoffmanni</emphasis>
</taxonomicName>
include the Upper Campanian to Upper Maastrichtian of
<collectingRegion id="3743F850D13DFF9EFC77F94AFBF94682" box="[965,1093,1715,1739]" country="United States of America" name="New Jersey" pageId="5" pageNumber="402">New Jersey</collectingRegion>
(
<collectingCountry id="8D907622D13DFF9EFBE6F94AFB324682" box="[1108,1166,1715,1739]" name="United States of America" pageId="5" pageNumber="402">USA</collectingCountry>
) (
<bibRefCitation id="91164B43D13DFF9EFB16F94BFA894682" author="Mulder, E. W. A." box="[1188,1333,1714,1739]" pageId="5" pageNumber="402" pagination="281 - 300" refId="ref7308" refString="Mulder, E. W. A. 1999. TransatlanticlatestCretaceousmosasaurs (Reptilia, Lacertilia) fromtheMaastrichtiantypeareaandNew Jersey. Geologie en Mijnbouw 78: 281 - 300." type="journal article" year="1999">Mulder 1999</bibRefCitation>
synonymised
<taxonomicName id="32874D31D13DFF9EFC77F92AFA8A46A3" authority="Cope, 1869" authorityName="Cope" authorityYear="1869" box="[965,1334,1746,1770]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="maximus">
<emphasis id="C7F3EAA0D13DFF9EFC77F92AFB0E46A3" box="[965,1202,1747,1770]" italics="true" pageId="5" pageNumber="402">Mosasaurus maximus</emphasis>
<bibRefCitation id="91164B43D13DFF9EFB05F92BFA8A46A3" author="Cope, E. D." box="[1207,1334,1746,1770]" pageId="5" pageNumber="402" pagination="250 - 266" refId="ref6092" refString="Cope, E. D. 1869. OnthereptilianordersPythonomorphaandStreptosauria. Proceedings of the Boston Society of Natural History 12: 250 - 266." type="journal article" year="1869">Cope,1869</bibRefCitation>
</taxonomicName>
,with
<taxonomicName id="32874D31D13DFF9EFAC4F92AFBB64740" authorityName="Mantell" authorityYear="1829" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFAC4F92AFBB64740" italics="true" pageId="5" pageNumber="402">M. hoffmanni</emphasis>
</taxonomicName>
), the uppermost Maastrichtian of
<collectingRegion id="3743F850D13DFF9EFA2BF908FC404760" country="United States of America" name="Missouri" pageId="5" pageNumber="402">Missouri</collectingRegion>
(
<bibRefCitation id="91164B43D13DFF9EFBA1F8E8FA874760" author="Campbell, C. &amp; Lee, T." box="[1043,1339,1809,1833]" pageId="5" pageNumber="402" refId="ref6006" refString="Campbell, C. and Lee, T. 2001. &quot; Tails of hoffmanni &quot;: mosasaur fossils in tsunamidepositatK / TboundaryofsoutheastMissouri. Journal of Vertebrate Paleontology 21 (Supplement to 3): 37 A." type="journal volume" year="2001">Campbell and Lee 2001</bibRefCitation>
), the Maastrichtian of
<collectingRegion id="3743F850D13DFF9EFBFAF8C9FB104701" box="[1096,1196,1840,1864]" country="United States of America" name="Alabama" pageId="5" pageNumber="402">Alabama</collectingRegion>
(
<bibRefCitation id="91164B43D13DFF9EFB0EF8C9FAE64700" author="Kiernan, C. R." box="[1212,1370,1840,1865]" pageId="5" pageNumber="402" pagination="91 - 103" refId="ref6620" refString="Kiernan, C. R. 2002. Stratigraphic distribution and habitat segregation of mosasaurs in the Upper Cretaceous of western and central Alabama, withan [sic] historicalreviewofAlabamamosasaurdiscoveries. Journal of Vertebrate Paleontology 22: 91 - 103." type="journal article" year="2002">Kiernan 2002</bibRefCitation>
),
<collectingCountry id="8D907622D13DFF9EFADDF8C9FA714701" box="[1391,1485,1840,1864]" name="Bulgaria" pageId="5" pageNumber="402">Bulgaria</collectingCountry>
</paragraph>
<paragraph id="F53836B2D13DFF9EFC9EF8A9FA714721" blockId="5.[812,1486,1652,2026]" box="[812,1485,1872,1896]" pageId="5" pageNumber="402">
(
<bibRefCitation id="91164B43D13DFF9EFC86F8A9FC6B4721" author="Tzankov, V." box="[820,983,1872,1896]" pageId="5" pageNumber="402" pagination="13 - 20" refId="ref7926" refString="Tzankov, V. 1939. Note sur la presence des reptiles fossiles du Cretace superieur de la Bulgarie du Nord. Geologica Balcanica 3: 13 - 20." type="book chapter" year="1939">Tzankov1939</bibRefCitation>
;seealsoNikolovandWestphal1976),north
</paragraph>
<paragraph id="F53836B2D13DFF9EFC9EF889FA7047C1" blockId="5.[812,1486,1652,2026]" box="[812,1484,1904,1928]" pageId="5" pageNumber="402">
ernDenmark(
<bibRefCitation id="91164B43D13DFF9EFC79F889FBEC47C1" author="Bonde, N." box="[971,1104,1904,1928]" pageId="5" pageNumber="402" pagination="26 - 30" refId="ref5961" refString="Bonde, N. 1997. En kaempemaessig mosasaur fra Israel. Varv 1: 26 - 30." type="journal article" year="1997">Bonde1997</bibRefCitation>
;Jagtpersonalobservation),
<collectingCountry id="8D907622D13DFF9EFACDF889FA7047C1" box="[1407,1484,1904,1928]" name="Democratic Republic of the Congo" pageId="5" pageNumber="402">Congo</collectingCountry>
</paragraph>
<paragraph id="F53836B2D13DFF9EFC9EF868FA7247E0" blockId="5.[812,1486,1652,2026]" box="[812,1486,1937,1961]" pageId="5" pageNumber="402">
(
<collectingCountry id="8D907622D13DFF9EFC86F868FCCD47E0" box="[820,881,1937,1961]" name="Democratic Republic of the Congo" pageId="5" pageNumber="402">Zaire</collectingCountry>
) (
<bibRefCitation id="91164B43D13DFF9EFC26F868FB1E47E0" author="Lingham-Soliar, T." box="[916,1186,1937,1961]" pageId="5" pageNumber="402" pagination="259 - 265" refId="ref6930" refString="Lingham-Soliar, T. 1994 b. FirstrecordofmosasaursfromtheMaastrichtian (UpperCretaceous) ofZaire. Palaontologische Zeitschrift 68: 259 - 265." type="journal article" year="1994">LinghamSoliar 1994b</bibRefCitation>
),
<collectingCountry id="8D907622D13DFF9EFB71F868FABE47E0" box="[1219,1282,1937,1961]" name="Niger" pageId="5" pageNumber="402">Niger</collectingCountry>
(
<bibRefCitation id="91164B43D13DFF9EFAACF868FA7247E0" author="Lingham-Soliar, T." box="[1310,1486,1937,1961]" pageId="5" pageNumber="402" pagination="653 - 670" refId="ref6851" refString="Lingham-Soliar, T. 1991. Mosasaurs from the Upper Cretaceous of Niger. Palaeontology 34: 653 - 670." type="journal article" year="1991">LinghamSoliar</bibRefCitation>
</paragraph>
<paragraph id="F53836B2D13DFF9EFC9EF848FA724780" blockId="5.[812,1486,1652,2026]" box="[812,1486,1969,1993]" pageId="5" pageNumber="402">
1991),andTurkey(BardetandTunoǧlu2002).
<bibRefCitation id="91164B43D13DFF9EFA94F848FA724780" author="Persson, P. O." box="[1318,1486,1969,1993]" pageId="5" pageNumber="402" pagination="431 - 478" refId="ref7537" refString="Persson, P. O. 1959. ReptilesfromtheSenonian (U. Cret.) ofScania (S. Sweden). Arkiv for Mineralogi och Geologi 2: 431 - 478." type="journal article" year="1959">Persson(1959:</bibRefCitation>
</paragraph>
<paragraph id="F53836B2D13DFF9EFC9EF82BFA7247A3" blockId="5.[812,1486,1652,2026]" box="[812,1486,2002,2026]" pageId="5" pageNumber="402">
462, fig. 10; pl. 15: 1, 2) recorded
<taxonomicName id="32874D31D13DFF9EFB1CF82BFADB47A3" authorityName="Mantell" authorityYear="1829" box="[1198,1383,2002,2026]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="402" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13DFF9EFB1CF82BFB7947A0" box="[1198,1221,2002,2025]" italics="true" pageId="5" pageNumber="402">M</emphasis>
. cf.
<emphasis id="C7F3EAA0D13DFF9EFB45F82BFADB47A3" box="[1271,1383,2002,2026]" italics="true" pageId="5" pageNumber="402">hoffmanni</emphasis>
</taxonomicName>
from the
</paragraph>
<paragraph id="F53836B2D13EFF9DFFD4FF10FEC74109" blockId="6.[102,775,232,510]" pageId="6" pageNumber="403">
Lower Campanian of southern
<collectingCountry id="8D907622D13EFF9DFE76FF11FDA34149" box="[452,543,232,256]" name="Sweden" pageId="6" pageNumber="403">Sweden</collectingCountry>
. This material, however, refers to
<taxonomicName id="32874D31D13EFF9DFEB6FEF1FD2C4169" authority="(Persson, 1963)" baseAuthorityName="Persson" baseAuthorityYear="1963" box="[260,656,264,288]" class="Reptilia" family="Mosasauridae" genus="Tylosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="403" phylum="Chordata" rank="species" species="ivoensis">
<emphasis id="C7F3EAA0D13EFF9DFEB6FEF1FE604169" box="[260,476,264,288]" italics="true" pageId="6" pageNumber="403">Tylosaurus ivoensis</emphasis>
(
<bibRefCitation id="91164B43D13EFF9DFE59FEF1FD3B4169" author="Persson, P. O." box="[491,647,264,288]" pageId="6" pageNumber="403" pagination="1 - 15" refId="ref7570" refString="Persson, P. O. 1963. StudiesonMesozoicmarinereptilefaunaswithparticular regardtothePlesiosauria. Publications from the Institutes of Mineralogy, Paleontology and Quaternary Geology, University of Lund 118: 1 - 15." type="journal article" year="1963">Persson, 1963</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="91164B43D13EFF9DFD7BFEF1FED34109" author="Lindgren, J. &amp; Siverson, M." pageId="6" pageNumber="403" pagination="73 - 93" refId="ref6816" refString="Lindgren, J. and Siverson, M. 2002. Tylosaurus ivoensis: a giant mosasaur fromtheearlyCampanianofSweden. Transactions of the Royal Society of Edinburgh, Earth Sciences 93: 73 - 93." type="journal article" year="2002">Lindgren and Siverson 2002</bibRefCitation>
).
</paragraph>
<paragraph id="F53836B2D13EFF9DFF39FEB1FEDC41B7" blockId="6.[102,775,232,510]" pageId="6" pageNumber="403">
InEurope,welldocumentedremainsof
<taxonomicName id="32874D31D13EFF9DFDFAFEB0FD624129" authorityName="Mantell" authorityYear="1829" box="[584,734,328,352]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="403" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13EFF9DFDFAFEB0FD624129" box="[584,734,328,352]" italics="true" pageId="6" pageNumber="403">M. hoffmanni</emphasis>
</taxonomicName>
appear to be confined to Upper Maastrichtian strata. However, the specimen from Maruszów suggests that
<taxonomicName id="32874D31D13EFF9DFDE3FE71FD5441D6" authorityName="Mantell" authorityYear="1829" box="[593,744,391,415]" class="Reptilia" family="Mosasauridae" genus="Mosasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="403" phylum="Chordata" rank="species" species="hoffmanni">
<emphasis id="C7F3EAA0D13EFF9DFDE3FE71FD5441D6" box="[593,744,391,415]" italics="true" pageId="6" pageNumber="403">M. hoffmanni</emphasis>
</taxonomicName>
or a closely related (ancestral?) species appeared in Europe as earlyastheLateCampanian.Thismaybeconfirmedonlyby more diagnostic finds.
</paragraph>
</subSubSection>
</treatment>
</document>