206 lines
23 KiB
XML
206 lines
23 KiB
XML
<document id="50D25D46748C4B9300517308DD1EC35F" ID-DOI="10.1649/0010-065X-76.4.577" ID-ISSN="1938-4394" ID-Zenodo-Dep="13252887" ID-ZooBank="urn:lsid:zoobank.org:pub:88AEA250-3D9E-40D3-B393-32C4FB04DAF3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723003834567" checkinUser="felipe" docAuthor="Ivie, Michael A., Fanti, Fabrizio & Ferreira, Vinicius S." docDate="2022" docId="363F87BCE051A776FF07FBFCFDECEA4D" docLanguage="en" docName="TheColeopBull.76.4.577-583.pdf" docOrigin="The Coleopterists Bulletin 76 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-76.4.577" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Tytthonyx (Tytthonyx) milleri Ivie, Fanti, and Ferreira 2022, new species" docType="treatment" docUuid="3175554E-150F-4ECF-9312-B46A65796F73" docUuidSource="ZooBank" docVersion="2" lastPageNumber="578" masterDocId="CA06FFC4E053A772FFB9FFDEFFC6EB35" masterDocTitle="A New Tytthonyx LeConte (Coleoptera: Cantharidae) from Dominican Amber, with Notes on Diagnostic Characters of West Indian Cantharids with Short Elytra" masterLastPageNumber="583" masterPageNumber="577" pageNumber="578" updateTime="1723006793479" updateUser="ExternalLinkService">
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<mods:title id="5B3B1A5319E222D24173F1CB1E59DDEF">A New Tytthonyx LeConte (Coleoptera: Cantharidae) from Dominican Amber, with Notes on Diagnostic Characters of West Indian Cantharids with Short Elytra</mods:title>
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<mods:namePart id="CE32502B89EDCE927EE875B0A457B24E">Ivie, Michael A.</mods:namePart>
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<mods:namePart id="64677DF14B4514D56252C37A2845C14C">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="12A1D68B0FA8FBBF27A405D4F1DB6623">Ferreira, Vinicius S.</mods:namePart>
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<mods:title id="37AE22AAD10C69D48F4BB236F53811A1">The Coleopterists Bulletin</mods:title>
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<mods:part id="98D39852A97A283C9A188BA13A2DD404">
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<mods:date id="28D7E5F0BB59CC03D73504BE03FD8DE9">2022</mods:date>
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<paragraph id="BE2936AAE051A770FF07FBFCFE61EF94" blockId="2.[175,614,1057,1186]" pageId="2" pageNumber="578">
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<taxonomicName id="79964D29E051A770FF07FBFCFE4BEF67" authority="Ivie, Fanti, and Ferreira" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri" status="sp. nov." subGenus="Tytthonyx">
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<emphasis id="8CE2EAB8E051A770FF07FBFCFEE7EF02" bold="true" box="[190,289,1058,1079]" italics="true" pageId="2" pageNumber="578">TYttHONYX</emphasis>
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(
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<emphasis id="8CE2EAB8E051A770FE96FBFFFE54EF02" bold="true" box="[303,402,1057,1079]" italics="true" pageId="2" pageNumber="578">Tytthonyx</emphasis>
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)
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<emphasis id="8CE2EAB8E051A770FE19FBFFFE27EF02" bold="true" box="[416,481,1057,1079]" italics="true" pageId="2" pageNumber="578">milleri</emphasis>
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Ivie, Fanti, and Ferreira
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</taxonomicName>
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,
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zoobank.org/
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<figureCitation id="26AD2A2FE051A770FED6FB52FE61EF94" box="[367,423,1164,1186]" captionStart="Fig" captionStartId="3.[197,228,1644,1663]" captionTargetBox="[173,1188,193,1617]" captionTargetId="figure-7@3.[173,1188,193,1617]" captionTargetPageId="3" captionText="Fig. 1. Tytthonyx (Tytthonyx) milleri, new species in Dominican amber. Holotype (WIBF 049821). A) Right lateral view, scale bar = 1.0 mm, B) Left lateral view, scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13252889" httpUri="https://zenodo.org/record/13252889/files/figure.png" pageId="2" pageNumber="578">Fig. 1</figureCitation>
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<subSubSection id="F68C6521E051A770FF09FB69FE6CEE58" pageId="2" pageNumber="578" type="materials_examined">
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<paragraph id="BE2936AAE051A770FF09FB69FE30EFD2" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
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<emphasis id="8CE2EAB8E051A770FF09FB69FE8AEFF8" bold="true" box="[176,332,1207,1229]" pageId="2" pageNumber="578">Type Specimen.</emphasis>
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<materialsCitation id="0EFE3CF7E051A770FEE8FB69FE30EFD2" collectionCode="WIBF" pageId="2" pageNumber="578" specimenCode="WIBF 049821" specimenCount="1" typeStatus="holotype">
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<typeStatus id="612D8808E051A770FEE8FB69FE6AEFF8" box="[337,428,1207,1229]" pageId="2" pageNumber="578" type="holotype">Holotype</typeStatus>
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: female, in Dominican amber, accession no.
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<specimenCode id="EE309ED1E051A770FEDDFB0CFE37EFD2" box="[356,497,1234,1255]" collectionCode="WIBF" country="USA" httpUri="http://grbio.org/cool/mzn7-y8yj" name="West Indian Beetle Fauna Project Collection" pageId="2" pageNumber="578">WIBF 049821</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="BE2936AAE051A770FF09FB32FED9EE28" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
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<materialsCitation id="0EFE3CF7E051A770FF09FB32FEDDEE28" country="Dominican Republic" location="Cordillera Septentrional" pageId="2" pageNumber="578" specimenCount="1" typeStatus="holotype">
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<emphasis id="8CE2EAB8E051A770FF09FB32FE87EE37" bold="true" box="[176,321,1260,1282]" pageId="2" pageNumber="578">
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<typeStatus id="612D8808E051A770FF09FB32FF24EE34" box="[176,226,1260,1281]" pageId="2" pageNumber="578">Type</typeStatus>
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Locality.
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</emphasis>
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||
<collectingCountry id="C681763AE051A770FEFEFB32FDD4EE37" box="[327,530,1260,1282]" name="Dominican Republic" pageId="2" pageNumber="578">Dominican Republic</collectingCountry>
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:
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<location id="BB496071E051A770FDA5FB32FEDDEE28" LSID="urn:lsid:plazi:treatment:363F87BCE051A776FF07FBFCFDECEA4D:BB496071E051A770FDA5FB32FEDDEE28" country="Dominican Republic" name="Cordillera Septentrional" pageId="2" pageNumber="578">Cordillera Septentrional</location>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="BE2936AAE051A770FF09FAFCFE6CEE58" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
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<emphasis id="8CE2EAB8E051A770FF09FAFCFE8DEE02" bold="true" box="[176,331,1313,1335]" pageId="2" pageNumber="578">
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<typeStatus id="612D8808E051A770FF09FAFCFF24EE02" box="[176,226,1314,1335]" pageId="2" pageNumber="578">Type</typeStatus>
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Horizon.
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</emphasis>
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Late Early/Middle Miocene; 15–20 million years ago (mya), probably close to 16 mya. La Toca Formation.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F68C6521E051A770FF09FAACFDD0ED38" pageId="2" pageNumber="578" type="etymology">
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||
<paragraph id="BE2936AAE051A770FF09FAACFDD0ED38" blockId="2.[152,638,1207,1709]" pageId="2" pageNumber="578">
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<emphasis id="8CE2EAB8E051A770FF09FAACFEE5EEB2" bold="true" box="[176,291,1394,1415]" pageId="2" pageNumber="578">Etymology.</emphasis>
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The species is named after the late Richard Stuart Miller (1945–2021). Rich’s interests were mainly focused on “cantharoids”, most strongly on
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<taxonomicName id="79964D29E051A770FF0CFA1CFEC7EEED" box="[181,257,1474,1496]" class="Insecta" family="Lycidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="family">Lycidae</taxonomicName>
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and
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<taxonomicName id="79964D29E051A770FE96FA1CFE60EEED" authorityName="Rafinesque" authorityYear="1815" box="[303,422,1474,1496]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="family">Lampyridae</taxonomicName>
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, and his extensive collection was donated to the
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<collectingRegion id="7C52F848E051A770FE11FA03FE3BEEC7" box="[424,509,1501,1522]" country="United States of America" name="Montana" pageId="2" pageNumber="578">Montana</collectingRegion>
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Entomology Collection at
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<collectingRegion id="7C52F848E051A770FEA0FA29FE63ED39" box="[281,421,1527,1548]" country="United States of America" name="Montana" pageId="2" pageNumber="578">Montana State</collectingRegion>
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University.
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</paragraph>
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</subSubSection>
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<subSubSection id="F68C6521E051A770FF09F9CFFBE4E9B7" pageId="2" pageNumber="578" type="diagnosis">
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<paragraph id="BE2936AAE051A770FF09F9CFFBE4E9B7" blockId="2.[152,638,1207,1709]" lastBlockId="2.[685,1173,194,1709]" pageId="2" pageNumber="578">
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<emphasis id="8CE2EAB8E051A770FF09F9CFFEDFED12" bold="true" box="[176,281,1553,1575]" pageId="2" pageNumber="578">Diagnosis.</emphasis>
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The generic placement of the species is discussed below. Currently, four other species of short-elytra cantharids are known from Dominican amber, two of which have been described:
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<taxonomicName id="79964D29E051A770FD9EF9BCFDB8ED42" authorityName="Poinar and Fanti" authorityYear="2016" box="[551,638,1634,1655]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
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<emphasis id="8CE2EAB8E051A770FD9EF9BCFDB8ED42" box="[551,638,1634,1655]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="79964D29E051A770FF7DF9A3FEDCEDA4" authorityName="Fanti and Damgaard" authorityYear="2019" box="[196,282,1660,1682]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
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<emphasis id="8CE2EAB8E051A770FF7DF9A3FEDCEDA4" box="[196,282,1660,1682]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="79964D29E051A770FE93F9A2FE09EDA4" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[298,463,1660,1681]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
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<emphasis id="8CE2EAB8E051A770FE93F9A2FE09EDA4" box="[298,463,1660,1681]" italics="true" pageId="2" pageNumber="578">Tytthonyx milleri</emphasis>
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</taxonomicName>
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can be separated from
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<taxonomicName id="79964D29E051A770FF6AF946FEEBED99" authorityName="Fanti and Damgaard" authorityYear="2019" box="[211,301,1687,1709]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
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<emphasis id="8CE2EAB8E051A770FF6AF946FEEBED99" box="[211,301,1687,1709]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
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</taxonomicName>
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by its smaller size (
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<quantity id="796E9B4FE051A770FDB4F949FD98ED99" box="[525,606,1687,1708]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="578" unit="mm" value="2.8">2.8 mm</quantity>
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in
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<taxonomicName id="79964D29E051A770FD14FF1CFCC3EBE2" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[685,773,194,215]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
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<emphasis id="8CE2EAB8E051A770FD14FF1CFCC3EBE2" box="[685,773,194,215]" italics="true" pageId="2" pageNumber="578">T. milleri</emphasis>
|
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</taxonomicName>
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and 6.0 mm in
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<taxonomicName id="79964D29E051A770FC24FF1CFC37EBE2" authorityName="Fanti and Damgaard" authorityYear="2019" box="[925,1009,194,215]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
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<emphasis id="8CE2EAB8E051A770FC24FF1CFC37EBE2" box="[925,1009,194,215]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
|
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</taxonomicName>
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) and the flavous color (
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<taxonomicName id="79964D29E051A770FD55FF03FC87EBC4" authorityName="Fanti and Damgaard" authorityYear="2019" box="[748,833,220,242]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
|
||
<emphasis id="8CE2EAB8E051A770FD55FF03FD3FEBC7" box="[748,761,221,242]" italics="true" pageId="2" pageNumber="578">T</emphasis>
|
||
.
|
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<emphasis id="8CE2EAB8E051A770FCBCFF02FC87EBC4" box="[773,833,220,241]" italics="true" pageId="2" pageNumber="578">stadili</emphasis>
|
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</taxonomicName>
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has a black face, elytra, venter, and basal eight antennomeres).
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<taxonomicName id="79964D29E051A770FC78FF29FBAEEA39" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[961,1128,247,268]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
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<emphasis id="8CE2EAB8E051A770FC78FF29FBAEEA39" box="[961,1128,247,268]" italics="true" pageId="2" pageNumber="578">Tytthonyx milleri</emphasis>
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</taxonomicName>
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has somewhat stronger serration of antennomeres 3–5 and a convex head (almost flat in
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<taxonomicName id="79964D29E051A770FC51FEF3FBFCEA74" authorityName="Fanti and Damgaard" authorityYear="2019" box="[1000,1082,300,322]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="stadili">
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<emphasis id="8CE2EAB8E051A770FC51FEF3FBFCEA74" box="[1000,1082,300,322]" italics="true" pageId="2" pageNumber="578">T. stadili</emphasis>
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</taxonomicName>
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although this character state is possibly an artifact of preservation) (Fanti and Damgaard 2019). From
|
||
<taxonomicName id="79964D29E051A770FB8EFEBCFB4BEA42" authorityName="Poinar and Fanti" authorityYear="2016" box="[1079,1165,354,375]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
|
||
<emphasis id="8CE2EAB8E051A770FB8EFEBCFB4BEA42" box="[1079,1165,354,375]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
|
||
</taxonomicName>
|
||
, which is similar in size, it is quite different (see Discussion below). The easiest way to distinguish them is by the length of the antenna, which is as long as the body in
|
||
<taxonomicName id="79964D29E051A770FCCBFE13FC0BEAD4" authorityName="Ivie, Fanti, and Ferreira" authorityYear="2022" box="[882,973,460,482]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="milleri">
|
||
<emphasis id="8CE2EAB8E051A770FCCBFE13FCB9EAD7" box="[882,895,461,482]" italics="true" pageId="2" pageNumber="578">T</emphasis>
|
||
.
|
||
<emphasis id="8CE2EAB8E051A770FC34FE12FC0BEAD4" box="[909,973,460,481]" italics="true" pageId="2" pageNumber="578">milleri</emphasis>
|
||
</taxonomicName>
|
||
and longer than the body in
|
||
<taxonomicName id="79964D29E051A770FD40FE36FC95EAC8" authorityName="Poinar and Fanti" authorityYear="2016" box="[761,851,487,509]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
|
||
<emphasis id="8CE2EAB8E051A770FD40FE36FCC0EAC8" box="[761,774,488,509]" italics="true" pageId="2" pageNumber="578">T</emphasis>
|
||
.
|
||
<emphasis id="8CE2EAB8E051A770FCABFE36FC95EAC8" box="[786,851,488,509]" italics="true" pageId="2" pageNumber="578">geiseri</emphasis>
|
||
</taxonomicName>
|
||
. The form of the pronotum is also distinctly different, being more rounded anterolaterally (square in
|
||
<taxonomicName id="79964D29E051A770FCF3FDC3FC67E907" authorityName="Poinar and Fanti" authorityYear="2016" box="[842,929,541,562]" class="Insecta" family="Cantharidae" genus="Tytthonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="geiseri">
|
||
<emphasis id="8CE2EAB8E051A770FCF3FDC3FC67E907" box="[842,929,541,562]" italics="true" pageId="2" pageNumber="578">T. geiseri</emphasis>
|
||
</taxonomicName>
|
||
) (Poinar and Fanti 2016). These characters are all potentially somewhat relat- ed to the different sexes of the
|
||
<typeStatus id="612D8808E051A770FC74FD8CFBC4E952" box="[973,1026,594,615]" pageId="2" pageNumber="578">types</typeStatus>
|
||
, but exceed the variation seen in most modern species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F68C6521E051A776FD7CFD59FDECEA4D" lastPageId="4" lastPageNumber="580" pageId="2" pageNumber="578" type="description">
|
||
<paragraph id="BE2936AAE051A776FD7CFD59FDECEA4D" blockId="2.[685,1173,194,1709]" lastBlockId="4.[152,638,194,376]" lastPageId="4" lastPageNumber="580" pageId="2" pageNumber="578">
|
||
<emphasis id="8CE2EAB8E051A770FD7CFD59FC84E9A8" bold="true" box="[709,834,647,669]" pageId="2" pageNumber="578">Description.</emphasis>
|
||
Female (based on the wide last abdominal ventrite), adult, alate, entirely flavous. Body length
|
||
<emphasis id="8CE2EAB8E051A770FC9FFD63FC85E9E7" box="[806,835,701,722]" italics="true" pageId="2" pageNumber="578">ca.</emphasis>
|
||
<quantity id="796E9B4FE051A770FCFEFD62FC54E9E7" box="[839,914,700,722]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="578" unit="mm" value="2.8">2.8 mm</quantity>
|
||
. Head completely exposed, elongate, narrowed behind the eyes, convex medially, pubescent and shallowly punctate. Eyes round, inserted laterally. Frontoclypeal region (face) constricted by antennal insertions near front of eyes, with a transverse depression between antennal insertions and a widened frontal margin, frontal margin nearly straight, slightly concave. Mandibles long, broad basal to the median strong retinaculum, thereafter long, thin, falciform. Maxillary palp 4-segmented; first palpomere robust, short; second elongate, cylindrical; third robust, intermediate in length between first two; fourth globular, apically pointed. Labial palp 3-segmented; terminal palpomere globular and apically pointed.Antenna with 11 antennomeres, serrate; relatively short, reaching
|
||
<emphasis id="8CE2EAB8E051A770FD14FBB6FD0DEF48" box="[685,715,1128,1149]" italics="true" pageId="2" pageNumber="578">ca.</emphasis>
|
||
midpoint of abdomen; scape stout, club-shaped; antennomere 2 filiform, about two-thirds length of first; antennomeres 3–8 (especially 4–7) serrate, subequal in length; antennomere 9 slightly serrate; antennomeres 10–11 filiform with terminal antennomere rounded at apex; all antennomeres pubescent. Pronotum transverse, surface slightly convex, pubescent, shallowly punctate, slightly wider than head, corners round, anterior and posterior margins and lateral beads strongly bordered, sides irregular and slightly enlarged near the anterior margin, lacking secretory pore. Elytra very short, reaching base of first ventrite; slightly dehiscent apically, apices round; setae short. Hind wings fully developed, surpassing last abdominal tergite; costal margin with series of dark bristles; radial cell open; r-m connected at both ends; Mr apparently narrowly erased at base; M
|
||
<subScript id="221234EFE051A770FCBCF9E6FCDBED73" attach="left" box="[773,797,1592,1606]" fontSize="6" pageId="2" pageNumber="578">3+4</subScript>
|
||
complete; cubital vein not visible. Ventrites transverse, pubescent, last ventrite wide. Legs long, slender; pro- and mesocoxae conical; trochanters elongate and offset, rounded apically; femora and tibiae cylindrical, straight; with small, fine apical spur on pro- and metatibiae, slightly stouter on mesotibia. Tarsal formula 5-5-5; metatarsus with first tarsomere elongate and thin, the second tarsomere about one-third length of first, third tarsomere small, fourth very short and strongly bilobed, fifth tarsomere slender, subequal to second; claws simple and minute. Male unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |