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<document id="B7B9E96DF5DCED4907BE5F41555C7DA4" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. &amp; Saxton, Natalie A." docDate="2025" docId="038987E8FFFE6E12FF4613CBFBADF801" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhytomus fervidus Saxton &amp; Genier 2025, new species" docType="treatment" docVersion="5" lastPageNumber="96" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="96" updateTime="1738747596743" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="8B0465BD1A37259D31D7C4485CB2E43E">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="D0BBE3E2BC9EA342A0BA9E902D3A2C45" type="personal">
<mods:role id="EC2A78EBC20281DF37B38F96E7759716">
<mods:roleTerm id="4077E12B3B7F9D4E108A94E365443CFC">Author</mods:roleTerm>
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<mods:namePart id="A35D7307F3FEB3E8183DE799A372FDE0">Génier, François</mods:namePart>
<mods:nameIdentifier id="8C7D242FC7900E4629223A2201BDF2FC" type="ORCID">0000-0002-5399-8412</mods:nameIdentifier>
<mods:nameIdentifier id="2B70E3A685BBF6F51CAB9FFFF3DB856E" type="email">enier@nature.ca</mods:nameIdentifier>
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<mods:name id="A6165EBF57216A1682C435E0E993DEC5" type="personal">
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</mods:role>
<mods:namePart id="FBF42E33407B9C455CF7DF86FFA06D8E">Gunter, Nicole L.</mods:namePart>
<mods:affiliation id="F1D646A0547EAE4944B54003F9CBCA2F">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:roleTerm id="6A0C9E7DA720F41945D984D083A9F753">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7702A5F3776D3C4BD7891521732398F1">Saxton, Natalie A.</mods:namePart>
<mods:affiliation id="1C8937A5F28FA1E4018D74EE0D272B93">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
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<mods:title id="6985FDDB91DB40019AD50CBF49FBF522">Zootaxa</mods:title>
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<mods:part id="9D05F40CD92402194882EF9A199D3CF0">
<mods:date id="22ADE816E4C0547B1ED5837ABA0A6C95">2025</mods:date>
<mods:detail id="3E548A927EC4A5BDD479A77213CF1C0B" type="pubDate">
<mods:number id="F0788408012B2F82AB272BFF29B20B66">2025-01-22</mods:number>
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<mods:number id="F2828DBD6E0D7776C2063325B86ED258">5575</mods:number>
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<mods:url id="7619D3075238C663173FEAE4292A2C17">https://doi.org/10.11646/zootaxa.5575.1.3</mods:url>
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<mods:classification id="42913A6833787107D0F7EA46C0A195A7">journal article</mods:classification>
<mods:identifier id="9106AB2ECE3A95BC631AD07158716C14" type="DOI">10.11646/zootaxa.5575.1.3</mods:identifier>
<mods:identifier id="5B3A262257FC5F684F302526B4C5BDC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="87CBF55B94CA86B8435892D95C695ED0" type="Zenodo-Dep">14746442</mods:identifier>
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<treatment id="038987E8FFFE6E12FF4613CBFBADF801" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801" httpUri="http://treatment.plazi.org/id/038987E8FFFE6E12FF4613CBFBADF801" lastPageNumber="96" pageId="9" pageNumber="96">
<subSubSection id="C33A6575FFFE6E12FF4613CBFD31FC58" box="[151,754,974,1001]" pageId="9" pageNumber="96" type="nomenclature">
<paragraph id="8B9F36FEFFFE6E12FF4613CBFD31FC58" blockId="9.[151,946,974,1073]" box="[151,754,974,1001]" pageId="9" pageNumber="96">
<heading id="D0D78192FFFE6E12FF4613CBFD31FC58" bold="true" box="[151,754,974,1001]" fontSize="11" level="1" pageId="9" pageNumber="96" reason="1">
<emphasis id="B954EAECFFFE6E12FF4613CBFD31FC58" bold="true" box="[151,754,974,1001]" pageId="9" pageNumber="96">
<taxonomicName id="4C204D7DFFFE6E12FF4613CBFD96FC59" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,597,974,1001]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="fervidus" status="sp. nov.">
<emphasis id="B954EAECFFFE6E12FF4613CBFEB8FC59" bold="true" box="[151,379,974,1001]" italics="true" pageId="9" pageNumber="96">Rhytomus fervidus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFE6E12FDB313CAFD31FC58" box="[610,754,974,1001]" pageId="9" pageNumber="96" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C33A6575FFFE6E12FF4613F7FDB4FB81" pageId="9" pageNumber="96" type="description">
<paragraph id="8B9F36FEFFFE6E12FF4613F7FC71FBBD" blockId="9.[151,946,974,1073]" box="[151,946,1010,1037]" pageId="9" pageNumber="96">
<uri id="FFB13AFCFFFE6E12FF4613F7FC71FBBD" box="[151,946,1010,1037]" pageId="9" pageNumber="96">
urn:lsid:zoobank.org:act:
<uuid id="FF860C2BFFFE6E12FE7613F7FC71FBBD" box="[423,946,1010,1037]" pageId="9" pageNumber="96">27C09598-572B-4AF0-904F-59067A688C08</uuid>
</uri>
</paragraph>
<paragraph id="8B9F36FEFFFE6E12FF461412FDB4FB81" blockId="9.[151,946,974,1073]" box="[151,631,1047,1073]" pageId="9" pageNumber="96">
<figureCitation id="131B2A7BFFFE6E12FF461412FF3BFB81" box="[151,248,1047,1073]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="9" pageNumber="96">Figs. 4B</figureCitation>
,
<figureCitation id="131B2A7BFFFE6E12FED51412FEEAFB81" box="[260,297,1047,1073]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="9" pageNumber="96">7A</figureCitation>
,
<figureCitation id="131B2A7BFFFE6E12FEE51412FE9BFB81" box="[308,344,1047,1073]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="9" pageNumber="96">8A</figureCitation>
,
<figureCitation id="131B2A7BFFFE6E12FEB41412FE44FB81" box="[357,391,1047,1073]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="9" pageNumber="96">9B</figureCitation>
,
<figureCitation id="131B2A7BFFFE6E12FE421412FDFFFB81" box="[403,572,1047,1073]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="9" pageNumber="96">11A, 11D, 11E</figureCitation>
,
<figureCitation id="131B2A7BFFFE6E12FD991412FDB4FB81" box="[584,631,1047,1073]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="9" pageNumber="96">12B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFE6E12FF46145FFA58FAFD" pageId="9" pageNumber="96" type="materials_examined">
<paragraph id="8B9F36FEFFFE6E12FF46145FFEDDFB51" blockId="9.[151,1437,1114,1969]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF46145FFE7DFBC5" bold="true" box="[151,446,1114,1141]" pageId="9" pageNumber="96">Name-bearing type data.</emphasis>
<materialsCitation id="3B483CA3FFFE6E12FE15145EFED8FB51" collectedFrom="oak forest dung" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" determinerName=". N. Saxton &amp; F. Genier" elevation="6000" location="Canadian Museum" longitude="144.23334" pageId="9" pageNumber="96" specimenCode="CMNEN 00049283" specimenCount="1" specimenCount-male="1" stateProvince="Western Highlands" typeStatus="holotype">
<typeStatus id="549B885CFFFE6E12FE15145EFDEFFBC5" box="[452,556,1115,1141]" pageId="9" pageNumber="96" type="holotype">Holotype</typeStatus>
<specimenCount id="9D26FD77FFFE6E12FDE2145EFD8BFBC5" box="[563,584,1115,1141]" count="1" pageId="9" pageNumber="96" type="male">♂</specimenCount>
(
<collectionCode id="ED31AE3BFFFE6E12FD87145EFD73FBC5" box="[598,688,1115,1141]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="9" pageNumber="96">CMNC</collectionCode>
): || PAPUA NEW GUINEA | Western highlands, Mt. | Hagen, 6000’ |
<date id="FF9E103EFFFE6E12FF74147BFEF9FB29" box="[165,314,1150,1177]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">
<collectingDate id="EFDAE9D6FFFE6E12FF74147BFEF9FB29" box="[165,314,1150,1177]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">5-8.VII.1974</collectingDate>
</date>
,
<collectorName id="26D55328FFFE6E12FE97147AFE57FB29" box="[326,404,1151,1177]" pageId="9" pageNumber="96">S.Peck</collectorName>
|
<collectedFrom id="383C7503FFFE6E12FE7B147AFD99FB29" box="[426,602,1151,1177]" pageId="9" pageNumber="96">oak forest dung</collectedFrom>
tps. 24-25 ||; ||
<location id="8EFF6025FFFE6E12FCDA147AFC1CFB29" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FCDA147AFC1CFB29" box="[779,991,1151,1177]" name="Canadian Museum" pageId="9" pageNumber="96">Canadian Museum</location>
of |
<location id="8EFF6025FFFE6E12FBC5147AFB9CFB29" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FBC5147AFB9CFB29" box="[1044,1119,1151,1177]" name="Musee" pageId="9" pageNumber="96">Musée</location>
canadien de la | NATURE |
<specimenCode id="DB869E85FFFE6E12FF4614A6FEB7FB0D" box="[151,372,1187,1213]" pageId="9" pageNumber="96">CMNEN 00049283</specimenCode>
|| [QR code label]; || HOLOTYPE ♂ |
<taxonomicName id="4C204D7DFFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" italics="true" pageId="9" pageNumber="96">
<location id="8EFF6025FFFE6E12FCFA14A6FC5AFB0D" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" name="Rhytomus" pageId="9" pageNumber="96">Rhytomus</location>
</emphasis>
</taxonomicName>
|
<taxonomicName id="4C204D7DFFFE6E12FC7E14A6FBCAFB0D" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[943,1033,1187,1213]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="fervidus">
<emphasis id="B954EAECFFFE6E12FC7E14A6FBCAFB0D" box="[943,1033,1187,1213]" italics="true" pageId="9" pageNumber="96">fervidus</emphasis>
</taxonomicName>
| Des
<determinerName id="7E43DA3AFFFE6E12FB9C14A6FB0AFB0D" box="[1101,1225,1187,1213]" pageId="9" pageNumber="96">. N. Saxton</determinerName>
&amp;
<determinerName id="7E43DA3AFFFE6E12FB3F14A6FA9BFB0D" box="[1262,1368,1187,1213]" pageId="9" pageNumber="96">F. Génier</determinerName>
, 2024 || [red card]
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F36FEFFFE6E12FF1614EFFA58FAFD" blockId="9.[151,1437,1114,1969]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF1614EFFE6CFAB4" bold="true" box="[199,431,1258,1284]" pageId="9" pageNumber="96">Material examined</emphasis>
(
<specimenCount id="9D26FD77FFFE6E12FE1514EEFDD9FAB5" box="[452,538,1259,1285]" count="20" pageId="9" pageNumber="96" type="male">20 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFFE6E12FDFD14EEFDADFAB5" box="[556,622,1259,1285]" count="9" pageId="9" pageNumber="96" type="female">9 ♀♀</specimenCount>
).
<emphasis id="B954EAECFFFE6E12FD5C14EFFC77FAB4" bold="true" box="[653,948,1258,1285]" pageId="9" pageNumber="96">
<collectingCountry id="F337766EFFFE6E12FD5C14EFFC77FAB4" box="[653,948,1258,1285]" name="Papua New Guinea" pageId="9" pageNumber="96">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<collectingRegion id="49E4F81CFFFE6E12FC1914EEFA5BFAB5" box="[968,1432,1259,1285]" country="Papua New Guinea" name="Western Highlands" pageId="9" pageNumber="96">WESTERN HIGHLANDS PROVINCE</collectingRegion>
,
<location id="8EFF6025FFFE6E12FF46150AFEF6FA99" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FF46150AFEF6FA99" box="[151,309,1295,1321]" country="Papua New Guinea" name="Mount Hagen" pageId="9" pageNumber="96" stateProvince="Western Highlands">Mount Hagen</location>
,
<quantity id="4CD89B1BFFFE6E12FE91150AFE5BFA99" box="[320,408,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.829" pageId="9" pageNumber="96" unit="m" value="1829.0">
<elevation id="000DD1CDFFFE6E12FE91150AFE5BFA99" box="[320,408,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.829" pageId="9" pageNumber="96" unit="m" value="1829.0">1829 m</elevation>
</quantity>
, [
<geoCoordinate id="EE145039FFFE6E12FE7A150BFE38FA99" box="[427,507,1294,1321]" degrees="05" direction="south" minutes="52" orientation="latitude" pageId="9" pageNumber="96" precision="925" value="-5.866667">05°52’</geoCoordinate>
<collectingRegion id="49E4F81CFFFE6E12FE2A150BFDC8FA99" box="[507,523,1294,1321]" country="Papua New Guinea" name="S" pageId="9" pageNumber="96">S</collectingRegion>
,
<geoCoordinate id="EE145039FFFE6E12FDC6150AFD44FA99" box="[535,647,1295,1321]" degrees="144" direction="east" minutes="14" orientation="longitude" pageId="9" pageNumber="96" precision="925" value="144.23334">144°14’E</geoCoordinate>
],
<date id="FF9E103EFFFE6E12FD4B150BFCE9FA99" box="[666,810,1294,1321]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">
<collectingDate id="EFDAE9D6FFFE6E12FD4B150BFCE9FA99" box="[666,810,1294,1321]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">5–8.vii.1974</collectingDate>
</date>
, oak forest,
<collectingMethod id="52614EE9FFFE6E12FC65150AFBEFFA99" box="[948,1068,1295,1321]" pageId="9" pageNumber="96">dung traps</collectingMethod>
,
<collectorName id="26D55328FFFE6E12FBE9150AFB4CFA99" box="[1080,1167,1295,1321]" pageId="9" pageNumber="96">S. Peck</collectorName>
, (
<collectingMethod id="52614EE9FFFE6E12FB73150AFADAFA99" box="[1186,1305,1295,1321]" pageId="9" pageNumber="96">dung traps</collectingMethod>
24-25)—
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<typeStatus id="549B885CFFFE6E12FF461536FF3FFAFD" box="[151,252,1331,1357]" pageId="9" pageNumber="96" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D26FD77FFFE6E12FED61536FEDFFAFD" box="[263,284,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">♀</specimenCount>
<typeStatus id="549B885CFFFE6E12FEF21536FE43FAFD" box="[291,384,1331,1357]" pageId="9" pageNumber="96" type="allotype">allotype</typeStatus>
,
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<specimenCount id="9D26FD77FFFE6E12FE5A1536FE19FAFD" box="[395,474,1331,1357]" count="12" pageId="9" pageNumber="96" type="male">12 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFFE6E12FE371536FDE6FAFD" box="[486,549,1331,1357]" count="6" pageId="9" pageNumber="96" type="female">6 ♀♀</specimenCount>
<typeStatus id="549B885CFFFE6E12FDFD1536FD54FAFD" box="[556,663,1331,1357]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFFE6E12FD761536FD3CFAFD" box="[679,767,1331,1357]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="9" pageNumber="96">CMNC</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B483CA3FFFE6E12FCC31536FBAEFAFD" box="[786,1133,1331,1357]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" elevation="1829" location="Mount Hagen" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="S" typeStatus="paratype">
<specimenCount id="9D26FD77FFFE6E12FCC31536FC90FAFD" box="[786,851,1331,1357]" count="6" pageId="9" pageNumber="96" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFFE6E12FC8E1536FC4AFAFD" box="[863,905,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFFE6E12FC411536FC38FAFD" box="[912,1019,1331,1357]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFFE6E12FBDB1536FBA6FAFD" box="[1034,1125,1331,1357]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="96">MNHN</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B483CA3FFFE6E12FBA91536FA55FAFD" box="[1144,1430,1331,1357]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="QM" collectorName="S. Peck" country="Papua New Guinea" elevation="1829" location="Mount Hagen" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="S" typeStatus="paratype">
<specimenCount id="9D26FD77FFFE6E12FBA91536FB67FAFD" box="[1144,1188,1331,1357]" count="1" pageId="9" pageNumber="96" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFFE6E12FB611536FB19FAFD" box="[1200,1242,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">1 ♀</specimenCount>
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(
<collectionCode id="ED31AE3BFFFE6E12FA8D1536FA4CFAFD" box="[1372,1423,1331,1357]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="9" pageNumber="96" type="Museum">QM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFE6E12FF161553FA91F8D9" pageId="9" pageNumber="96" type="description">
<paragraph id="8B9F36FEFFFE6E12FF161553FA91F8D9" blockId="9.[151,1437,1114,1969]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF161553FDDBFAC1" bold="true" box="[199,536,1366,1393]" pageId="9" pageNumber="96">Description. Measurements.</emphasis>
Body length 4.2–5.0 mm, maximum body width
<quantity id="4CD89B1BFFFE6E12FB821552FB1CFAC1" box="[1107,1247,1367,1393]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="9" pageNumber="96" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6–3.1 mm</quantity>
.
<typeStatus id="549B885CFFFE6E12FB3A1552FA9BFAC1" box="[1259,1368,1367,1393]" pageId="9" pageNumber="96" type="holotype">Holotype</typeStatus>
: body length
<quantity id="4CD89B1BFFFE6E12FF32157EFEF9FA25" box="[227,314,1403,1429]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="9" pageNumber="96" unit="mm" value="4.7">4.7 mm</quantity>
, maximum body width 3.0 mm.
<emphasis id="B954EAECFFFE6E12FD71157EFD27FA25" bold="true" box="[672,740,1403,1429]" pageId="9" pageNumber="96">Body.</emphasis>
Overall shape in dorsal view oval, elytral edge evenly rounded from base to apex.
<emphasis id="B954EAECFFFE6E12FEAA159AFE01FA09" bold="true" box="[379,450,1439,1465]" pageId="9" pageNumber="96">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination (
<figureCitation id="131B2A7BFFFE6E12FBAD159AFB1BFA09" box="[1148,1240,1439,1465]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="9" pageNumber="96">Fig. 4C</figureCitation>
), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width.
<emphasis id="B954EAECFFFE6E12FE2315E2FDB1F9B1" bold="true" box="[498,626,1511,1537]" pageId="9" pageNumber="96">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFFE6E12FE7A162AFE3DF9F9" bold="true" box="[427,510,1583,1609]" pageId="9" pageNumber="96">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (
<figureCitation id="131B2A7BFFFE6E12FDD81672FDAEF921" box="[521,621,1655,1681]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="9" pageNumber="96">Fig. 12B</figureCitation>
), basal elytral pit slightly wider than stria 10 and shallow (
<figureCitation id="131B2A7BFFFE6E12FB3A1672FA93F921" box="[1259,1360,1655,1681]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="9" pageNumber="96">Fig. 11D</figureCitation>
), basal surface of pseudepipleuron with irregularly distributed more or less defined punctures (
<figureCitation id="131B2A7BFFFE6E12FBB2169EFB06F905" box="[1123,1221,1691,1717]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="9" pageNumber="96">Fig. 11E</figureCitation>
), interstriae 3–5 on basal fifth with scattered unaligned punctures.
<emphasis id="B954EAECFFFE6E12FD7316BAFD22F969" bold="true" box="[674,737,1727,1753]" pageId="9" pageNumber="96">Legs.</emphasis>
Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge.
<emphasis id="B954EAECFFFE6E12FCD816E7FC40F94C" bold="true" box="[777,899,1762,1788]" pageId="9" pageNumber="96">Pygidium.</emphasis>
Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (
<figureCitation id="131B2A7BFFFE6E12FEAE172EFE1FF8F5" box="[383,476,1835,1861]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="9" pageNumber="96">Fig. 8A</figureCitation>
), pygidium of female depressed apically.
<emphasis id="B954EAECFFFE6E12FC15172FFBFDF8F4" bold="true" box="[964,1086,1834,1860]" pageId="9" pageNumber="96">Aedeagus.</emphasis>
Left paramere apex truncated apically, distinctly angular externally (
<figureCitation id="131B2A7BFFFE6E12FD94174AFD5DF8D9" box="[581,670,1871,1897]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="9" pageNumber="96">Fig. 9B</figureCitation>
).
<emphasis id="B954EAECFFFE6E12FD63174AFCBDF8D8" bold="true" box="[690,894,1870,1897]" pageId="9" pageNumber="96">Female genitalia.</emphasis>
Spermatheca with two apodemes basally.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFE6E12FF161776FBADF801" pageId="9" pageNumber="96" type="etymology">
<paragraph id="8B9F36FEFFFE6E12FF161776FBADF801" blockId="9.[151,1437,1114,1969]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF161776FE8DF83D" bold="true" box="[199,334,1907,1933]" pageId="9" pageNumber="96">Etymology.</emphasis>
The specific epithet for this species is the masculine nominative Latin adjective meaning “boiling hot” or “fiery” in reference to the
<typeStatus id="549B885CFFFE6E12FDC21792FD80F801" box="[531,579,1943,1969]" pageId="9" pageNumber="96">type</typeStatus>
locality, which is near an old composite volcano.
</paragraph>
</subSubSection>
</treatment>
</document>
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