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<document id="168BBDA11B2EAE9C879A06736258CD61" ID-DOI="10.11646/zootaxa.5575.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="14746442" ID-ZooBank="521D404D-EB9D-4110-989E-A8A273C10152" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1737970514015" checkinUser="plazi" docAuthor="Génier, François, Gunter, Nicole L. &amp; Saxton, Natalie A." docDate="2025" docId="038987E8FFE36E0FFF461092FA5FFAAF" docLanguage="en" docName="zootaxa.5575.1.3.pdf" docOrigin="Zootaxa 5575 (1)" docSource="https://doi.org/10.11646/zootaxa.5575.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhytomus" docType="key" docVersion="5" lastPageNumber="107" masterDocId="FFB0FF90FFF76E1BFFD11005FFC3FFB0" masterDocTitle="Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="110" masterPageNumber="87" pageNumber="107" updateTime="1738747596743" updateUser="carolina" zenodo-license-document="CLOSED">
<mods:mods id="A62F57246FF40F4B7268B25830EBD1D5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E99BBC6040CDAF20B26DE98A0F324325">
<mods:title id="3307EC5B050988D5A98C35260CE0F8C1">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="66F68EFDD14D495A9FA5F920EF0D0FFA" type="personal">
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<mods:namePart id="3D2873FDB0AF57F197E373D56F1669FE">Génier, François</mods:namePart>
<mods:nameIdentifier id="DAD318F4B1A24A72B23AAF30C67F4173" type="ORCID">0000-0002-5399-8412</mods:nameIdentifier>
<mods:nameIdentifier id="07DB640D399297C583EA4F6705C3D19A" type="email">enier@nature.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="C0EFEC4D8A47D832BEF9AB51C4795813" type="personal">
<mods:role id="624B7460B09DD8AE3ED0DF1A55E2925F">
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<mods:namePart id="A55F49E86EC08DAF3089371FDB1AD2C7">Gunter, Nicole L.</mods:namePart>
<mods:affiliation id="60C7FB99B72B97D1CA5AD4AF0AD1E4C5">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:name id="8B6158A6C9496CA5BDD7E1AC7F06ABEA" type="personal">
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</mods:role>
<mods:namePart id="6FAFF13AB52B9B5013733EF15440AFD1">Saxton, Natalie A.</mods:namePart>
<mods:affiliation id="8719A3DCED6BEF182AEEBFB2DA3008D6">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
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<mods:title id="734D0F199E7364DBE88E3507370C90A4">Zootaxa</mods:title>
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<mods:date id="684265308A660FDD4E9CF638D0CBF29F">2025</mods:date>
<mods:detail id="9107D9DDAB61357607605693CC55730F" type="pubDate">
<mods:number id="236760A512058D51C8B85C7908692E2F">2025-01-22</mods:number>
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<mods:number id="00F99897F4C5129BA50FA487DD51306E">5575</mods:number>
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<mods:identifier id="28FB74E6A795FF4ABC06AE396A3215A7" type="DOI">10.11646/zootaxa.5575.1.3</mods:identifier>
<mods:identifier id="C3952174426E1E80416C90618F4C4B77" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038987E8FFE36E0FFF461092FA5FFAAF" LSID="urn:lsid:plazi:treatment:038987E8FFE36E0FFF461092FA5FFAAF" httpUri="http://treatment.plazi.org/id/038987E8FFE36E0FFF461092FA5FFAAF" lastPageNumber="107" pageId="20" pageNumber="107">
<subSubSection id="C33A6575FFE36E0FFF461092FCF2FF01" box="[151,817,151,178]" pageId="20" pageNumber="107" type="nomenclature">
<paragraph id="8B9F36FEFFE36E0FFF461092FCF2FF01" blockId="20.[151,1436,151,250]" box="[151,817,151,178]" pageId="20" pageNumber="107">
<heading id="D0D78192FFE36E0FFF461092FCF2FF01" bold="true" box="[151,817,151,178]" fontSize="11" level="1" pageId="20" pageNumber="107" reason="1">
<emphasis id="B954EAECFFE36E0FFF461092FCF2FF01" bold="true" box="[151,817,151,178]" pageId="20" pageNumber="107">
Key to adult mainland New
<collectingCountry id="F337766EFFE36E0FFE271092FD92FF01" box="[502,593,151,177]" name="Guinea-Bissau" pageId="20" pageNumber="107">Guinea</collectingCountry>
<taxonomicName id="4C204D7DFFE36E0FFD881092FD10FF02" box="[601,723,151,178]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFE36E0FFD881092FD10FF02" bold="true" box="[601,723,151,178]" italics="true" pageId="20" pageNumber="107">Rhytomus</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE36E0FFF4610BEFE7BFF49" pageId="20" pageNumber="107" type="multiple">
<paragraph id="8B9F36FEFFE36E0FFF4610BEFE7BFF49" blockId="20.[151,1436,151,250]" pageId="20" pageNumber="107">
<emphasis id="B954EAECFFE36E0FFF4610BEFE50FF65" bold="true" box="[151,403,187,213]" pageId="20" pageNumber="107">Access to online key:</emphasis>
https://keys.lucidcentral.org/keys/v4/rhytomus/ (click on the magic wand icon to select the best character to use first)
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE36E0FFF461121FA5FFAAF" pageId="20" pageNumber="107" type="key">
<key id="FFB161CFFFE36E0FFF461121FA5FFAAF" pageId="20" pageNumber="107">
<keyStep id="30D42E5BFFE36E0FFF461121FA5FFE22" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF461121FA5FFEE7" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461121FA5FFEE7" pageId="20" pageNumber="107">
1 Interstriae 3–5 on basal fifth all having unaligned scattered punctures (
<figureCitation id="131B2A7BFFE36E0FFCAE1121FC15FE89" box="[895,982,292,314]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11A</figureCitation>
); basal pseudepipleural pit present, shallow and slightly larger than stria 10 (
<figureCitation id="131B2A7BFFE36E0FFE201144FD84FEE7" box="[497,583,321,343]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11D</figureCitation>
).................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF46115AFA5FFE22" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF46115AFA5FFE22" pageId="20" pageNumber="107">
- Interstriae 3–5 with at least one interval having two distinct rows of punctures on basal fifth (11B); basal pseudepipleural pit deep and distinctly larger than stria 10 (
<figureCitation id="131B2A7BFFE36E0FFD8F1179FD70FE21" box="[606,691,380,402]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11C</figureCitation>
) OR pseudepipleural pit is absent................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF46119CFA5FFDB7" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF46119CFA5FFE7C" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF46119CFA5FFE7C" pageId="20" pageNumber="107">2 Elytral interval 1 with basal punctures much smaller in size to those in intervals 2–3; pronotal lateral edge obsolete or sharply defined............................................................................................. 3</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF4611D1FA5FFDB7" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4611D1FA5FFDB7" pageId="20" pageNumber="107">- Elytral interval 1 with basal punctures similar in size and coarseness to those in intervals 2–3; pronotal lateral edge always sharply defined...................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF46120AFA5FFDCD" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF46120AFA5FFDF2" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF46120AFA5FFDF2" pageId="20" pageNumber="107">
3 Pronotal lateral edge obsolete; male clypeal edge without emargination lateral to clypeal teeth (
<figureCitation id="131B2A7BFFE36E0FFB56120AFB17FD94" box="[1159,1236,527,549]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="20" pageNumber="107">Fig. 4B</figureCitation>
); Body length
<quantity id="4CD89B1BFFE36E0FFAB7120AFEC4FDF2" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="5.3" metricValueMin="4.0" pageId="20" pageNumber="107" unit="mm" value="4.65" valueMax="5.3" valueMin="4.0">4–5.3 mm</quantity>
....................................................................
<taxonomicName id="4C204D7DFFE36E0FFB911229FA5FFDF2" authority="(Paulian, 1985)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1985" box="[1088,1436,556,578]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B954EAECFFE36E0FFB911229FAC1FDF2" box="[1088,1282,556,578]" italics="true" pageId="20" pageNumber="107">Rhytomus papuanus</emphasis>
(
<bibRefCitation id="EFB14B0FFFE36E0FFADE1229FA56FDF2" author="Paulian, R." box="[1295,1429,556,578]" pageId="20" pageNumber="107" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF46124FFA5FFDCD" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF46124FFA5FFDCD" pageId="20" pageNumber="107">
- Pronotal lateral edge sharply defined; male clypeal edge with emargination lateral to clypeal teeth (
<figureCitation id="131B2A7BFFE36E0FFB6E124CFAD3FDEE" box="[1215,1296,585,607]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="20" pageNumber="107">Fig. 4A</figureCitation>
); Body length
<quantity id="4CD89B1BFFE36E0FFF331262FE96FDCC" box="[226,341,615,637]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.1" metricValueMin="3.6" pageId="20" pageNumber="107" unit="mm" value="3.8499999999999996" valueMax="4.1" valueMin="3.6">3.6–4.1 mm</quantity>
..............................................................
<taxonomicName id="4C204D7DFFE36E0FFB9D1262FA5FFDCD" authority="(Paulian, 1985)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1985" box="[1100,1436,615,637]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B954EAECFFE36E0FFB9D1262FAC1FDCC" box="[1100,1282,615,636]" italics="true" pageId="20" pageNumber="107">Rhytomus howdeni</emphasis>
(
<bibRefCitation id="EFB14B0FFFE36E0FFADE1262FA56FDCD" author="Paulian, R." box="[1295,1429,615,637]" pageId="20" pageNumber="107" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF461281FA5FFD41" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF461281FA5FFD07" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461281FA5FFD07" pageId="20" pageNumber="107">
4 Basal pseudepipleural surface with irregularly distributed more or less defined punctures (
<figureCitation id="131B2A7BFFE36E0FFBE21281FB45FD29" box="[1075,1158,644,666]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11E</figureCitation>
); female pygidium depressed apically......................................................
<taxonomicName id="4C204D7DFFE36E0FFC1212A4FAE3FD07" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[963,1312,673,695]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="fervidus" status="sp. nov.">
<emphasis id="B954EAECFFE36E0FFC1212A4FBB6FD06" box="[963,1141,673,694]" italics="true" pageId="20" pageNumber="107">Rhytomus fervidus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFE36E0FFAF812A7FA5FFD07" box="[1321,1436,673,695]" pageId="20" pageNumber="107" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF4612BAFA5FFD41" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4612BAFA5FFD41" pageId="20" pageNumber="107">
- Basal pseudepipleural surface with sharply defined ocellate punctures (
<figureCitation id="131B2A7BFFE36E0FFC5C12BAFC22FD64" box="[909,993,703,725]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11F</figureCitation>
); female pygidium evenly convex..................................................................................
<taxonomicName id="4C204D7DFFE36E0FFB8E12D9FA5FFD41" authority="(Paulian, 1937)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1937" box="[1119,1436,732,754]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="heurni">
<emphasis id="B954EAECFFE36E0FFB8E12D9FAC0FD41" box="[1119,1283,732,753]" italics="true" pageId="20" pageNumber="107">Rhytomus heurni</emphasis>
(
<bibRefCitation id="EFB14B0FFFE36E0FFADE12D9FA56FD41" author="Paulian, R." box="[1295,1429,732,754]" pageId="20" pageNumber="107" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian, 1937</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF4612FCFA5FFCD7" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF4612FCFA5FFC9D" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4612FCFA5FFC9D" pageId="20" pageNumber="107">
5 Base of pronotum lacking fine longitudinal depression medially; basal pseudepipleural pit deep and distinctly larger than stria 10 (
<figureCitation id="131B2A7BFFE36E0FFED61312FE9EFC9C" box="[263,349,791,813]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11C</figureCitation>
); male pygidial groove not distinctly narrowed medially (
<figureCitation id="131B2A7BFFE36E0FFC561312FBCEFC9C" box="[903,1037,791,813]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="20" pageNumber="107">Figs. 8D, H, I</figureCitation>
)............................... 6
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF461331FA5FFCD7" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461331FA5FFCD7" pageId="20" pageNumber="107">
- Base of pronotum with fine longitudinal depression medially (Fig. E); basal pseudepipleural pit absent; male pygidial groove distinctly narrowed medially and terminating in pits (
<figureCitation id="131B2A7BFFE36E0FFD331354FCEAFCD7" box="[738,809,849,871]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="20" pageNumber="107">Fig. 8E</figureCitation>
)........
<taxonomicName id="4C204D7DFFE36E0FFC4E1354FAE3FCD7" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[927,1312,849,871]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="latidentatus" status="sp. nov.">
<emphasis id="B954EAECFFE36E0FFC4E1354FBB6FCD6" box="[927,1141,849,870]" italics="true" pageId="20" pageNumber="107">Rhytomus latidentatus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFE36E0FFAF81357FA5FFCD7" box="[1321,1436,849,871]" pageId="20" pageNumber="107" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<keyStep id="30D42E5BFFE36E0FFF46136AFA5FFC0F" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF46136AFA5FFC34" blockId="20.[151,1437,292,1311]" box="[151,1436,879,901]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF46136AFA5FFC34" box="[151,1436,879,901]" pageId="20" pageNumber="107">
6 Interstriae 3, 4 and 5 having two distinct rows of punctures; elytral pubescence long and conspicuous (
<figureCitation id="131B2A7BFFE36E0FFB68136AFACDFC34" box="[1209,1294,879,901]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11B</figureCitation>
)......... 7
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF461389FA5FFC0F" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461389FA5FFC0F" pageId="20" pageNumber="107">
- Interstriae 3 and 5 having unaligned and scattered punctures, interstriae 4 having two distinct rows of punctures; elytral pubescence short and fine (as in
<figureCitation id="131B2A7BFFE36E0FFDCA13ACFDB3FC0F" box="[539,624,937,959]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">Fig. 11A</figureCitation>
)................................................................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF4613C2FA5FFB84" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF4613C2FA5FFC4A" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4613C2FA5FFC4A" pageId="20" pageNumber="107">
<specimenCount id="9D26FD77FFE36E0FFF4613C2FED0FC6D" box="[151,275,967,989]" count="7" pageId="20" pageNumber="107" type="male">7 Male</specimenCount>
metatibial apicointernal tooth approximately in line with posterior tibial edge (
<figureCitation id="131B2A7BFFE36E0FFBDE13C2FBA4FC6C" box="[1039,1127,967,989]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12C</figureCitation>
); pygidium of female depressed basally; body length 3.0–
<quantity id="4CD89B1BFFE36E0FFE0513E1FDDDFC4A" box="[468,542,996,1018]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="20" pageNumber="107" unit="mm" value="4.6">4.6 mm</quantity>
..................................
<taxonomicName id="4C204D7DFFE36E0FFC1C13E1FAE3FC4A" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[973,1312,996,1018]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="sodalis" status="sp. nov.">
<emphasis id="B954EAECFFE36E0FFC1C13E1FBB6FC49" box="[973,1141,996,1017]" italics="true" pageId="20" pageNumber="107">Rhytomus sodalis</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFE36E0FFAF813E0FA5FFC4A" box="[1321,1436,996,1018]" pageId="20" pageNumber="107" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF461407FA5FFB84" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461407FA5FFB84" pageId="20" pageNumber="107">
- Male metatibial apicointernal tooth set anteriorly to posterior tibial edge (
<figureCitation id="131B2A7BFFE36E0FFC741404FC3DFBA6" box="[933,1022,1025,1047]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12D</figureCitation>
); pygidium of female evenly convex: body length 4.1–6.0 mm.................................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF461439FA5FFB1A" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF461439FA5FFBDF" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461439FA5FFBDF" pageId="20" pageNumber="107">
8 Elytral surface between punctures dull, with distinct microsculptures (
<figureCitation id="131B2A7BFFE36E0FFC521439FC0CFBE1" box="[899,975,1084,1106]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="20" pageNumber="107">Fig. 7D</figureCitation>
,
<figureCitation id="131B2A7BFFE36E0FFC081439FBC0FBE1" box="[985,1027,1084,1105]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="20" pageNumber="107">11B</figureCitation>
); Body length
<quantity id="4CD89B1BFFE36E0FFB401439FAC7FBE1" box="[1169,1284,1084,1106]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.4" metricValueMin="4.1" pageId="20" pageNumber="107" unit="mm" value="4.75" valueMax="5.4" valueMin="4.1">4.1–5.4 mm</quantity>
.........................................................................
<taxonomicName id="4C204D7DFFE36E0FFC1F145CFAE3FBDF" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[974,1312,1113,1135]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="larseni" status="sp. nov.">
<emphasis id="B954EAECFFE36E0FFC1F145CFBB5FBDE" box="[974,1142,1113,1134]" italics="true" pageId="20" pageNumber="107">Rhytomus larseni</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFE36E0FFAF8145FFA5FFBDF" box="[1321,1436,1113,1135]" pageId="20" pageNumber="107" rank="species">new species</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF461472FA5FFB1A" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF461472FA5FFB1A" pageId="20" pageNumber="107">
- Elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (
<figureCitation id="131B2A7BFFE36E0FFBB61472FB76FB3D" box="[1127,1205,1143,1165]" captionStart="FIGURE 6" captionStartId="8.[152,255,2017,2041]" captionTargetBox="[151,1436,1071,1991]" captionTargetId="figure-192@8.[151,1436,1071,1991]" captionTargetPageId="8" captionText="FIGURE 6. Lectotype of Rhytomus elongatus. A, dorsal habitus; B, dorsal head; C, label data." figureDoi="http://doi.org/10.5281/zenodo.14746454" httpUri="https://zenodo.org/record/14746454/files/figure.png" pageId="20" pageNumber="107">Fig. 6A</figureCitation>
); Body length
<quantity id="4CD89B1BFFE36E0FFA931472FAB0FB3D" box="[1346,1395,1143,1165]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="107" unit="mm" value="6.0">6mm</quantity>
...................................................................
<taxonomicName id="4C204D7DFFE36E0FFC2D1491FA5FFB1A" authority="(van Lansberge, 1885)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="van Lansberge" baseAuthorityYear="1885" box="[1020,1436,1172,1194]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B954EAECFFE36E0FFC2D1491FB7DFB19" box="[1020,1214,1172,1193]" italics="true" pageId="20" pageNumber="107">Rhytomus elongatus</emphasis>
(
<bibRefCitation id="EFB14B0FFFE36E0FFB1A1491FA56FB1A" author="Van Lansberge, J. W." box="[1227,1429,1172,1194]" pageId="20" pageNumber="107" pagination="375 - 400" refId="ref16891" refString="Van Lansberge, J. W. (1885) Description d'especes nouvelles de coleopteres appartenant au Musee civique de Genes. Annali del Museo civico di storia naturale di Genova, 2 (2), 375 - 400." type="journal article" year="1885">van Lansberge, 1885</bibRefCitation>
)
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30D42E5BFFE36E0FFF4614B4FA5FFAAF" pageId="20" pageNumber="107">
<paragraph id="8B9F36FEFFE36E0FFF4614B4FA5FFB55" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4614B4FA5FFB55" pageId="20" pageNumber="107">
9 Elytral surface appearing glossy, at most with ill-defined microsculpture (as in
<figureCitation id="131B2A7BFFE36E0FFC1E14B4FBE6FB76" box="[975,1061,1201,1223]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12B</figureCitation>
); Elytra strongly convex in lateral view (
<figureCitation id="131B2A7BFFE36E0FFF3814CAFEFDFB54" box="[233,318,1231,1253]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12E</figureCitation>
)..............................................................
<taxonomicName id="4C204D7DFFE36E0FFBE514CAFA5FFB55" authority="(Balthasar, 1965)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Balthasar" baseAuthorityYear="1965" box="[1076,1436,1231,1253]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="setulosus">
<emphasis id="B954EAECFFE36E0FFBE514CAFB32FB54" box="[1076,1265,1231,1252]" italics="true" pageId="20" pageNumber="107">Rhytomus setulosus</emphasis>
(
<bibRefCitation id="EFB14B0FFFE36E0FFB2C14CAFA56FB55" author="Balthasar, V." box="[1277,1429,1231,1253]" pageId="20" pageNumber="107" pagination="14 - 23" refId="ref14343" refString="Balthasar, V. (1965) Eine neue Untergattung und neue Arten der Familie Scarabaeidae. Acta Entomologica Bohemoslovaca, 62 (1), 14 - 23." type="journal article" year="1965">Balthasar, 1965</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36FEFFE36E0FFF4614E9FA5FFAAF" blockId="20.[151,1437,292,1311]" pageId="20" pageNumber="107">
<keyLead id="30D195CBFFE36E0FFF4614E9FA5FFAAF" pageId="20" pageNumber="107">
- Elytral surface dull, with distinct microsculpture (
<figureCitation id="131B2A7BFFE36E0FFD1114E9FCD9FAB1" box="[704,794,1260,1282]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12A</figureCitation>
); Elytra moderately convex in lateral view (
<figureCitation id="131B2A7BFFE36E0FFB6E14E9FAD0FAB1" box="[1215,1299,1260,1282]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="20" pageNumber="107">Fig. 12F</figureCitation>
)........................................................................
<taxonomicName id="4C204D7DFFE36E0FFC08150CFAE3FAAF" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[985,1312,1289,1311]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="107" phylum="Arthropoda" rank="species" species="vanus" status="sp. nov.">
<emphasis id="B954EAECFFE36E0FFC08150CFBB6FAAF" box="[985,1141,1289,1311]" italics="true" pageId="20" pageNumber="107">Rhytomus vanus</emphasis>
Saxton &amp; Génier
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,
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</keyLead>
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349
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<emphasis id="B954EAECFFE76E0BFF4614CFFE47FB52" bold="true" box="[151,388,1226,1250]" italics="true" pageId="16" pageNumber="103">Tesserodon papuanum</emphasis>
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(original description)
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<emphasis id="B954EAECFFE76E0BFF461514FE06FA9B" bold="true" box="[151,453,1297,1323]" pageId="16" pageNumber="103">Name-bearing type data.</emphasis>
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(
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): ||
<collectingCountry id="F337766EFFE76E0BFD3B1514FC3DFA9B" box="[746,1022,1297,1323]" name="Papua New Guinea" pageId="16" pageNumber="103">PAPUA NEW GUINEA</collectingCountry>
|
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, 1400 m. |
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.
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|
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18-19 ||; || PAPUA NEW GUINEA | Wau,
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. |
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| S. Peck, carT18-19 || [photocopy of handwritten label]; || H. &amp; A.Howden | Collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011856 || [QR code label]; ||
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| papuanum
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|
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det. || [partly handwritten].
<emphasis id="B954EAECFFE76E0BFDF215A4FDBCFA0B" bold="true" box="[547,639,1441,1467]" pageId="16" pageNumber="103">Studied</emphasis>
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<emphasis id="B954EAECFFE76E0BFF1615C0FC57FA6F" bold="true" box="[199,916,1476,1506]" pageId="16" pageNumber="103">
Material examined (
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,
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).
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:
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,
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W Bululo
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,
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,
<collectedFrom id="383C7503FFE76E0BFCF815ECFB9EF9B3" box="[809,1117,1513,1539]" pageId="16" pageNumber="103">mixed montane forest litter</collectedFrom>
,
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, (RSA2000- 039a)—
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(
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,
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</quantity>
, [
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,
<geoCoordinate id="EE145039FFE76E0BFC821608FC00F998" box="[851,963,1549,1576]" degrees="146" direction="east" minutes="25" orientation="longitude" pageId="16" pageNumber="103" precision="925" value="146.41667">146°25’E</geoCoordinate>
],
<date id="FF9E103EFFE76E0BFC051608FB8DF997" box="[980,1102,1549,1575]" pageId="16" pageNumber="103" value="1984-11-18">
<collectingDate id="EFDAE9D6FFE76E0BFC051608FB8DF997" box="[980,1102,1549,1575]" pageId="16" pageNumber="103" value="1984-11-18">18.xi.1984</collectingDate>
</date>
,
<collectingMethod id="52614EE9FFE76E0BFB881608FB01F997" box="[1113,1218,1549,1575]" pageId="16" pageNumber="103">infly trap</collectingMethod>
,
<collectorName id="26D55328FFE76E0BFB1D1608FA84F997" box="[1228,1351,1549,1575]" pageId="16" pageNumber="103">H. Roberts</collectorName>
, (C.I.E. A16609)—
<specimenCount id="9D26FD77FFE76E0BFECD1634FEFCF9FB" box="[284,319,1585,1611]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED31AE3BFFE76E0BFE9A1634FE7FF9FB" box="[331,444,1585,1611]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="16" pageNumber="103" type="Museum">NHMUK</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFE1C1634FC99F9DF" collectedFrom="montane forest litter" collectingDate="2000-02-06" collectingDateMax="2000-02-08" collectingDateMin="1974-06-29" collectionCode="CMNC" collectorName="R. S. Anderson" country="Papua New Guinea" county="Aseki Road" elevation="2050" latitude="-7.3333335" location="Mount Kaindi" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Morobe" typeStatus="holotype">
<collectingMunicipality id="6BFBAC84FFE76E0BFE1C1634FDA8F9FB" box="[461,619,1585,1611]" pageId="16" pageNumber="103">Mount Kaindi</collectingMunicipality>
,
<locationDeviation id="9A2D2FDCFFE76E0BFDA51634FCEEF9FB" box="[628,813,1585,1611]" location="Wau" pageId="16" pageNumber="103">
<quantity id="4CD89B1BFFE76E0BFDA51634FD68F9FB" box="[628,683,1585,1611]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="103" unit="km" value="9.0">9 km</quantity>
NNW Wau
</locationDeviation>
,
<quantity id="4CD89B1BFFE76E0BFCE41634FC49F9FB" box="[821,906,1585,1612]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="16" pageNumber="103" unit="m" value="2050.0">
<elevation id="000DD1CDFFE76E0BFCE41634FC49F9FB" box="[821,906,1585,1612]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="16" pageNumber="103" unit="m" value="2050.0">2050 m</elevation>
</quantity>
,
<geoCoordinate id="EE145039FFE76E0BFC451634FBDDF9FB" box="[916,1054,1585,1611]" degrees="07" direction="south" minutes="20" orientation="latitude" pageId="16" pageNumber="103" precision="15" seconds="24" value="-7.34">07°20’24’’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFE76E0BFBF91634FB00F9FB" box="[1064,1219,1585,1611]" degrees="146" direction="east" minutes="40" orientation="longitude" pageId="16" pageNumber="103" precision="15" seconds="01" value="146.66695">146°40’01’’E</geoCoordinate>
,
<date id="FF9E103EFFE76E0BFB1D1634FAF0F9FB" box="[1228,1331,1585,1611]" pageId="16" pageNumber="103" value="2000-02-06">
<collectingDate id="EFDAE9D6FFE76E0BFB1D1634FAF0F9FB" box="[1228,1331,1585,1611]" pageId="16" pageNumber="103" value="2000-02-06">6.ii.2000</collectingDate>
</date>
,
<collectedFrom id="383C7503FFE76E0BFAED1634FED4F9DF" pageId="16" pageNumber="103">montane forest litter</collectedFrom>
,
<collectorName id="26D55328FFE76E0BFEF51650FE0FF9DF" box="[292,460,1621,1647]" pageId="16" pageNumber="103">R.S. Anderson</collectorName>
, (RSA2000-035b)—
<specimenCount id="9D26FD77FFE76E0BFD6C1650FD27F9DF" box="[701,740,1621,1648]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED31AE3BFFE76E0BFD241650FC8DF9DF" box="[757,846,1621,1647]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="16" pageNumber="103">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFCB21650FD57F923" collectedFrom="forest litter" collectingDate="1974-07-01" collectingDateMax="1974-07-03" collectingDateMin="1974-07-01" collectingMethod="berlese" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1219" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="1" specimenCount-female="1" specimenCount-male="6" stateProvince="Morobe" typeStatus="holotype">
<location id="8EFF6025FFE76E0BFCB21650FC5AF9DF" LSID="urn:lsid:plazi:treatment:038987E8FFE76E0AFF4614CFFE52FE1D:8EFF6025FFE76E0BFCB21650FC5AF9DF" box="[867,921,1621,1647]" country="Papua New Guinea" county="Aseki Road" latitude="-7.3333335" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" name="Wau" pageId="16" pageNumber="103" stateProvince="Morobe">Wau</location>
,
<quantity id="4CD89B1BFFE76E0BFC771650FC3CF9DF" box="[934,1023,1621,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="16" pageNumber="103" unit="m" value="1219.0">
<elevation id="000DD1CDFFE76E0BFC771650FC3CF9DF" box="[934,1023,1621,1647]" metricMagnitude="3" metricUnit="m" metricValue="1.219" pageId="16" pageNumber="103" unit="m" value="1219.0">1219 m</elevation>
</quantity>
, [
<geoCoordinate id="EE145039FFE76E0BFBC51650FBB6F9DF" box="[1044,1141,1621,1647]" degrees="07" direction="south" minutes="20" orientation="latitude" pageId="16" pageNumber="103" precision="925" value="-7.3333335">07°20’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFE76E0BFB521650FB30F9DF" box="[1155,1267,1621,1647]" degrees="146" direction="east" minutes="43" orientation="longitude" pageId="16" pageNumber="103" precision="925" value="146.71666">146°43’E</geoCoordinate>
],
<date id="FF9E103EFFE76E0BFAD91650FA5BF9DF" box="[1288,1432,1621,1647]" pageId="16" pageNumber="103" value="1974-07-01" valueMax="1974-07-03" valueMin="1974-07-01">
<collectingDate id="EFDAE9D6FFE76E0BFAD91650FA5BF9DF" box="[1288,1432,1621,1647]" pageId="16" pageNumber="103" value="1974-07-01" valueMax="1974-07-03" valueMin="1974-07-01">1–3.vii.1974</collectingDate>
</date>
,
<collectedFrom id="383C7503FFE76E0BFF46167CFED5F923" box="[151,278,1657,1683]" pageId="16" pageNumber="103">forest litter</collectedFrom>
,
<collectingMethod id="52614EE9FFE76E0BFEF2167CFEB6F923" box="[291,373,1657,1683]" pageId="16" pageNumber="103">berlese</collectingMethod>
,
<collectorName id="26D55328FFE76E0BFE50167CFE18F923" box="[385,475,1657,1683]" pageId="16" pageNumber="103">S. Peck</collectorName>
—
<specimenCount id="9D26FD77FFE76E0BFE28167CFDDDF923" box="[505,542,1657,1683]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED31AE3BFFE76E0BFDFE167CFD4BF923" box="[559,648,1657,1683]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="16" pageNumber="103">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFD4D167CFA99F923" box="[668,1370,1657,1683]" collectedFrom="forest litter" collectingDate="1974-07-01" collectingDateMax="1974-07-03" collectingDateMin="1974-07-01" collectingMethod="berlese" collectionCode="MNHN" collectorName="H. Howden" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Morobe" typeStatus="holotype">
same locality,
<date id="FF9E103EFFE76E0BFC91167CFC13F923" box="[832,976,1657,1683]" pageId="16" pageNumber="103" value="1974-07-01" valueMax="1974-07-03" valueMin="1974-07-01">
<collectingDate id="EFDAE9D6FFE76E0BFC91167CFC13F923" box="[832,976,1657,1683]" pageId="16" pageNumber="103" value="1974-07-01" valueMax="1974-07-03" valueMin="1974-07-01">1–3.vii.1974</collectingDate>
</date>
,
<collectorName id="26D55328FFE76E0BFC0C167CFBA7F923" box="[989,1124,1657,1683]" pageId="16" pageNumber="103">H. Howden</collectorName>
—
<specimenCount id="9D26FD77FFE76E0BFB50167CFB6AF923" box="[1153,1193,1657,1683]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFB66167CFB21F923" box="[1207,1250,1657,1683]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED31AE3BFFE76E0BFB23167CFA8DF923" box="[1266,1358,1657,1683]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="103">MNHN</collectionCode>
);
</materialsCitation>
<location id="8EFF6025FFE76E0BFAB2167CFA5AF923" LSID="urn:lsid:plazi:treatment:038987E8FFE76E0AFF4614CFFE52FE1D:8EFF6025FFE76E0BFAB2167CFA5AF923" box="[1379,1433,1657,1683]" country="Papua New Guinea" county="Aseki Road" latitude="-7.3333335" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" name="Wau" pageId="16" pageNumber="103" stateProvince="Morobe">Wau</location>
,
<quantity id="4CD89B1BFFE76E0BFF461698FF2DF907" box="[151,238,1693,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="103" unit="m" value="1400.0">
<elevation id="000DD1CDFFE76E0BFF461698FF2DF907" box="[151,238,1693,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="103" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, [
<geoCoordinate id="EE145039FFE76E0BFED01698FEA2F907" box="[257,353,1693,1719]" degrees="07" direction="south" minutes="20" orientation="latitude" pageId="16" pageNumber="103" precision="925" value="-7.3333335">07°20’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFE76E0BFEBC1698FE1EF907" box="[365,477,1693,1719]" degrees="146" direction="east" minutes="43" orientation="longitude" pageId="16" pageNumber="103" precision="925" value="146.71666">146°43’E</geoCoordinate>
],
<date id="FF9E103EFFE76E0BFE211698FD72F907" box="[496,689,1693,1719]" pageId="16" pageNumber="103" value="1974-06-29" valueMax="1974-07-02" valueMin="1974-06-29">
<collectingDate id="EFDAE9D6FFE76E0BFE211698FD72F907" box="[496,689,1693,1719]" pageId="16" pageNumber="103" value="1974-06-29" valueMax="1974-07-02" valueMin="1974-06-29">29.vi.–2.vii.1974</collectingDate>
</date>
,
<collectingMethod id="52614EE9FFE76E0BFD6C1698FC8FF907" box="[701,844,1693,1719]" pageId="16" pageNumber="103">carrion traps</collectingMethod>
,
<collectorName id="26D55328FFE76E0BFC861698FC6DF907" box="[855,942,1693,1719]" pageId="16" pageNumber="103">S. Peck</collectorName>
, (carrion traps 18-19)—
<specimenCount id="9D26FD77FFE76E0BFB6C1698FB12F907" box="[1213,1233,1693,1719]" count="1" pageId="16" pageNumber="103" type="male">♂</specimenCount>
<typeStatus id="549B885CFFE76E0BFB091698FAFEF907" box="[1240,1341,1693,1719]" pageId="16" pageNumber="103" type="holotype">holotype</typeStatus>
,
<materialsCitation id="3B483CA3FFE76E0BFA991698FE7DF96B" collectingDate="1974-06-29" collectingDateMax="1974-07-02" collectingDateMin="1974-06-29" collectingMethod="carrion traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="15" specimenCount-female="3" specimenCount-male="12" stateProvince="Morobe" typeStatus="paratype">
<specimenCount id="9D26FD77FFE76E0BFA991698FA54F907" box="[1352,1431,1693,1719]" count="12" pageId="16" pageNumber="103" type="male">12 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFF4616C4FF14F96B" box="[151,215,1729,1755]" count="3" pageId="16" pageNumber="103" type="female">3 ♀♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFF0E16C4FE89F96B" box="[223,330,1729,1755]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFE8B16C4FE71F96B" box="[346,434,1729,1755]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="16" pageNumber="103">CMNC</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFE1716C4FCE8F96B" box="[454,811,1729,1755]" collectingDate="1974-06-29" collectingDateMax="1974-07-02" collectingDateMin="1974-06-29" collectingMethod="carrion traps" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Morobe" typeStatus="paratype">
<specimenCount id="9D26FD77FFE76E0BFE1716C4FDCBF96B" box="[454,520,1729,1755]" count="2" pageId="16" pageNumber="103" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFE76E0BFDC416C4FD83F96B" box="[533,576,1729,1755]" count="1" pageId="16" pageNumber="103" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFD9916C4FD70F96B" box="[584,691,1729,1755]" pageId="16" pageNumber="103" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFD1216C4FCDCF96B" box="[707,799,1729,1755]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="103">MNHN</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFCE216C4FEB1F94F" collectingDate="1974-06-29" collectingDateMax="1974-07-02" collectingDateMin="1974-06-29" collectingMethod="carrion traps / carrion trap" collectionCode="CMNC, MNHN" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="1" specimenCount-female="5" specimenCount-male="15" stateProvince="Morobe" typeStatus="paratype">
same locality,
<date id="FF9E103EFFE76E0BFC0716C4FB54F96B" box="[982,1175,1729,1755]" pageId="16" pageNumber="103" value="1974-06-29" valueMax="1974-07-02" valueMin="1974-06-29">
<collectingDate id="EFDAE9D6FFE76E0BFC0716C4FB54F96B" box="[982,1175,1729,1755]" pageId="16" pageNumber="103" value="1974-06-29" valueMax="1974-07-02" valueMin="1974-06-29">29.vi.–2.vii.1974</collectingDate>
</date>
,
<collectingMethod id="52614EE9FFE76E0BFB7516C4FAF7F96B" box="[1188,1332,1729,1755]" pageId="16" pageNumber="103">carrion traps</collectingMethod>
,
<collectorName id="26D55328FFE76E0BFA9116C4FA5BF96B" box="[1344,1432,1729,1755]" pageId="16" pageNumber="103">S. Peck</collectorName>
, (
<collectingMethod id="52614EE9FFE76E0BFF4E16E0FEE0F94F" box="[159,291,1765,1791]" pageId="16" pageNumber="103">carrion trap</collectingMethod>
20)—
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFEA316E0FD97F94F" box="[370,596,1765,1791]" collectingDate="1974-06-29" collectingDateMax="1974-07-02" collectingDateMin="1974-06-29" collectingMethod="carrion traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="1" specimenCount-female="1" specimenCount-male="15" stateProvince="Morobe" typeStatus="allotype">
<specimenCount id="9D26FD77FFE76E0BFEA316E0FE43F94F" box="[370,384,1765,1791]" count="1" pageId="16" pageNumber="103" type="female">♀</specimenCount>
<typeStatus id="549B885CFFE76E0BFE5616E0FE22F94F" box="[391,481,1765,1791]" pageId="16" pageNumber="103" type="allotype">allotype</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFE2116E0FD8BF94F" box="[496,584,1765,1791]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="16" pageNumber="103">CMNC</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B483CA3FFE76E0BFD8A16E0FC9CF94F" box="[603,863,1765,1791]" collectingDate="1974-06-29" collectingDateMax="1974-07-02" collectingDateMin="1974-06-29" collectingMethod="carrion traps" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" county="Aseki Road" elevation="1750" latitude="-7.3333335" location="Wau" longLatPrecision="1303" longitude="146.71666" municipality="Mount Kaindi" pageId="16" pageNumber="103" specimenCount="1" specimenCount-female="5" specimenCount-male="1" stateProvince="Morobe" typeStatus="paratype">
<specimenCount id="9D26FD77FFE76E0BFD8A16E0FD46F94F" box="[603,645,1765,1791]" count="1" pageId="16" pageNumber="103" type="male">1 ♂</specimenCount>
<typeStatus id="549B885CFFE76E0BFD5D16E0FD2FF94F" box="[652,748,1765,1791]" pageId="16" pageNumber="103" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED31AE3BFFE76E0BFD2A16E0FC94F94F" box="[763,855,1765,1791]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="103">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFE76E0AFF16170CFE52FE1D" lastPageId="17" lastPageNumber="104" pageId="16" pageNumber="103" type="description">
<paragraph id="8B9F36FEFFE76E0AFF16170CFE52FE1D" blockId="16.[151,1437,1297,2043]" lastBlockId="17.[151,1436,151,429]" lastPageId="17" lastPageNumber="104" pageId="16" pageNumber="103">
<emphasis id="B954EAECFFE76E0BFF16170CFEAEF893" bold="true" box="[199,365,1801,1827]" pageId="16" pageNumber="103">Redescription</emphasis>
.
<emphasis id="B954EAECFFE76E0BFEAC170CFDF0F893" bold="true" box="[381,563,1801,1827]" pageId="16" pageNumber="103">Measurements.</emphasis>
Body length
<quantity id="4CD89B1BFFE76E0BFD1F170CFC9BF893" box="[718,856,1801,1828]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="16" pageNumber="103" unit="mm" value="4.699999999999999" valueMax="5.3" valueMin="4.1">4.1–5.3 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFE76E0BFBB7170CFB32F893" box="[1126,1265,1801,1827]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="16" pageNumber="103" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
.
<emphasis id="B954EAECFFE76E0BFB2C170CFA83F893" bold="true" box="[1277,1344,1801,1827]" pageId="16" pageNumber="103">Body.</emphasis>
Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex.
<emphasis id="B954EAECFFE76E0BFB951728FB48F8F7" bold="true" box="[1092,1163,1837,1863]" pageId="16" pageNumber="103">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width.
<emphasis id="B954EAECFFE76E0BFB811770FB13F83F" bold="true" box="[1104,1232,1909,1935]" pageId="16" pageNumber="103">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.9, lateral pronotal edge rounded, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFE76E0BFCE717B8FC49F867" bold="true" box="[822,906,1981,2007]" pageId="16" pageNumber="103">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures.
<emphasis id="B954EAECFFE66E0AFCCC10DAFC9FFF49" bold="true" box="[797,860,223,249]" pageId="17" pageNumber="104">Legs.</emphasis>
Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge.
<emphasis id="B954EAECFFE66E0AFC5D1107FBC4FEAC" bold="true" box="[908,1031,258,284]" pageId="17" pageNumber="104">Pygidium.</emphasis>
Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (
<figureCitation id="131B2A7BFFE66E0AFE03114EFDE4FED5" box="[466,551,331,357]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="17" pageNumber="104">Fig. 8F</figureCitation>
), pygidium of female evenly convex.
<emphasis id="B954EAECFFE66E0AFC00114FFB8FFED4" bold="true" box="[977,1100,330,356]" pageId="17" pageNumber="104">Aedeagus.</emphasis>
Left paramere apex truncated apically, distinctly angular externally (
<figureCitation id="131B2A7BFFE66E0AFD9A116AFD70FE39" box="[587,691,367,393]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="17" pageNumber="104">Fig. 10B</figureCitation>
).
<emphasis id="B954EAECFFE66E0AFD19116AFC55FE38" bold="true" box="[712,918,366,393]" pageId="17" pageNumber="104">Female genitalia.</emphasis>
Spermatheca with two apodemes basally and one apodeme apically.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="728018FDD2B0C4593F199915B97B37DC">Rhytomus Génier &amp; Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:affiliation id="E634CB8F4883DF2ADAB4F8E31C041F5E">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. &amp; Corresponding author</mods:affiliation>
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<subSubSection id="C33A6575FFF16E1DFF461092FDBEFF01" box="[151,637,150,178]" pageId="6" pageNumber="93" type="nomenclature">
<paragraph id="8B9F36FEFFF16E1DFF461092FDBEFF01" blockId="6.[151,963,150,214]" box="[151,637,150,178]" pageId="6" pageNumber="93">
<heading id="D0D78192FFF16E1DFF461092FDBEFF01" bold="true" box="[151,637,150,178]" fontSize="11" level="1" pageId="6" pageNumber="93" reason="1">
<emphasis id="B954EAECFFF16E1DFF461092FDBEFF01" bold="true" box="[151,637,150,178]" pageId="6" pageNumber="93">
<taxonomicName id="4C204D7DFFF16E1DFF461092FE2DFF01" authority="Genier &amp; Saxton" authorityName="Genier &amp; Saxton" authorityYear="2025" box="[151,494,150,178]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B954EAECFFF16E1DFF461092FED2FF02" bold="true" box="[151,273,151,178]" italics="true" pageId="6" pageNumber="93">Rhytomus</emphasis>
Génier &amp; Saxton
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,
<taxonomicNameLabel id="A2675797FFF16E1DFE2A1092FDBEFF01" box="[507,637,151,177]" pageId="6" pageNumber="93" rank="genus">new genus</taxonomicNameLabel>
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<paragraph id="8B9F36FEFFF16E1DFF461105FD30FEA7" blockId="6.[151,755,255,280]" box="[151,755,255,280]" pageId="6" pageNumber="93">
<typeStatus id="549B885CFFF16E1DFF461105FF10FEA8" box="[151,211,256,280]" pageId="6" pageNumber="93">Types</typeStatus>
species.
<taxonomicName id="4C204D7DFFF16E1DFEFE1105FD2CFEA7" authority="van Lansberge, 1885" authorityName="van Lansberge" authorityYear="1885" box="[303,751,255,280]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="elongatum">
<emphasis id="B954EAECFFF16E1DFEFE1105FDD1FEA8" box="[303,530,256,280]" italics="true" pageId="6" pageNumber="93">Tesserodon elongatum</emphasis>
<bibRefCitation id="EFB14B0FFFF16E1DFDC81105FD2CFEA7" author="Van Lansberge, J. W." box="[537,751,255,280]" pageId="6" pageNumber="93" pagination="375 - 400" refId="ref16891" refString="Van Lansberge, J. W. (1885) Description d'especes nouvelles de coleopteres appartenant au Musee civique de Genes. Annali del Museo civico di storia naturale di Genova, 2 (2), 375 - 400." type="journal article" year="1885">van Lansberge, 1885</bibRefCitation>
</taxonomicName>
.
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<subSubSection id="C33A6575FFF16E1DFF461140FECFFE40" pageId="6" pageNumber="93" type="diagnosis">
<paragraph id="8B9F36FEFFF16E1DFF461140FECFFE40" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF461140FED1FEEF" bold="true" box="[151,274,325,351]" pageId="6" pageNumber="93">Diagnosis.</emphasis>
The presence of a trochanterofemoral pit combined with a pseudepipleuron, the presence of a posterior hypomeral carina and well-defined elytral striae 9 and 10 will separate
<taxonomicName id="4C204D7DFFF16E1DFC66116FFBE6FE34" box="[951,1061,362,388]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF16E1DFC66116FFBE6FE34" box="[951,1061,362,388]" italics="true" pageId="6" pageNumber="93">Rhytomus</emphasis>
</taxonomicName>
from all the other genera in New
<collectingCountry id="F337766EFFF16E1DFF46118BFF2FFE18" box="[151,236,398,424]" name="Guinea-Bissau" pageId="6" pageNumber="93">Guinea</collectingCountry>
. Within the Australasian endemic clade,
<taxonomicName id="4C204D7DFFF16E1DFD7F118BFCDFFE18" box="[686,796,398,424]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF16E1DFD7F118BFCDFFE18" box="[686,796,398,424]" italics="true" pageId="6" pageNumber="93">Rhytomus</emphasis>
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can be distinguished from other genera by those previous characters listed in conjunction with having 9 elytral striae, and wide eyes with an interocular space 3–6 times the eye width.
</paragraph>
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<subSubSection id="C33A6575FFF16E1DFF1611FCFD32FA28" pageId="6" pageNumber="93" type="description">
<paragraph id="8B9F36FEFFF16E1DFF1611FCFA5EFD78" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF1611FCFE61FDA4" bold="true" box="[199,418,505,532]" pageId="6" pageNumber="93">Description. Body.</emphasis>
Total length 3.0–
<quantity id="4CD89B1BFFF16E1DFDB511FFFD79FDA4" box="[612,698,506,532]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="93" unit="mm" value="6.3">6.3 mm</quantity>
. Tegument hard, brittle. Color dark reddish brown to black (teneral specimens lighter in color), ventral surface and legs usually lighter in color, lacking metallic sheen.
<emphasis id="B954EAECFFF16E1DFB20121BFAFBFD88" bold="true" box="[1265,1336,542,568]" pageId="6" pageNumber="93">Head.</emphasis>
Anterior edge of clypeus with two clypeal teeth, with or without emargination on external side of each tooth. Dorsal part of eyes wide (
<figureCitation id="131B2A7BFFF16E1DFEEB1263FE6CFD30" box="[314,431,614,640]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4A,B</figureCitation>
), inner edge straight basally, outer edge arcuate, with at least 12 facets along its widest portion. Ocular canthus incomplete. Occiput of head simply rounded (
<figureCitation id="131B2A7BFFF16E1DFC60128FFBE5FD14" box="[945,1062,650,676]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4C,D</figureCitation>
). Antennae nine segmented, with 3-antennomere club, club antennomeres approximately equal width. Antennomeres 7 and 8 lacking median cavity.
</paragraph>
<paragraph id="8B9F36FEFFF16E1DFF4612D7FD32FA28" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF4612D7FED4FD5C" bold="true" box="[151,279,722,748]" pageId="6" pageNumber="93">Pronotum.</emphasis>
Basal edge of pronotum broadly arcuate. Posterior hypomeral carina present, almost as long as half lateral pronotal edge. Anterior edge lacking membranous imbrication, anterior marginal bead very narrow.
<emphasis id="B954EAECFFF16E1DFA9812F3FA5FFCA0" bold="true" box="[1353,1436,758,784]" pageId="6" pageNumber="93">Elytra.</emphasis>
Elytra with 9 striae, 7 on disc and 2 on the pseudepipleuron adjacent to the epipleural edge (
<figureCitation id="131B2A7BFFF16E1DFB54131FFB1DFC84" box="[1157,1246,794,820]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4D</figureCitation>
). Stria 8 missing. Stria 9 effaced anteriorly, fused or very closely set with tenth stria past basal third to posterior declivity where it is distinct again. With a well-defined pseudepipleural fold from base to apical declivity outside stria 7. Laterobasal elytral pit present or absent (
<figureCitation id="131B2A7BFFF16E1DFE381383FD8BFC10" box="[489,584,902,928]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4A</figureCitation>
).
<emphasis id="B954EAECFFF16E1DFD8F1380FD32FC2F" bold="true" box="[606,753,901,927]" pageId="6" pageNumber="93">Hind wings.</emphasis>
Present, fully developed.
<emphasis id="B954EAECFFF16E1DFBF51380FB5DFC2F" bold="true" box="[1060,1182,901,927]" pageId="6" pageNumber="93">Pygidium.</emphasis>
Deeply and narrowly sulcate across the base, sulcus usually wider and deeper laterally (
<figureCitation id="131B2A7BFFF16E1DFCBE13AFFC70FC74" box="[879,947,938,964]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="6" pageNumber="93">Fig. 8</figureCitation>
).
<emphasis id="B954EAECFFF16E1DFC1613ACFB17FC73" bold="true" box="[967,1236,937,963]" pageId="6" pageNumber="93">Pterothoracic ventrite.</emphasis>
Mesepimera with coarse semicircular ocellate punctures.
<emphasis id="B954EAECFFF16E1DFD8A13C8FC84FC57" bold="true" box="[603,839,973,999]" pageId="6" pageNumber="93">Abdominal ventrite</emphasis>
. Ventrite 6 approximately as long as 1–5 combined along midline.
<emphasis id="B954EAECFFF16E1DFE9113F7FEBCFBBC" bold="true" box="[320,383,1010,1036]" pageId="6" pageNumber="93">Legs.</emphasis>
Protibiae (
<figureCitation id="131B2A7BFFF16E1DFE2B13F7FD8DFBBC" box="[506,590,1010,1036]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4E</figureCitation>
) of male abruptly internally expanded apically, distal edge of internal angle with a row of contiguous spiniform setae flanked posteriorly by a row of long squamiform setae, with more or less elongated triangular spur. Protibiae distally straight with 3 teeth on outer edge, each tooth with two groups of setae dorsally. Prothoracic legs with trochanterofemoral anterior pit (
<figureCitation id="131B2A7BFFF16E1DFC61145BFBC8FBC8" box="[944,1035,1118,1144]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="6" pageNumber="93">Fig. 4C</figureCitation>
), trochanter excavated dorsally for reception of inner apical edge of protibia. Protarsi with 5 tarsomeres, tarsomere 1 and 5 longer. Long, setae on all tarsomeres or tarsomere 1 lacking setae. Male mesotibiae with dense setae on ventral edge. Metatibiae of males with well-developed tooth on inner apical angle. Mesotarsi with 5 tarsomeres, tarsomere 2 about same length as tarsomere 5. Metatarsomere 1 widest, metatarsomere 2 equal to metatarsomere
<quantity id="4CD89B1BFFF16E1DFBE014EBFB9CFAB8" box="[1073,1119,1262,1289]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="6" pageNumber="93" unit="in" value="5.0">5 in</quantity>
length, all tarsomeres with long setae ventrally. Claws simple or slightly angulate at base. Metacoxal connecting membrane apodeme present, oval (internal).
<emphasis id="B954EAECFFF16E1DFE981530FE7CFAFF" bold="true" box="[329,447,1333,1359]" pageId="6" pageNumber="93">Genitalia.</emphasis>
Aedeagus with parameres asymmetrical, right paramere apically bent downwards and expanded, left paramere more or less bent downward apically or simply rounded (
<figureCitation id="131B2A7BFFF16E1DFB81155FFB61FAC4" box="[1104,1186,1370,1396]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="6" pageNumber="93">Figs. 9</figureCitation>
,
<figureCitation id="131B2A7BFFF16E1DFB60155FFB13FAC4" box="[1201,1232,1370,1396]" captionStart="FIGURE 10" captionStartId="14.[152,255,1979,2003]" captionTargetBox="[155,1431,490,1949]" captionTargetId="figure-166@14.[151,1435,486,1953]" captionTargetPageId="14" captionText="FIGURE 10. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral. A, R. latidentatus; B, R. papuanus; C, R. sodalis; D, R. vanus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746466" httpUri="https://zenodo.org/record/14746466/files/figure.png" pageId="6" pageNumber="93">10</figureCitation>
). Spermatheca Vshaped, with or without apodeme distally and basally.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFF16E1DFF1615A7FED4F998" pageId="6" pageNumber="93" type="etymology">
<paragraph id="8B9F36FEFFF16E1DFF1615A7FED4F998" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF1615A7FE8DFA0C" bold="true" box="[199,334,1442,1468]" pageId="6" pageNumber="93">Etymology.</emphasis>
Based on the combination of two Greek words. The feminine noun ῥῠτῐ́ς (rhytís) meaning wrinkle, crease and the masculine noun ὦμος (omos) meaning shoulder, the terminal component transliterated in Latin with the masculine ending -us (ICZN art. 30.1.2). The name refers to the presence of a crease on the humeral surface of the elytron.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFF16E1DFF161634FDAEF9C1" pageId="6" pageNumber="93" type="distribution">
<paragraph id="8B9F36FEFFF16E1DFF161634FDAEF9C1" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF161634FEA2F9FB" bold="true" box="[199,353,1585,1611]" pageId="6" pageNumber="93">Distribution.</emphasis>
<collectingCountry id="F337766EFFF16E1DFEB71637FDFFF9FC" box="[358,572,1586,1612]" name="Papua New Guinea" pageId="6" pageNumber="93">Papua New Guinea</collectingCountry>
and
<collectingCountry id="F337766EFFF16E1DFDA11637FD1EF9FC" box="[624,733,1586,1612]" name="Indonesia" pageId="6" pageNumber="93">Indonesia</collectingCountry>
(Seram Island, Waigeo Island and
<collectingRegion id="49E4F81CFFF16E1DFB8C1637FB64F9FC" box="[1117,1191,1586,1612]" country="Indonesia" name="Papua Barat" pageId="6" pageNumber="93">Papua</collectingRegion>
). Specimens collected at elevations from
<quantity id="4CD89B1BFFF16E1DFEB91653FE50F9C0" box="[360,403,1622,1648]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="93" unit="m" value="3.0">3 m</quantity>
to
<quantity id="4CD89B1BFFF16E1DFE661653FDCFF9C0" box="[439,524,1622,1649]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="6" pageNumber="93" unit="m" value="2050.0">2050 m</quantity>
(
<figureCitation id="131B2A7BFFF16E1DFDCA1653FD9CF9C1" box="[539,607,1622,1649]" captionStart="FIGURE 5" captionStartId="7.[152,255,1897,1921]" captionTargetBox="[199,1388,1125,1872]" captionTargetId="figure-84@7.[199,1388,1125,1872]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Rhytomus in mainland New Guinea. Note that some distribution points are extrapolated based on best available data, view treatment of each species for more detail." figureDoi="http://doi.org/10.5281/zenodo.14746452" httpUri="https://zenodo.org/record/14746452/files/figure.png" pageId="6" pageNumber="93">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFF16E1DFF16167FFEC7F804" pageId="6" pageNumber="93" type="discussion">
<paragraph id="8B9F36FEFFF16E1DFF16167FFEC7F804" blockId="6.[151,1437,325,1972]" pageId="6" pageNumber="93">
<emphasis id="B954EAECFFF16E1DFF16167FFEF8F924" bold="true" box="[199,315,1658,1684]" pageId="6" pageNumber="93">Remarks.</emphasis>
<taxonomicName id="4C204D7DFFF16E1DFE99167FFC6BF924" authority="Krikken &amp; Huijbregts, 2009" authorityName="Krikken &amp; Huijbregts" authorityYear="2009" box="[328,936,1658,1684]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="seramicum">
<emphasis id="B954EAECFFF16E1DFE99167FFD8EF924" box="[328,589,1658,1684]" italics="true" pageId="6" pageNumber="93">Tesserodon seramicum</emphasis>
<bibRefCitation id="EFB14B0FFFF16E1DFD8B167FFC6BF924" author="Krikken, J. &amp; Huijbregts, J." box="[602,936,1658,1684]" pageId="6" pageNumber="93" pagination="237 - 266" refId="ref15470" refString="Krikken, J. &amp; Huijbregts, J. (2009) A new Haroldius Boucomont and a new Tesserodon Hope from the Moluccas (Coleoptera: Scarabaeidae: Scarabaeinae). Koleopterologische Rundschau, 79, 237 - 266." type="journal article" year="2009">Krikken &amp; Huijbregts, 2009</bibRefCitation>
</taxonomicName>
(described from Seram Island,
<collectingCountry id="F337766EFFF16E1DFAF7167FFA56F924" box="[1318,1429,1658,1684]" name="Indonesia" pageId="6" pageNumber="93">Indonesia</collectingCountry>
) and
<taxonomicName id="4C204D7DFFF16E1DFF1B169BFCE2F908" authority="Ochi, Kon &amp; Hartini, 2017" authorityName="Ochi, Kon &amp; Hartini" authorityYear="2017" box="[202,801,1694,1720]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="waigeoensis">
<emphasis id="B954EAECFFF16E1DFF1B169BFE18F908" box="[202,475,1694,1720]" italics="true" pageId="6" pageNumber="93">Tesserodon waigeoensis</emphasis>
<bibRefCitation id="EFB14B0FFFF16E1DFE34169BFCE2F908" author="Ochi, T. &amp; Kon, M. &amp; Hartini, S." box="[485,801,1694,1720]" pageId="6" pageNumber="93" pagination="59 - 66" refId="ref16278" refString="Ochi, T., Kon, M. &amp; Hartini, S. (2017) Descriptions of a new species of the genus Tesserodon and two new species of the genus Temnoplectron from West Papua, Indonesia (Coleoptera, Scarabaeidae, Scarabaeinae). Kogane, 20, 59 - 66." type="journal article" year="2017">Ochi, Kon &amp; Hartini, 2017</bibRefCitation>
</taxonomicName>
(described from Waigeo Island,
<collectingRegion id="49E4F81CFFF16E1DFB4E169BFAE4F908" box="[1183,1319,1694,1720]" country="Indonesia" name="Papua Barat" pageId="6" pageNumber="93">West Papua</collectingRegion>
) are from outside the study area. However, examination of the illustrations provided in the original descriptions and images of a specimen (RMNH.INS.1488474) substantiate their transfer to the genus
<taxonomicName id="4C204D7DFFF16E1DFC2616E3FBA6F8B0" box="[1015,1125,1766,1792]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF16E1DFC2616E3FBA6F8B0" box="[1015,1125,1766,1792]" italics="true" pageId="6" pageNumber="93">Rhytomus</emphasis>
</taxonomicName>
and should be combined as follows:
<taxonomicName id="4C204D7DFFF16E1DFF2A170FFCFEF894" authority="(Krikken &amp; Huijbregts, 2009)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Krikken &amp; Huijbregts" baseAuthorityYear="2009" box="[251,829,1802,1828]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="seramicus" status="comb. nov.">
<emphasis id="B954EAECFFF16E1DFF2A170FFE20F894" box="[251,483,1802,1828]" italics="true" pageId="6" pageNumber="93">Rhytomus seramicus</emphasis>
(
<bibRefCitation id="EFB14B0FFFF16E1DFE20170FFCF6F894" author="Krikken, J. &amp; Huijbregts, J." box="[497,821,1802,1828]" pageId="6" pageNumber="93" pagination="237 - 266" refId="ref15470" refString="Krikken, J. &amp; Huijbregts, J. (2009) A new Haroldius Boucomont and a new Tesserodon Hope from the Moluccas (Coleoptera: Scarabaeidae: Scarabaeinae). Koleopterologische Rundschau, 79, 237 - 266." type="journal article" year="2009">Krikken &amp; Huijbregts, 2009</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B954EAECFFF16E1DFC98170FFBD5F893" bold="true" box="[841,1046,1801,1828]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A2675797FFF16E1DFC98170FFBD5F893" box="[841,1046,1801,1828]" pageId="6" pageNumber="93" rank="species">new combination</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C204D7DFFF16E1DFB9E170FFE51F8F8" authority="(Ochi, Kon &amp; Hartini, 2017)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Ochi, Kon &amp; Hartini" baseAuthorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="waigeoensis" status="comb. nov.">
<emphasis id="B954EAECFFF16E1DFB9E170FFA8DF894" box="[1103,1358,1802,1828]" italics="true" pageId="6" pageNumber="93">Rhytomus waigeoensis</emphasis>
(
<bibRefCitation id="EFB14B0FFFF16E1DFA8C170FFE49F8F8" author="Ochi, T. &amp; Kon, M. &amp; Hartini, S." pageId="6" pageNumber="93" pagination="59 - 66" refId="ref16278" refString="Ochi, T., Kon, M. &amp; Hartini, S. (2017) Descriptions of a new species of the genus Tesserodon and two new species of the genus Temnoplectron from West Papua, Indonesia (Coleoptera, Scarabaeidae, Scarabaeinae). Kogane, 20, 59 - 66." type="journal article" year="2017">Ochi, Kon &amp; Hartini, 2017</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B954EAECFFF16E1DFE4E172BFDAEF8F7" bold="true" box="[415,621,1837,1864]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A2675797FFF16E1DFE4E172BFDAEF8F7" box="[415,621,1837,1864]" pageId="6" pageNumber="93" rank="species">new combination</taxonomicNameLabel>
</emphasis>
. Spermathecal morphology was studied only for species where enough material was available. Males of
<taxonomicName id="4C204D7DFFF16E1DFDCB1756FCEEF8DC" authority="Paulian, 1937" authorityName="Paulian" authorityYear="1937" box="[538,813,1874,1900]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="heurni" status="new combination and R.">
<emphasis id="B954EAECFFF16E1DFDCB1756FD46F8DC" box="[538,645,1874,1900]" italics="true" pageId="6" pageNumber="93">R. heurni</emphasis>
<bibRefCitation id="EFB14B0FFFF16E1DFD411757FCEEF8DC" author="Paulian, R." box="[656,813,1874,1900]" pageId="6" pageNumber="93" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian, 1937</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A2675797FFF16E1DFCE61757FB9EF8DC" box="[823,1117,1873,1900]" pageId="6" pageNumber="93" rank="subSpecies">
<emphasis id="B954EAECFFF16E1DFCE61757FBC4F8DB" bold="true" box="[823,1031,1873,1900]" pageId="6" pageNumber="93">new combination</emphasis>
and
<taxonomicName id="4C204D7DFFF16E1DFB951756FB9EF8DC" authority="(Balthasar, 1965)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Balthasar" baseAuthorityYear="1965" box="[1092,1117,1875,1900]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="setulosus" status="comb. nov.">
<emphasis id="B954EAECFFF16E1DFB951756FB9EF8DC" box="[1092,1117,1875,1900]" italics="true" pageId="6" pageNumber="93">R.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="B954EAECFFF16E1DFBB61757FB0DF8DC" box="[1127,1230,1874,1900]" italics="true" pageId="6" pageNumber="93">setulosus</emphasis>
(
<bibRefCitation id="EFB14B0FFFF16E1DFB0E1757FA57F8DD" author="Balthasar, V." box="[1247,1428,1874,1901]" pageId="6" pageNumber="93" pagination="14 - 23" refId="ref14343" refString="Balthasar, V. (1965) Eine neue Untergattung und neue Arten der Familie Scarabaeidae. Acta Entomologica Bohemoslovaca, 62 (1), 14 - 23." type="journal article" year="1965">Balthasar, 1965</bibRefCitation>
)
<emphasis id="B954EAECFFF16E1DFF461773FEA6F83F" bold="true" box="[151,357,1909,1936]" pageId="6" pageNumber="93">
<taxonomicNameLabel id="A2675797FFF16E1DFF461773FEA6F83F" box="[151,357,1909,1936]" pageId="6" pageNumber="93" rank="species">new combination</taxonomicNameLabel>
</emphasis>
are unknown; the online identification key will be updated when additional material becomes available.
</paragraph>
</subSubSection>
<caption id="DF5F6676FFF06E1CFF4913A3FC2CFBB7" ID-DOI="http://doi.org/10.5281/zenodo.14746450" ID-Zenodo-Dep="14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="7" pageNumber="94" startId="7.[152,255,934,958]" targetBox="[151,1435,182,906]" targetPageId="7" targetType="figure">
<paragraph id="8B9F36FEFFF06E1CFF4913A3FC2CFBB7" blockId="7.[151,1437,934,1031]" pageId="7" pageNumber="94">
<emphasis id="B954EAECFFF06E1CFF4913A3FEDAFC0E" bold="true" box="[152,281,934,958]" pageId="7" pageNumber="94">FIGURE 4.</emphasis>
Diagnostic characters of
<taxonomicName id="4C204D7DFFF06E1CFDCE13A2FD40FC0F" box="[543,643,935,959]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF06E1CFDCE13A2FD40FC0F" box="[543,643,935,959]" italics="true" pageId="7" pageNumber="94">Rhytomus</emphasis>
</taxonomicName>
. A, dorsal head showing emargination and interocular space (
<taxonomicName id="4C204D7DFFF06E1CFB2D13A2FAACFC0F" authorityName="Paulian" authorityYear="1985" box="[1276,1391,935,959]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B954EAECFFF06E1CFB2D13A2FAACFC0F" box="[1276,1391,935,959]" italics="true" pageId="7" pageNumber="94">R. howdeni</emphasis>
</taxonomicName>
); B, dorsal head showing lack of emargination and interocular space (
<taxonomicName id="4C204D7DFFF06E1CFCFF13CEFC5EFC53" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[814,925,971,995]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="fervidus">
<emphasis id="B954EAECFFF06E1CFCFF13CEFC5EFC53" box="[814,925,971,995]" italics="true" pageId="7" pageNumber="94">R. fervidus</emphasis>
</taxonomicName>
) C, ventral view of male foreleg (
<taxonomicName id="4C204D7DFFF06E1CFB2813CEFA4EFC53" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1273,1421,971,995]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="latidentatus">
<emphasis id="B954EAECFFF06E1CFB2813CEFA4EFC53" box="[1273,1421,971,995]" italics="true" pageId="7" pageNumber="94">R. latidentatus</emphasis>
</taxonomicName>
); D, lateral view of elytra (
<taxonomicName id="4C204D7DFFF06E1CFE4613EAFE2DFBB6" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[407,494,1007,1030]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="vanus">
<emphasis id="B954EAECFFF06E1CFE4613EAFE2DFBB6" box="[407,494,1007,1030]" italics="true" pageId="7" pageNumber="94">R. vanus</emphasis>
</taxonomicName>
); E, dorsal view of male protibia (
<taxonomicName id="4C204D7DFFF06E1CFC9C13EAFC22FBB7" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[845,993,1007,1031]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="latidentatus">
<emphasis id="B954EAECFFF06E1CFC9C13EAFC22FBB7" box="[845,993,1007,1031]" italics="true" pageId="7" pageNumber="94">R. latidentatus</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<caption id="DF5F6676FFF06E1CFF49176CFCF9F815" ID-DOI="http://doi.org/10.5281/zenodo.14746452" ID-Zenodo-Dep="14746452" httpUri="https://zenodo.org/record/14746452/files/figure.png" pageId="7" pageNumber="94" startId="7.[152,255,1897,1921]" targetBox="[199,1388,1125,1872]" targetPageId="7" targetType="figure">
<paragraph id="8B9F36FEFFF06E1CFF49176CFCF9F815" blockId="7.[151,1436,1897,1957]" pageId="7" pageNumber="94">
<emphasis id="B954EAECFFF06E1CFF49176CFEDAF832" bold="true" box="[152,281,1897,1922]" pageId="7" pageNumber="94">FIGURE 5.</emphasis>
Distribution of
<taxonomicName id="4C204D7DFFF06E1CFE6F176CFDE1F831" box="[446,546,1897,1921]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFF06E1CFE6F176CFDE1F831" box="[446,546,1897,1921]" italics="true" pageId="7" pageNumber="94">Rhytomus</emphasis>
</taxonomicName>
in mainland New Guinea. Note that some distribution points are extrapolated based on best available data, view treatment of each species for more detail.
</paragraph>
</caption>
<subSubSection id="C33A6575FFFF6E13FF461092FC6EFD35" pageId="8" pageNumber="95" type="taxon_list">
<paragraph id="8B9F36FEFFFF6E13FF461092FEA4FF01" blockId="8.[151,359,151,177]" box="[151,359,151,177]" pageId="8" pageNumber="95">
<emphasis id="B954EAECFFFF6E13FF461092FEA4FF01" bold="true" box="[151,359,151,177]" pageId="8" pageNumber="95">Species checklist</emphasis>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF4610DAFD7CFEAD" blockId="8.[151,1006,223,646]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF4610DAFDBCFF4A" authority="(van Lansberge, 1885)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="van Lansberge" baseAuthorityYear="1885" box="[151,639,223,250]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="elongatus" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF4610DAFEB9FF49" box="[151,378,223,249]" italics="true" pageId="8" pageNumber="95">Rhytomus elongatus</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FE5910DAFDB5FF4A" author="Van Lansberge, J. W." box="[392,630,223,250]" pageId="8" pageNumber="95" pagination="375 - 400" refId="ref16891" refString="Van Lansberge, J. W. (1885) Description d'especes nouvelles de coleopteres appartenant au Musee civique de Genes. Annali del Museo civico di storia naturale di Genova, 2 (2), 375 - 400." type="journal article" year="1885">van Lansberge, 1885</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FD5A10E5FC8FFF49" box="[651,844,223,250]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
(
<typeStatus id="549B885CFFFF6E13FC8A10DAFC4EFF49" box="[859,909,223,249]" pageId="8" pageNumber="95">type</typeStatus>
species)
<taxonomicName id="4C204D7DFFFF6E13FF461106FDEEFEAD" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,557,259,285]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="fervidus" status="sp. nov.">
<emphasis id="B954EAECFFFF6E13FF461106FEA4FEAD" box="[151,359,259,285]" italics="true" pageId="8" pageNumber="95">Rhytomus fervidus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDE91101FD7CFEAD" box="[568,703,259,286]" pageId="8" pageNumber="95" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B9F36FEFFFF6E13FF461122FD1AFEF1" blockId="8.[151,1006,223,646]" box="[151,729,295,322]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF461122FDCFFEF1" authority="(Paulian, 1937)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1937" box="[151,524,295,321]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="heurni" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF461122FE95FEF1" box="[151,342,295,321]" italics="true" pageId="8" pageNumber="95">Rhytomus heurni</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FEB51122FDC7FEF1" author="Paulian, R." box="[356,516,295,321]" pageId="8" pageNumber="95" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian, 1937</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDC6112DFD1AFEF1" box="[535,729,295,322]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF46114EFD2CFED5" blockId="8.[151,1006,223,646]" box="[151,751,331,358]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF46114EFDE2FED6" authority="(Paulian, 1985)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1985" box="[151,545,331,358]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="howdeni" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF46114EFEAFFED5" box="[151,364,331,357]" italics="true" pageId="8" pageNumber="95">Rhytomus howdeni</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FEAB114EFDDAFED6" author="Paulian, R." box="[378,537,331,358]" pageId="8" pageNumber="95" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDFC1149FD2CFED5" box="[557,751,331,358]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF46116AFD70FE39" blockId="8.[151,1006,223,646]" box="[151,691,367,394]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF46116AFDE2FE39" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,545,367,393]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="larseni" status="sp. nov.">
<emphasis id="B954EAECFFFF6E13FF46116AFE98FE39" box="[151,347,367,393]" italics="true" pageId="8" pageNumber="95">Rhytomus larseni</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDFD1175FD70FE39" box="[556,691,367,394]" pageId="8" pageNumber="95" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF461196FD2AFE1D" blockId="8.[151,1006,223,646]" box="[151,745,403,430]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF461196FD94FE1D" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,599,403,429]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="latidentatus" status="sp. nov.">
<emphasis id="B954EAECFFFF6E13FF461196FE52FE1D" box="[151,401,403,429]" italics="true" pageId="8" pageNumber="95">Rhytomus latidentatus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDB31191FD2AFE1D" box="[610,745,403,430]" pageId="8" pageNumber="95" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF4611B2FD3EFE61" blockId="8.[151,1006,223,646]" box="[151,765,439,466]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF4611B2FDECFE62" authority="(Paulian, 1985)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1985" box="[151,559,439,466]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="papuanus" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF4611B2FEB9FE61" box="[151,378,439,465]" italics="true" pageId="8" pageNumber="95">Rhytomus papuanus</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FE5911B2FDE4FE62" author="Paulian, R." box="[392,551,439,466]" pageId="8" pageNumber="95" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985</bibRefCitation>
)
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,
<taxonomicNameLabel id="A2675797FFFF6E13FDEA11BDFD3EFE61" box="[571,765,439,466]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
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<paragraph id="8B9F36FEFFFF6E13FF4611DEFCC8FDA9" blockId="8.[151,1006,223,646]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF4611DEFD15FE45" authority="(Krikken &amp; Huijbregts, 2009)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Krikken &amp; Huijbregts" baseAuthorityYear="2009" box="[151,726,475,501]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="seramicus" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF4611DEFEBCFE45" box="[151,383,475,501]" italics="true" pageId="8" pageNumber="95">Rhytomus seramicus</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FE5D11DEFD0DFE45" author="Krikken, J. &amp; Huijbregts, J." box="[396,718,475,501]" pageId="8" pageNumber="95" pagination="237 - 266" refId="ref15470" refString="Krikken, J. &amp; Huijbregts, J. (2009) A new Haroldius Boucomont and a new Tesserodon Hope from the Moluccas (Coleoptera: Scarabaeidae: Scarabaeinae). Koleopterologische Rundschau, 79, 237 - 266." type="journal article" year="2009">Krikken &amp; Huijbregts, 2009</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FD3311D9FC60FE45" box="[738,931,475,502]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
<taxonomicName id="4C204D7DFFFF6E13FF4611FAFDFDFDAA" authority="(Balthasar, 1965)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Balthasar" baseAuthorityYear="1965" box="[151,574,511,538]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="setulosus" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF4611FAFEB7FDA9" box="[151,372,511,537]" italics="true" pageId="8" pageNumber="95">Rhytomus setulosus</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FE5311FAFDF6FDAA" author="Balthasar, V." box="[386,565,511,538]" pageId="8" pageNumber="95" pagination="14 - 23" refId="ref14343" refString="Balthasar, V. (1965) Eine neue Untergattung und neue Arten der Familie Scarabaeidae. Acta Entomologica Bohemoslovaca, 62 (1), 14 - 23." type="journal article" year="1965">Balthasar, 1965</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FD9B1205FCC8FDA9" box="[586,779,511,538]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF461226FD77FD8D" blockId="8.[151,1006,223,646]" box="[151,692,547,574]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF461226FDE0FD8D" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,547,547,573]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="sodalis" status="sp. nov.">
<emphasis id="B954EAECFFFF6E13FF461226FE9FFD8D" box="[151,348,547,573]" italics="true" pageId="8" pageNumber="95">Rhytomus sodalis</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDFF1221FD77FD8D" box="[558,692,547,574]" pageId="8" pageNumber="95" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF461242FD65FDD1" blockId="8.[151,1006,223,646]" box="[151,678,583,610]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF461242FDD6FDD1" authority="Saxton &amp; Genier" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[151,533,583,609]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="vanus" status="sp. nov.">
<emphasis id="B954EAECFFFF6E13FF461242FE8DFDD1" box="[151,334,583,609]" italics="true" pageId="8" pageNumber="95">Rhytomus vanus</emphasis>
Saxton &amp; Génier
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FDF1124DFD65FDD1" box="[544,678,583,610]" pageId="8" pageNumber="95" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9F36FEFFFF6E13FF46126EFC6EFD35" blockId="8.[151,1006,223,646]" box="[151,941,619,646]" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FF46126EFD23FD35" authority="(Ochi, Kon &amp; Hartini, 2017)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Ochi, Kon &amp; Hartini" baseAuthorityYear="2017" box="[151,736,619,646]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="waigeoensis" status="comb. nov.">
<emphasis id="B954EAECFFFF6E13FF46126EFE55FD35" box="[151,406,619,645]" italics="true" pageId="8" pageNumber="95">Rhytomus waigeoensis</emphasis>
(
<bibRefCitation id="EFB14B0FFFFF6E13FE75126EFD1BFD35" author="Ochi, T. &amp; Kon, M. &amp; Hartini, S." box="[420,728,619,646]" pageId="8" pageNumber="95" pagination="59 - 66" refId="ref16278" refString="Ochi, T., Kon, M. &amp; Hartini, S. (2017) Descriptions of a new species of the genus Tesserodon and two new species of the genus Temnoplectron from West Papua, Indonesia (Coleoptera, Scarabaeidae, Scarabaeinae). Kogane, 20, 59 - 66." type="journal article" year="2017">Ochi, Kon &amp; Hartini, 2017</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2675797FFFF6E13FD3A1269FC6EFD35" box="[747,941,619,646]" pageId="8" pageNumber="95" rank="species">new combination</taxonomicNameLabel>
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316
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<emphasis id="B954EAECFFFC6E10FF461757FCE7F8DA" bold="true" box="[151,804,1874,1898]" pageId="11" pageNumber="98">
<treatmentCitation id="0A8110EFFFFC6E10FF461757FDFAF8DA" author="Paulian, R." box="[151,569,1874,1898]" page="222" pageId="11" pageNumber="98" year="1985">
<taxonomicName id="4C204D7DFFFC6E10FF461757FDFAF8DA" ID-CoL="55JGP" authority="Paulian, 1985: 222" authorityName="Paulian" authorityPageNumber="222" authorityYear="1985" box="[151,569,1874,1898]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B954EAECFFFC6E10FF461757FEAFF8DA" bold="true" box="[151,364,1874,1898]" italics="true" pageId="11" pageNumber="98">Tesserodon howdeni</emphasis>
<bibRefCitation id="EFB14B0FFFFC6E10FEA31757FDFAF8DA" author="Paulian, R." box="[370,569,1874,1898]" pageId="11" pageNumber="98" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian, 1985: 222</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description)
</emphasis>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFC6E16FF46179DFD5BFED4" lastPageId="13" lastPageNumber="100" pageId="11" pageNumber="98" type="materials_examined">
<paragraph id="8B9F36FEFFFC6E17FF46179DFB63FF65" blockId="11.[151,1437,1944,2043]" lastBlockId="12.[151,1436,151,213]" lastPageId="12" lastPageNumber="99" pageId="11" pageNumber="98">
<emphasis id="B954EAECFFFC6E10FF46179DFE09F803" bold="true" box="[151,458,1944,1971]" pageId="11" pageNumber="98">Name-bearing type data.</emphasis>
<materialsCitation id="3B483CA3FFFC6E17FE07179CFB5AFF64" collectingDate="1974-07-15" collectingDateMax="1974-07-16" collectingDateMin="1974-07-15" collectionCode="CMNC" collectorName="S. Peck" country="Papua N. Guinea" determinerName="R. Paulian" lastPageId="12" lastPageNumber="99" location="Papua N. Guinea" municipality="Musee" pageId="11" pageNumber="98" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549B885CFFFC6E10FE07179CFDFDF803" box="[470,574,1945,1971]" pageId="11" pageNumber="98" type="holotype">Holotype</typeStatus>
<specimenCount id="9D26FD77FFFC6E10FD9B179CFD9CF803" box="[586,607,1945,1971]" count="1" pageId="11" pageNumber="98" type="male">♂</specimenCount>
(
<collectionCode id="ED31AE3BFFFC6E10FDA2179CFD0EF803" box="[627,717,1945,1971]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="11" pageNumber="98">CMNC</collectionCode>
): ||
<collectingCountry id="F337766EFFFC6E10FD2D179CFC7BF803" box="[764,952,1945,1971]" pageId="11" pageNumber="98">Papua N.Guinea</collectingCountry>
| Brown R.
<quantity id="4CD89B1BFFFC6E10FB97179CFB4BF803" box="[1094,1160,1945,1971]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="11" pageNumber="98" unit="mi" value="30.0">30mi.</quantity>
N | Fort Moresby |
<date id="FF9E103EFFFC6E10FAA6179DFEE3F867" pageId="11" pageNumber="98" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">
<collectingDate id="EFDAE9D6FFFC6E10FAA6179DFEE3F867" pageId="11" pageNumber="98" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">15- 16.VII.1974</collectingDate>
</date>
||; ||
<collectorName id="26D55328FFFC6E10FE8B17B8FE73F867" box="[346,432,1981,2007]" pageId="11" pageNumber="98">S. Peck</collectorName>
| collector ||; || H. &amp; A. Howden | collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011854 ||; ||
<taxonomicName id="4C204D7DFFFC6E10FC1A17E4FB89F84B" authorityName="Hope" authorityYear="1837" box="[971,1098,2017,2043]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="genus">Tesserodon</taxonomicName>
| howdeni | n. sp. |
<determinerName id="7E43DA3AFFFC6E10FAF617E4FA5FF84B" box="[1319,1436,2017,2043]" pageId="11" pageNumber="98">R. Paulian</determinerName>
det. || [partly handwritten].
<emphasis id="B954EAECFFFB6E17FE1C1092FDC6FF01" bold="true" box="[461,517,151,177]" pageId="12" pageNumber="99">Note</emphasis>
. Previously stated as: “Nouvelle-Guinée (Papouasie): Brown River,
<quantity id="4CD89B1BFFFB6E17FAC91092FABAFF01" box="[1304,1401,151,177]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="12" pageNumber="99" unit="mi" value="20.0">20 miles</quantity>
N. Port Moresby, 14/
<date id="FF9E103EFFFB6E17FEB110BEFE2DFF65" box="[352,494,187,213]" pageId="12" pageNumber="99" value="1974-07-16">16-VII-1974</date>
(S. Peck), piège à excréments, en forêt” by Paulian.
<emphasis id="B954EAECFFFB6E17FBEC10BFFB5AFF64" bold="true" box="[1085,1177,186,212]" pageId="12" pageNumber="99">Studied</emphasis>
</materialsCitation>
.
</paragraph>
<caption id="DF5F6676FFFB6E17FF4917A2FE30F853" ID-DOI="http://doi.org/10.5281/zenodo.14746462" ID-Zenodo-Dep="14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="12" pageNumber="99" startId="12.[152,255,1959,1983]" targetBox="[155,1431,238,1929]" targetPageId="12" targetType="figure">
<paragraph id="8B9F36FEFFFB6E17FF4917A2FE30F853" blockId="12.[151,1436,1959,2019]" pageId="12" pageNumber="99">
<emphasis id="B954EAECFFFB6E17FF4917A2FED4F80F" bold="true" box="[152,279,1959,1983]" pageId="12" pageNumber="99">FIGURE 9.</emphasis>
Male aedeagus of
<taxonomicName id="4C204D7DFFFB6E17FE0317A2FD44F80F" box="[466,647,1959,1983]" pageId="12" pageNumber="99">
<emphasis id="B954EAECFFFB6E17FE0317A2FDF5F80F" box="[466,566,1959,1983]" italics="true" pageId="12" pageNumber="99">Rhytomus</emphasis>
species
</taxonomicName>
, left lateral, dorsal, right lateral. A,
<taxonomicName id="4C204D7DFFFB6E17FC3617ADFBA6F80F" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="van Lansberge" baseAuthorityYear="1885" box="[999,1125,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B954EAECFFFB6E17FC3617ADFBA6F80F" box="[999,1125,1959,1983]" italics="true" pageId="12" pageNumber="99">R. elongatus</emphasis>
</taxonomicName>
; B,
<taxonomicName id="4C204D7DFFFB6E17FB5F17ADFB38F80F" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[1166,1275,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="fervidus">
<emphasis id="B954EAECFFFB6E17FB5F17ADFB38F80F" box="[1166,1275,1959,1983]" italics="true" pageId="12" pageNumber="99">R. fervidus</emphasis>
</taxonomicName>
; C,
<emphasis id="B954EAECFFFB6E17FAF217ADFA5FF80F" box="[1315,1436,1959,1983]" italics="true" pageId="12" pageNumber="99">
<taxonomicName id="4C204D7DFFFB6E17FAF217ADFA55F80F" authorityName="Paulian" authorityYear="1985" box="[1315,1430,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="howdeni">R. howdeni</taxonomicName>
;
</emphasis>
D,
<emphasis id="B954EAECFFFB6E17FF6717C9FEE3F853" box="[182,288,1995,2019]" italics="true" pageId="12" pageNumber="99">
<taxonomicName id="4C204D7DFFFB6E17FF6717C9FEDFF853" authorityName="Saxton &amp; Genier" authorityYear="2025" box="[182,284,1995,2019]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="99" phylum="Arthropoda" rank="species" species="larseni">R. larseni</taxonomicName>
.
</emphasis>
Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9F36FEFFFA6E16FF161093FD5BFED4" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
<emphasis id="B954EAECFFFA6E16FF161093FE6BFF00" bold="true" box="[199,424,150,176]" pageId="13" pageNumber="100">Material examined</emphasis>
(
<specimenCount id="9D26FD77FFFA6E16FE661093FE34FF01" box="[439,503,150,177]" count="5" pageId="13" pageNumber="100" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D26FD77FFFA6E16FDD31093FDFEFF00" box="[514,573,150,176]" count="2" pageId="13" pageNumber="100" type="female">2 ♀♀</specimenCount>
).
<materialsCitation id="3B483CA3FFFA6E16FD841093FE9AFF48" collectedFrom="forest" collectingDate="1974-07-14" collectingDateMax="1974-07-18" collectingDateMin="1974-07-14" collectingMethod="dung traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.204166" location="Brown River" longLatPrecision="21" longitude="147.2875" pageId="13" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="CENTRAL PROVINCE">
<emphasis id="B954EAECFFFA6E16FD841093FCACFF00" bold="true" box="[597,879,150,176]" pageId="13" pageNumber="100">
<collectingCountry id="F337766EFFFA6E16FD841093FCACFF00" box="[597,879,150,176]" name="Papua New Guinea" pageId="13" pageNumber="100">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<collectingRegion id="49E4F81CFFFA6E16FCAD1093FB52FF00" box="[892,1169,150,176]" country="Papua New Guinea" name="Central" pageId="13" pageNumber="100">CENTRAL PROVINCE</collectingRegion>
,
<locationDeviation id="9A2D2FDCFFFA6E16FB4A1093FA5AFF00" box="[1179,1433,150,177]" location="port Moresby" pageId="13" pageNumber="100">
<quantity id="4CD89B1BFFFA6E16FB4A1093FB22FF00" box="[1179,1249,150,176]" metricMagnitude="4" metricUnit="m" metricValue="2.8968192" pageId="13" pageNumber="100" unit="mi" value="18.0">18 mi.</quantity>
N port Moresby
</locationDeviation>
,
<location id="8EFF6025FFFA6E16FF4610BEFEE9FF64" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FF4610BEFEE9FF64" box="[151,298,186,213]" country="Papua New Guinea" latitude="-9.204166" longLatPrecision="21" longitude="147.2875" name="Brown River" pageId="13" pageNumber="100" stateProvince="CENTRAL PROVINCE">Brown River</location>
,
<quantity id="4CD89B1BFFFA6E16FEE410BFFEA2FF65" box="[309,353,186,213]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">
<elevation id="000DD1CDFFFA6E16FEE410BFFEA2FF65" box="[309,353,186,213]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">3 m</elevation>
</quantity>
, [
<geoCoordinate id="EE145039FFFA6E16FEA510BFFDC3FF65" box="[372,512,186,213]" degrees="09" direction="south" minutes="12" orientation="latitude" pageId="13" pageNumber="100" precision="15" seconds="15" value="-9.204166">09°12’15’’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFFA6E16FDDD10BFFD6BFF65" box="[524,680,186,213]" degrees="147" direction="east" minutes="17" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="15" value="147.2875">147°17’15’’E</geoCoordinate>
],
<date id="FF9E103EFFFA6E16FD6A10BFFCA4FF64" box="[699,871,186,212]" pageId="13" pageNumber="100" value="1974-07-14" valueMax="1974-07-18" valueMin="1974-07-14">
<collectingDate id="EFDAE9D6FFFA6E16FD6A10BFFCA4FF64" box="[699,871,186,212]" pageId="13" pageNumber="100" value="1974-07-14" valueMax="1974-07-18" valueMin="1974-07-14">14–18.vii.1974</collectingDate>
</date>
,
<collectedFrom id="383C7503FFFA6E16FCA310BFFC76FF64" box="[882,949,186,212]" pageId="13" pageNumber="100">forest</collectedFrom>
,
<collectingMethod id="52614EE9FFFA6E16FC6E10BFFBF4FF65" box="[959,1079,186,213]" pageId="13" pageNumber="100">dung traps</collectingMethod>
,
<collectorName id="26D55328FFFA6E16FB9310BFFB5BFF64" box="[1090,1176,186,212]" pageId="13" pageNumber="100">S. Peck</collectorName>
, (dung traps 39-42)—
<specimenCount id="9D26FD77FFFA6E16FA4210BFFF6DFF49" count="1" pageId="13" pageNumber="100" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D26FD77FFFA6E16FF6D10DBFF24FF49" box="[188,231,222,249]" count="1" pageId="13" pageNumber="100" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED31AE3BFFFA6E16FF2910DBFE92FF48" box="[248,337,222,248]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="100">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CA3FFFA6E16FEB410DBFD13FEAC" collectingDate="1974-07-16" collectingDateMax="1974-07-18" collectingDateMin="1974-07-16" collectingMethod="dung traps" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.191667" location="Brown River" longLatPrecision="21" longitude="147.31667" pageId="13" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="National Capital">
<locationDeviation id="9A2D2FDCFFFA6E16FEB410DBFDB3FF48" box="[357,624,222,249]" location="Port Moresby" pageId="13" pageNumber="100">
<quantity id="4CD89B1BFFFA6E16FEB410DBFE6CFF48" box="[357,431,222,248]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="13" pageNumber="100" unit="mi" value="20.0">20 mi.</quantity>
N Port Moresby
</locationDeviation>
,
<location id="8EFF6025FFFA6E16FDAD10DAFCD2FF48" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FDAD10DAFCD2FF48" box="[636,785,222,249]" country="Papua New Guinea" latitude="-9.191667" longLatPrecision="21" longitude="147.31667" name="Brown River" pageId="13" pageNumber="100" stateProvince="National Capital">Brown River</location>
,
<quantity id="4CD89B1BFFFA6E16FCCF10DBFC8FFF49" box="[798,844,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">
<elevation id="000DD1CDFFFA6E16FCCF10DBFC8FFF49" box="[798,844,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="100" unit="m" value="3.0">3 m</elevation>
</quantity>
, [
<geoCoordinate id="EE145039FFFA6E16FCB010DBFC2FFF48" box="[865,1004,222,248]" degrees="09" direction="south" minutes="11" orientation="latitude" pageId="13" pageNumber="100" precision="15" seconds="30" value="-9.191667">09°11’30’’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFFA6E16FC2B10DBFB55FF48" box="[1018,1174,222,248]" degrees="147" direction="east" minutes="19" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="00" value="147.31667">147°19’00’’E</geoCoordinate>
],
<date id="FF9E103EFFFA6E16FB7A10DBFA94FF48" box="[1195,1367,222,248]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">
<collectingDate id="EFDAE9D6FFFA6E16FB7A10DBFA94FF48" box="[1195,1367,222,248]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">16–18.vii.1974</collectingDate>
</date>
,
<collectingMethod id="52614EE9FFFA6E16FAB510DBFF13FEAD" pageId="13" pageNumber="100">dung traps</collectingMethod>
,
<collectorName id="26D55328FFFA6E16FF0C1107FEF5FEAC" box="[221,310,258,284]" pageId="13" pageNumber="100">S. Peck</collectorName>
, (dung traps 39-41)—
<specimenCount id="9D26FD77FFFA6E16FDE61107FD9DFEAD" box="[567,606,258,285]" count="1" pageId="13" pageNumber="100" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED31AE3BFFFA6E16FDBE1107FD0BFEAC" box="[623,712,258,284]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="100">CMNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CA3FFFA6E16FD0C1106FDDDFEF0" collectingDate="1974-07-16" collectingDateMax="1974-07-18" collectingDateMin="1974-07-16" collectingMethod="dung trap" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" elevation="3" latitude="-9.191667" location="Brown River" longLatPrecision="21" longitude="147.31667" pageId="13" pageNumber="100" specimenCount="2" specimenCount-female="1" specimenCount-male="2" stateProvince="Central" typeStatus="paratype">
same locality,
<date id="FF9E103EFFFA6E16FC501107FBEEFEAC" box="[897,1069,258,284]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">
<collectingDate id="EFDAE9D6FFFA6E16FC501107FBEEFEAC" box="[897,1069,258,284]" pageId="13" pageNumber="100" value="1974-07-16" valueMax="1974-07-18" valueMin="1974-07-16">16–18.vii.1974</collectingDate>
</date>
,
<collectingMethod id="52614EE9FFFA6E16FBEB1107FB6AFEAD" box="[1082,1193,258,285]" pageId="13" pageNumber="100">dung trap</collectingMethod>
,
<collectorName id="26D55328FFFA6E16FB671107FACCFEAC" box="[1206,1295,258,284]" pageId="13" pageNumber="100">S. Peck</collectorName>
, (dung traps 39-41)—
<specimenCount id="9D26FD77FFFA6E16FF2F1123FEFFFEF1" box="[254,316,294,321]" count="2" pageId="13" pageNumber="100" type="male">2 ♂♂</specimenCount>
<typeStatus id="549B885CFFFA6E16FE901122FE6FFEF1" box="[321,428,295,321]" pageId="13" pageNumber="100" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED31AE3BFFFA6E16FE6B1123FDD5FEF0" box="[442,534,294,320]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="100">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CA3FFFA6E16FDF91123FE4BFED4" collectingDate="1974-07-15" collectingDateMax="1974-07-16" collectingDateMin="1974-07-15" collectorName="S. Peck" country="Papua New Guinea" latitude="-9.183333" location="Brown River" longLatPrecision="925" longitude="147.34166" pageId="13" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="National Capital" typeStatus="holotype">
<locationDeviation id="9A2D2FDCFFFA6E16FDF91123FCE4FEF0" box="[552,807,294,321]" location="Port Moresby" pageId="13" pageNumber="100">
<quantity id="4CD89B1BFFFA6E16FDF91123FDACFEF0" box="[552,623,294,320]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="13" pageNumber="100" unit="mi" value="30.0">30 mi.</quantity>
N Port Moresby
</locationDeviation>
,
<location id="8EFF6025FFFA6E16FCE01122FC00FEF0" LSID="urn:lsid:plazi:treatment:038987E8FFFC6E16FF461757FAABFB10:8EFF6025FFFA6E16FCE01122FC00FEF0" box="[817,963,294,321]" country="Papua New Guinea" latitude="-9.183333" longLatPrecision="925" longitude="147.34166" name="Brown River" pageId="13" pageNumber="100" stateProvince="National Capital">Brown River</location>
, [
<geoCoordinate id="EE145039FFFA6E16FC051123FBF1FEF0" box="[980,1074,294,320]" degrees="09" direction="south" minutes="11" orientation="latitude" pageId="13" pageNumber="100" precision="925" value="-9.183333">09°11’S</geoCoordinate>
,
<geoCoordinate id="EE145039FFFA6E16FBEC1123FB19FEF0" box="[1085,1242,294,320]" degrees="147" direction="east" minutes="20" orientation="longitude" pageId="13" pageNumber="100" precision="15" seconds="30" value="147.34166">147°20’30’’E</geoCoordinate>
],
<date id="FF9E103EFFFA6E16FB3D1123FA5BFEF1" box="[1260,1432,294,321]" pageId="13" pageNumber="100" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">
<collectingDate id="EFDAE9D6FFFA6E16FB3D1123FA5BFEF1" box="[1260,1432,294,321]" pageId="13" pageNumber="100" value="1974-07-15" valueMax="1974-07-16" valueMin="1974-07-15">15–16.vii.1974</collectingDate>
</date>
,
<collectorName id="26D55328FFFA6E16FF46114FFF2DFED4" box="[151,238,330,356]" pageId="13" pageNumber="100">S. Peck</collectorName>
—
<specimenCount id="9D26FD77FFFA6E16FEDB114FFEDFFED4" box="[266,284,330,356]" count="1" pageId="13" pageNumber="100" type="male">♂</specimenCount>
holotype
</materialsCitation>
,
<materialsCitation id="3B483CA3FFFA6E16FE42114FFD57FED4" box="[403,660,330,357]" collectionCode="CMNC" country="Papua New Guinea" pageId="13" pageNumber="100" specimenCount="1" specimenCount-male="1" stateProvince="Central" typeStatus="paratype">
<specimenCount id="9D26FD77FFFA6E16FE42114FFE7EFED5" box="[403,445,330,357]" count="1" pageId="13" pageNumber="100" type="male">1 ♂</specimenCount>
<typeStatus id="549B885CFFFA6E16FE15114EFDE7FED5" box="[452,548,331,357]" pageId="13" pageNumber="100" type="paratype">paratype</typeStatus>
(
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFA6E16FF16116BFD85FC30" pageId="13" pageNumber="100" type="description">
<paragraph id="8B9F36FEFFFA6E16FF16116BFD85FC30" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
<emphasis id="B954EAECFFFA6E16FF16116BFDF0FE38" bold="true" box="[199,563,366,392]" pageId="13" pageNumber="100">Redescription. Measurements.</emphasis>
Body length
<quantity id="4CD89B1BFFFA6E16FD1F116BFC9BFE39" box="[718,856,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="4.1" metricValueMin="3.6" pageId="13" pageNumber="100" unit="mm" value="3.8499999999999996" valueMax="4.1" valueMin="3.6">3.6–4.1 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFFA6E16FBB7116BFB32FE39" box="[1126,1265,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="13" pageNumber="100" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
.
<emphasis id="B954EAECFFFA6E16FB2C116BFA83FE38" bold="true" box="[1277,1344,366,392]" pageId="13" pageNumber="100">Body.</emphasis>
Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex.
<emphasis id="B954EAECFFFA6E16FB951197FB48FE1C" bold="true" box="[1092,1163,402,428]" pageId="13" pageNumber="100">Head.</emphasis>
Clypeal edge lateral to clypeal teeth emarginate (
<figureCitation id="131B2A7BFFFA6E16FE6F11B3FDD9FE60" box="[446,538,438,464]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="13" pageNumber="100">Fig. 4D</figureCitation>
), head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.6 times eye width.
<emphasis id="B954EAECFFFA6E16FBA311DFFB31FE44" bold="true" box="[1138,1266,474,500]" pageId="13" pageNumber="100">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFFA6E16FBFC1227FB42FD8C" bold="true" box="[1069,1153,546,572]" pageId="13" pageNumber="100">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures.
<emphasis id="B954EAECFFFA6E16FC5312B7FC02FD7C" bold="true" box="[898,961,690,716]" pageId="13" pageNumber="100">Legs.</emphasis>
Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge.
<emphasis id="B954EAECFFFA6E16FF0C12FFFE94FCA4" bold="true" box="[221,343,762,788]" pageId="13" pageNumber="100">Pygidium.</emphasis>
Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (
<figureCitation id="131B2A7BFFFA6E16FAE512FFFA4DFCA4" box="[1332,1422,762,788]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="13" pageNumber="100">Fig. 8C</figureCitation>
), pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex.
<emphasis id="B954EAECFFFA6E16FCBB1347FC27FCEC" bold="true" box="[874,996,834,860]" pageId="13" pageNumber="100">Aedeagus.</emphasis>
Left paramere apex truncated apically, distinctly angular externally (
<figureCitation id="131B2A7BFFFA6E16FE301363FDFAFC30" box="[481,569,870,896]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="13" pageNumber="100">Fig. 9C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFA6E16FF16138FFAABFB10" pageId="13" pageNumber="100" type="discussion">
<paragraph id="8B9F36FEFFFA6E16FF16138FFAABFB10" blockId="13.[151,1437,150,1185]" pageId="13" pageNumber="100">
<emphasis id="B954EAECFFFA6E16FF16138FFEF7FC14" bold="true" box="[199,308,906,932]" pageId="13" pageNumber="100">Remarks</emphasis>
. Examination of the
<typeStatus id="549B885CFFFA6E16FDF9138FFD41FC14" box="[552,642,906,932]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
revealed that the specimen labeled by Paulian is actually a male and is here considered a male
<typeStatus id="549B885CFFFA6E16FE1913AAFDEBFC79" box="[456,552,943,969]" pageId="13" pageNumber="100" type="paratype">paratype</typeStatus>
in the material examined. Additionally, the date ranges provided by
<bibRefCitation id="EFB14B0FFFFA6E16FA9913ABFF21FC5D" author="Paulian, R." pageId="13" pageNumber="100" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian (1985)</bibRefCitation>
in the original description do not match any label data from the specimens studied. Recent examination of the specimens at the MNHN (by FG) confirmed that both are males, indicating that the female specimen described by Paulian is either missing or that Paulian misidentified the gender of the
<typeStatus id="549B885CFFFA6E16FC38141FFB84FB84" box="[1001,1095,1050,1076]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
.
<specimenCount id="9D26FD77FFFA6E16FB83141EFB14FB84" box="[1106,1239,1050,1077]" count="2" pageId="13" pageNumber="100" type="female">Two female</specimenCount>
specimens in the CMNC are not labeled as
<typeStatus id="549B885CFFFA6E16FE13143AFDF2FBE9" box="[450,561,1087,1113]" pageId="13" pageNumber="100" type="paratype">paratypes</typeStatus>
. While one of these could theoretically be the
<typeStatus id="549B885CFFFA6E16FB94143BFB61FBE8" box="[1093,1186,1086,1112]" pageId="13" pageNumber="100" type="allotype">allotype</typeStatus>
, it is unclear if Henry Howden ever sent them to Paulian for study, as he often retained some specimens as a precaution against parcels being lost in the mail. Consequently, these specimens cannot be definitively considered part of the type series.
</paragraph>
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</treatment>
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190
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<paragraph id="8B9F36FEFFFD6E11FF46173AFCCFF8EA" blockId="10.[151,780,1855,1919]" box="[151,780,1855,1882]" pageId="10" pageNumber="97">
<heading id="D0D78192FFFD6E11FF46173AFCCFF8EA" bold="true" box="[151,780,1855,1882]" fontSize="11" level="1" pageId="10" pageNumber="97" reason="1">
<emphasis id="B954EAECFFFD6E11FF46173AFCCFF8EA" bold="true" box="[151,780,1855,1882]" pageId="10" pageNumber="97">
<taxonomicName id="4C204D7DFFFD6E11FF46173AFDEDF8EA" authority="(Paulian, 1937)" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="Paulian" baseAuthorityYear="1937" box="[151,558,1855,1882]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="97" phylum="Arthropoda" rank="species" species="heurni" status="comb. nov.">
<emphasis id="B954EAECFFFD6E11FF46173AFEA8F8EA" bold="true" box="[151,363,1855,1882]" italics="true" pageId="10" pageNumber="97">Rhytomus heurni</emphasis>
(
<bibRefCitation id="EFB14B0FFFFD6E11FEAB1745FDE5F8EA" author="Paulian, R." box="[378,550,1856,1882]" pageId="10" pageNumber="97" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian, 1937</bibRefCitation>
)
</taxonomicName>
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<paragraph id="8B9F36FEFFFD6E11FF461761FE91F8CE" blockId="10.[151,780,1855,1919]" box="[151,338,1892,1919]" pageId="10" pageNumber="97">
<figureCitation id="131B2A7BFFFD6E11FF461761FF2EF8CF" box="[151,237,1892,1919]" captionStart="FIGURE 7" captionStartId="10.[152,255,1714,1738]" captionTargetBox="[156,1425,184,1681]" captionTargetId="figure-22@10.[151,1435,179,1688]" captionTargetPageId="10" captionText="FIGURE 7. Habitus of Rhytomus species in mainland New Guinea. A, R. fervidus; B, R. heurni; C, R. howdeni; D, R. larseni; E, R. latidentatus; F, R. papuanus; G, R. setulosum; H, R. sodalis; I, R. vanus; Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14746458" httpUri="https://zenodo.org/record/14746458/files/figure.png" pageId="10" pageNumber="97">Fig. 7B</figureCitation>
,
<figureCitation id="131B2A7BFFFD6E11FF281761FEDFF8CE" box="[249,284,1892,1918]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="10" pageNumber="97">8B</figureCitation>
,
<figureCitation id="131B2A7BFFFD6E11FEF91761FE91F8CE" box="[296,338,1892,1918]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="10" pageNumber="97">11F</figureCitation>
</paragraph>
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<subSubSection id="C33A6575FFFD6E11FF4617ADFE31F86F" pageId="10" pageNumber="97" type="reference_group">
<paragraph id="8B9F36FEFFFD6E11FF4617ADFE31F86F" blockId="10.[151,1436,1960,2015]" pageId="10" pageNumber="97">
<treatmentCitationGroup id="AB3011D0FFFD6E11FF4617ADFE31F86F" pageId="10" pageNumber="97">
<emphasis id="B954EAECFFFD6E11FF4617ADFCEFF870" bold="true" box="[151,812,1960,1984]" pageId="10" pageNumber="97">
<treatmentCitation id="0A8110EFFFFD6E11FF4617ADFDE4F870" author="Paulian, R." box="[151,551,1960,1984]" page="122" pageId="10" pageNumber="97" year="1937">
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<emphasis id="B954EAECFFFD6E11FF4617ADFE9AF870" bold="true" box="[151,345,1960,1984]" italics="true" pageId="10" pageNumber="97">Tesserodon heurni</emphasis>
<bibRefCitation id="EFB14B0FFFFD6E11FEB117ADFDE4F870" author="Paulian, R." box="[352,551,1960,1984]" pageId="10" pageNumber="97" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian, 1937: 122</bibRefCitation>
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(original description)—
</emphasis>
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<bibRefCitation id="EFB14B0FFFFD6E11FCFD17ADFC26F871" author="Paulian, R." box="[812,997,1960,1985]" pageId="10" pageNumber="97" pagination="219 - 238" refId="ref16422" refString="Paulian, R. (1985) Les coleopteres Scarabaeidae Canthonides de Nouvelle-Guinee. Annales de la Societe Entomologique de France, Nouvelle Serie, 21 (2), 219 - 238. https: // doi. org / 10.1080 / 21686351.1985.12278409" type="journal article" year="1985">Paulian 1985: 221</bibRefCitation>
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(diagnosis, distribution, comment); Gunter
<emphasis id="B954EAECFFFD6E11FF1617CDFF37F86F" box="[199,244,1991,2015]" italics="true" pageId="10" pageNumber="97">et al</emphasis>
. 2019: 453 (distribution)
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</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFC6E10FF461093FE54FEAD" pageId="11" pageNumber="98" type="materials_examined">
<paragraph id="8B9F36FEFFFC6E10FF461093FE54FEAD" blockId="11.[151,1437,150,861]" pageId="11" pageNumber="98">
<emphasis id="B954EAECFFFC6E10FF461093FE0BFF01" bold="true" box="[151,456,150,177]" pageId="11" pageNumber="98">Name-bearing type data.</emphasis>
<materialsCitation id="3B483CA3FFFC6E10FE051092FE54FEAD" collectingDate="1920-10" collectionCode="SDEI" collectorName="W. C. van Heurn" country="N. Guinea" determinerName="R. Paulian" elevation="1410" location="N. Guinea" pageId="11" pageNumber="98" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="549B885CFFFC6E10FE051092FDFFFF01" box="[468,572,151,177]" pageId="11" pageNumber="98" type="holotype">Holotype</typeStatus>
<specimenCount id="9D26FD77FFFC6E10FD991092FD9EFF01" box="[584,605,151,177]" count="1" pageId="11" pageNumber="98" type="female">♀</specimenCount>
(
<collectionCode id="ED31AE3BFFFC6E10FDA11092FD76FF01" box="[624,693,151,177]" pageId="11" pageNumber="98">SDEI</collectionCode>
): ||
<collectingCountry id="F337766EFFFC6E10FD321092FC98FF01" box="[739,859,151,177]" pageId="11" pageNumber="98">N. Guinea</collectingCountry>
Exped. |
<collectorName id="26D55328FFFC6E10FC011092FB4DFF01" box="[976,1166,151,177]" pageId="11" pageNumber="98">W.C. van Heurn</collectorName>
|
<quantity id="4CD89B1BFFFC6E10FB7A1092FAC4FF01" box="[1195,1287,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="11" pageNumber="98" unit="m" value="1410.0">
<elevation id="000DD1CDFFFC6E10FB7A1092FAC4FF01" box="[1195,1287,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.41" pageId="11" pageNumber="98" unit="m" value="1410.0">1410 m</elevation>
</quantity>
.
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<collectingDate id="EFDAE9D6FFFC6E10FAC61092FA48FF01" box="[1303,1419,151,177]" pageId="11" pageNumber="98" value="1920-10">Okt. 1920</collectingDate>
</date>
| Dormanpadbivak ||; || Holotypus || [red card]; || DEI Müncheberg | Col-05059 || [blue card]; || WORLD SCARAB. | DATABASE | WSD00028176 || [QR code label]; ||
<taxonomicName id="4C204D7DFFFC6E10FD0810DAFC9BFF49" ID-CoL="63V9L" authorityName="Hope" authorityYear="1837" box="[729,856,223,249]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="98" phylum="Arthropoda" rank="genus">Tesserodon</taxonomicName>
| Heurni
<taxonomicNameLabel id="A2675797FFFC6E10FC1310DAFC3CFF49" box="[962,1023,223,249]" pageId="11" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
Type | det.
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R.
<bibRefCitation id="EFB14B0FFFFC6E10FB7510DAFAF4FF49" author="Paulian, R." box="[1188,1335,223,249]" pageId="11" pageNumber="98" pagination="121 - 122" refId="ref16378" refString="Paulian, R. (1937) Deux nouveaux Canthonides de la region australienne. Bulletin de la Societe Entomologique de France, 42 (8), 121 - 122. https: // doi. org / 10.3406 / bsef. 1937.15140" type="journal article" year="1937">Paulian 1937</bibRefCitation>
</determinerName>
|| [partly handwritten].
<emphasis id="B954EAECFFFC6E10FEE51107FE53FEAC" bold="true" box="[308,400,258,284]" pageId="11" pageNumber="98">Studied</emphasis>
.
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</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFC6E10FF161123FCB2FD7D" pageId="11" pageNumber="98" type="description">
<paragraph id="8B9F36FEFFFC6E10FF161123FCB2FD7D" blockId="11.[151,1437,150,861]" pageId="11" pageNumber="98">
<emphasis id="B954EAECFFFC6E10FF161123FEAEFEF0" bold="true" box="[199,365,294,320]" pageId="11" pageNumber="98">Redescription</emphasis>
.
<emphasis id="B954EAECFFFC6E10FEAE1122FDF5FEF1" bold="true" box="[383,566,295,321]" pageId="11" pageNumber="98">Measurements.</emphasis>
Body length
<quantity id="4CD89B1BFFFC6E10FD091122FCF7FEF1" box="[728,820,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="11" pageNumber="98" unit="mm" value="4.3">4.3 mm</quantity>
, maximum body width
<quantity id="4CD89B1BFFFC6E10FB9D1122FB6AFEF1" box="[1100,1193,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="98" unit="mm" value="3.1">3.1 mm</quantity>
.
<emphasis id="B954EAECFFFC6E10FB661122FB38FEF1" bold="true" box="[1207,1275,295,321]" pageId="11" pageNumber="98">Body.</emphasis>
Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex.
<emphasis id="B954EAECFFFC6E10FC14114EFBC8FED5" bold="true" box="[965,1035,331,357]" pageId="11" pageNumber="98">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 4.1 times eye width.
<emphasis id="B954EAECFFFC6E10FC6A1196FBF8FE1D" bold="true" box="[955,1083,403,429]" pageId="11" pageNumber="98">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.8, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFFC6E10FCAF11DEFC12FE45" bold="true" box="[894,977,475,501]" pageId="11" pageNumber="98">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined ocellate punctures (
<figureCitation id="131B2A7BFFFC6E10FB691242FADBFDD1" box="[1208,1304,583,609]" captionStart="FIGURE 11" captionStartId="15.[152,255,1503,1527]" captionTargetBox="[151,1435,179,1477]" captionTargetId="figure-17@15.[151,1435,179,1477]" captionTargetPageId="15" captionText="FIGURE 11. Characters used for species identification. A, dorsal elytral striae showing unaligned scattered punctures (R. fervidus); B, dorsal elytral striae showing two rows of punctuation on intervals 3–5 (R. larseni); C, basal pseudepipleural pit large and deep (R. larseni); D, basal pseudepipleural pit shallow and slightly larger than stria 10 (R. fervidus); E, basal pseudepipleural surface with irregularly distributed punctuation (R. fervidus); F, basal pseudepipleural surface with ocellate punctuation (R. heurni)." figureDoi="http://doi.org/10.5281/zenodo.14746468" httpUri="https://zenodo.org/record/14746468/files/figure.png" pageId="11" pageNumber="98">Fig. 11F</figureCitation>
), interstriae 3–5 on basal fifth with scattered unaligned punctures.
<emphasis id="B954EAECFFFC6E10FCD0126EFC83FD35" bold="true" box="[769,832,619,645]" pageId="11" pageNumber="98">Legs.</emphasis>
Profemoral anterior surface flat.
<emphasis id="B954EAECFFFC6E10FB6D126FFAF4FD34" bold="true" box="[1212,1335,618,644]" pageId="11" pageNumber="98">Pygidium.</emphasis>
Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex (
<figureCitation id="131B2A7BFFFC6E10FCDD12B6FCA7FD7D" box="[780,868,691,717]" captionStart="FIGURE 8" captionStartId="11.[152,255,1592,1616]" captionTargetBox="[154,1432,921,1562]" captionTargetId="figure-331@11.[151,1435,918,1565]" captionTargetPageId="11" captionText="FIGURE 8. Pygidium of Rhytomus species. A, male R. fervidus; B. female R. heurni; C, male R. howdeni; D, male R. larseni; E, male R. latidentatus; F, male R. papuanus; G, female R. setulosus; H, male R. sodalis; I, male R. vanus. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14746460" httpUri="https://zenodo.org/record/14746460/files/figure.png" pageId="11" pageNumber="98">Fig. 8B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFC6E10FF1612D2FC56FCED" pageId="11" pageNumber="98" type="discussion">
<paragraph id="8B9F36FEFFFC6E10FF1612D2FC56FCED" blockId="11.[151,1437,150,861]" pageId="11" pageNumber="98">
<emphasis id="B954EAECFFFC6E10FF1612D2FEF8FD41" bold="true" box="[199,315,727,753]" pageId="11" pageNumber="98">Remarks.</emphasis>
The original description of this species is based on a single female specimen collected during W. C. van Heurn New
<collectingCountry id="F337766EFFFC6E10FEAA12FEFE0FFCA5" box="[379,460,763,789]" name="Guinea-Bissau" pageId="11" pageNumber="98">Guinea</collectingCountry>
expedition in
<date id="FF9E103EFFFC6E10FDA512FEFCD0FCA5" box="[628,787,763,789]" pageId="11" pageNumber="98" value="1920-10">October 1920</date>
. The
<typeStatus id="549B885CFFFC6E10FC8412FEFC46FCA5" box="[853,901,763,789]" pageId="11" pageNumber="98">type</typeStatus>
locality “Dormanpadbivak” roughly translates to “Doorman path bivouac.” According to
<bibRefCitation id="EFB14B0FFFFC6E10FD58131AFC13FC89" author="Holthuis, L. B. &amp; Husson, A. M." box="[649,976,799,825]" pageId="11" pageNumber="98" pagination="3 - 67" refId="ref15225" refString="Holthuis, L. B. &amp; Husson, A. M. (1973) Jonkheer Drs. Willem Cornelis van Heurn (1887 - 1972). Zoologische Bijdragen, 16 (1), 3 - 67." type="journal article" year="1973">Holthuis and Husson (1973)</bibRefCitation>
this locality was approximating
<geoCoordinate id="EE145039FFFC6E10FA80131AFA5FFC89" box="[1361,1436,799,825]" degrees="3" direction="south" minutes="24" orientation="latitude" pageId="11" pageNumber="98" precision="925" value="-3.4">3º24’S</geoCoordinate>
<geoCoordinate id="EE145039FFFC6E10FF461346FEC3FCED" box="[151,256,835,861]" degrees="138" direction="east" minutes="38" orientation="longitude" pageId="11" pageNumber="98" precision="925" value="138.63333">138º38’E</geoCoordinate>
at
<quantity id="4CD89B1BFFFC6E10FEF21347FE77FCED" box="[291,436,834,861]" metricMagnitude="3" metricUnit="m" metricValue="1.43" metricValueMax="1.45" metricValueMin="1.41" pageId="11" pageNumber="98" unit="m" value="1430.0" valueMax="1450.0" valueMin="1410.0">1410-1450m</quantity>
, which is about
<quantity id="4CD89B1BFFFC6E10FDBB1347FD69FCED" box="[618,682,834,861]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="98" unit="km" value="25.0">25km</quantity>
S. of Prauwenbivak.
</paragraph>
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</treatment>
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90
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(
<collectionCode id="ED31AE3BFFFF6E13FD8713B2FD69FC61" box="[598,682,951,977]" collectionName="Italy, Genova, Museo Civico di Storia Naturale &quot;Giacomo Doria&quot;" pageId="8" pageNumber="95">MSNG</collectionCode>
) by present designation: || Nuova Guinea | Fly River | L.M.D Albertis 1876-77 ||; || TYPUS || [bordered and printed in red]; ||
<taxonomicName id="4C204D7DFFFF6E13FCD113DEFCBCFC45" ID-CoL="63V9L" authorityName="Hope" authorityYear="1837" box="[768,895,987,1013]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="genus">Tesserodon</taxonomicName>
| elongatum | Lansberge Type || [handwritten]; || elongatum | Lansb. || [handwritten with a border]; || SYNTYPUS |
<taxonomicName id="4C204D7DFFFE6E12FC701092FBE3FF01" ID-CoL="63V9L" authorityName="Hope" authorityYear="1837" box="[929,1056,151,177]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">Tesserodon</taxonomicName>
| elongatum |
<bibRefCitation id="EFB14B0FFFFE6E12FB1B1092FA44FF01" author="Van Lansberge, J. W." box="[1226,1415,150,177]" pageId="9" pageNumber="96" pagination="375 - 400" refId="ref16891" refString="Van Lansberge, J. W. (1885) Description d'especes nouvelles de coleopteres appartenant au Musee civique de Genes. Annali del Museo civico di storia naturale di Genova, 2 (2), 375 - 400." type="journal article" year="1885">Lansberge, 1885</bibRefCitation>
|| [partly handwritten]; || WORLD SCARAB. | DATABASE | WSD00028173 || [QR code label]; || LECTOTYPE ♂ |
<taxonomicName id="4C204D7DFFFE6E12FF4610DAFED7FF49" ID-CoL="63V9L" authorityName="Hope" authorityYear="1837" box="[151,276,223,249]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAECFFFE6E12FF4610DAFED7FF49" box="[151,276,223,249]" italics="true" pageId="9" pageNumber="96">Tesserodon</emphasis>
</taxonomicName>
|
<taxonomicName id="4C204D7DFFFE6E12FEF910DAFE5DFF49" ID-CoL="55JGF" authorityName="van Lansberge" authorityYear="1885" box="[296,414,223,249]" class="Insecta" family="Scarabaeidae" genus="Tesserodon" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="elongatum">
<emphasis id="B954EAECFFFE6E12FEF910DAFE5DFF49" box="[296,414,223,249]" italics="true" pageId="9" pageNumber="96">elongatum</emphasis>
</taxonomicName>
|
<bibRefCitation id="EFB14B0FFFFE6E12FE6310DAFD5FFF49" author="Van Lansberge, J. W." box="[434,668,222,249]" pageId="9" pageNumber="96" pagination="375 - 400" refId="ref16891" refString="Van Lansberge, J. W. (1885) Description d'especes nouvelles de coleopteres appartenant au Musee civique de Genes. Annali del Museo civico di storia naturale di Genova, 2 (2), 375 - 400." type="journal article" year="1885">van Lansberge, 1885</bibRefCitation>
| des.
<determinerName id="7E43DA3AFFFE6E12FD3210DAFC61FF49" box="[739,930,223,249]" pageId="9" pageNumber="96">Génier &amp; Larsen</determinerName>
, 2015 || [red card].
<emphasis id="B954EAECFFFE6E12FBA810DBFB16FF48" bold="true" box="[1145,1237,222,248]" pageId="9" pageNumber="96">Studied</emphasis>
.
</materialsCitation>
</paragraph>
<caption id="DF5F6676FFFF6E13FF4917E4FB9EF849" ID-DOI="http://doi.org/10.5281/zenodo.14746454" ID-Zenodo-Dep="14746454" box="[152,1117,2017,2041]" httpUri="https://zenodo.org/record/14746454/files/figure.png" pageId="8" pageNumber="95" startId="8.[152,255,2017,2041]" targetBox="[151,1436,1071,1991]" targetPageId="8" targetType="figure">
<paragraph id="8B9F36FEFFFF6E13FF4917E4FB9EF849" blockId="8.[152,1117,2017,2041]" box="[152,1117,2017,2041]" pageId="8" pageNumber="95">
<emphasis id="B954EAECFFFF6E13FF4917E4FEDBF849" bold="true" box="[152,280,2017,2041]" pageId="8" pageNumber="95">FIGURE 6.</emphasis>
Lectotype of
<emphasis id="B954EAECFFFF6E13FE7717E4FDB9F849" box="[422,634,2017,2041]" italics="true" pageId="8" pageNumber="95">
<taxonomicName id="4C204D7DFFFF6E13FE7717E4FDB5F849" authorityName="Génier &amp; Gunter &amp; Saxton" authorityYear="2025" baseAuthorityName="van Lansberge" baseAuthorityYear="1885" box="[422,630,2017,2041]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="95" phylum="Arthropoda" rank="species" species="elongatus">Rhytomus elongatus</taxonomicName>
.
</emphasis>
A, dorsal habitus; B, dorsal head; C, label data.
</paragraph>
</caption>
<paragraph id="8B9F36FEFFFE6E12FF161107FB8BFEF1" blockId="9.[151,1437,150,897]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF161107FE73FEAC" bold="true" box="[199,432,258,284]" pageId="9" pageNumber="96">Material examined</emphasis>
(
<specimenCount id="9D26FD77FFFE6E12FE161106FE28FEAD" box="[455,491,259,285]" count="1" pageId="9" pageNumber="96" type="male">1♂</specimenCount>
,
<specimenCount id="9D26FD77FFFE6E12FE2C1106FDDDFEAD" box="[509,542,259,285]" count="1" pageId="9" pageNumber="96" type="female">1♀</specimenCount>
).
<emphasis id="B954EAECFFFE6E12FDEF1107FCA5FEAC" bold="true" box="[574,870,258,285]" pageId="9" pageNumber="96">
<collectingCountry id="F337766EFFFE6E12FDEF1107FCA5FEAC" box="[574,870,258,285]" name="Papua New Guinea" pageId="9" pageNumber="96">PAPUA NEW GUINEA</collectingCountry>
</emphasis>
:
<collectingRegion id="49E4F81CFFFE6E12FCAB1106FB58FEAD" box="[890,1179,259,285]" country="Papua New Guinea" name="Western" pageId="9" pageNumber="96">WESTERN PROVINCE</collectingRegion>
,
<location id="8EFF6025FFFE6E12FB7D1106FADCFEAD" LSID="urn:lsid:plazi:treatment:038987E8FFFF6E12FF4612CAFDC2FC31:8EFF6025FFFE6E12FB7D1106FADCFEAD" box="[1196,1311,259,285]" country="Papua New Guinea" name="Fly River" pageId="9" pageNumber="96" stateProvince="Western">Fly River</location>
, [
<geoCoordinate id="EE145039FFFE6E12FAE61106FA5BFEAD" box="[1335,1432,259,285]" degrees="08" direction="south" minutes="00" orientation="latitude" pageId="9" pageNumber="96" precision="925" value="-8.0">
<geoCoordinate id="EE145039FFFE6E12FAE61106FA4BFEAD" box="[1335,1416,259,285]" degrees="08" direction="south" minutes="00" orientation="latitude" pageId="9" pageNumber="96" precision="925" value="-8.0">08°00’</geoCoordinate>
<collectingRegion id="49E4F81CFFFE6E12FA591106FA5BFEAD" box="[1416,1432,259,285]" country="Papua New Guinea" name="S" pageId="9" pageNumber="96">S</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="EE145039FFFE6E12FF461122FECBFEF1" box="[151,264,295,321]" degrees="142" direction="east" minutes="00" orientation="longitude" pageId="9" pageNumber="96" precision="925" value="142.0">142°00’E</geoCoordinate>
], 1876–1877,
<collectorName id="26D55328FFFE6E12FE771122FD9DFEF1" box="[422,606,295,321]" pageId="9" pageNumber="96">L.M. D’Albertis</collectorName>
—
<specimenCount id="9D26FD77FFFE6E12FDAA1122FD4EFEF1" box="[635,653,295,321]" count="1" pageId="9" pageNumber="96" type="male">♂</specimenCount>
<typeStatus id="549B885CFFFE6E12FD441122FD3CFEF1" box="[661,767,295,321]" pageId="9" pageNumber="96" type="lectotype">lectotype</typeStatus>
,
<materialsCitation id="3B483CA3FFFE6E12FCDB1122FB87FEF1" box="[778,1092,295,321]" collectionCode="MHNG" collectorName="L. M. D'Albertis" country="Papua New Guinea" location="Fly River" longitude="142.0" pageId="9" pageNumber="96" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="S" typeStatus="paralectotype">
<specimenCount id="9D26FD77FFFE6E12FCDB1122FCF7FEF1" box="[778,820,295,321]" count="1" pageId="9" pageNumber="96" type="female">1 ♀</specimenCount>
<typeStatus id="549B885CFFFE6E12FCEA1122FC11FEF1" box="[827,978,295,321]" pageId="9" pageNumber="96" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="ED31AE3BFFFE6E12FC311122FBFFFEF1" box="[992,1084,295,321]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="96" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFE6E12FF16114FFD31FC89" pageId="9" pageNumber="96" type="description">
<paragraph id="8B9F36FEFFFE6E12FF16114FFD31FC89" blockId="9.[151,1437,150,897]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF16114FFDF3FED5" bold="true" box="[199,560,330,357]" pageId="9" pageNumber="96">Redescription. Measurements.</emphasis>
Body length 6.0 mm, maximum body width
<quantity id="4CD89B1BFFFE6E12FBCD114EFBB7FED5" box="[1052,1140,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="96" unit="mm" value="2.9">2.9 mm</quantity>
.
<emphasis id="B954EAECFFFE6E12FBAC114EFB02FED5" bold="true" box="[1149,1217,331,357]" pageId="9" pageNumber="96">Body.</emphasis>
Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third.
<emphasis id="B954EAECFFFE6E12FC29116AFBFDFE39" bold="true" box="[1016,1086,367,393]" pageId="9" pageNumber="96">Head.</emphasis>
Clypeal edge lateral to clypeal teeth lacking emargination (
<figureCitation id="131B2A7BFFFE6E12FE1F1196FDE6FE1D" box="[462,549,403,429]" captionStart="FIGURE 6" captionStartId="8.[152,255,2017,2041]" captionTargetBox="[151,1436,1071,1991]" captionTargetId="figure-192@8.[151,1436,1071,1991]" captionTargetPageId="8" captionText="FIGURE 6. Lectotype of Rhytomus elongatus. A, dorsal habitus; B, dorsal head; C, label data." figureDoi="http://doi.org/10.5281/zenodo.14746454" httpUri="https://zenodo.org/record/14746454/files/figure.png" pageId="9" pageNumber="96">Fig. 6B</figureCitation>
), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.3 times eye width.
<emphasis id="B954EAECFFFE6E12FBBB11B2FB29FE61" bold="true" box="[1130,1258,439,465]" pageId="9" pageNumber="96">Pronotum.</emphasis>
Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence long and conspicuous, pronotal basal portion along midline lacking fine longitudinal depression.
<emphasis id="B954EAECFFFE6E12FBAA11FAFB0DFDA9" bold="true" box="[1147,1230,511,537]" pageId="9" pageNumber="96">Elytra.</emphasis>
Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (
<figureCitation id="131B2A7BFFFE6E12FAE61242FA4CFDD1" box="[1335,1423,583,609]" captionStart="FIGURE 6" captionStartId="8.[152,255,2017,2041]" captionTargetBox="[151,1436,1071,1991]" captionTargetId="figure-192@8.[151,1436,1071,1991]" captionTargetPageId="8" captionText="FIGURE 6. Lectotype of Rhytomus elongatus. A, dorsal habitus; B, dorsal head; C, label data." figureDoi="http://doi.org/10.5281/zenodo.14746454" httpUri="https://zenodo.org/record/14746454/files/figure.png" pageId="9" pageNumber="96">Fig. 6A</figureCitation>
), basal elytral pit wide and deep, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures.
<emphasis id="B954EAECFFFE6E12FB08128AFADBFD19" bold="true" box="[1241,1304,655,681]" pageId="9" pageNumber="96">Legs.</emphasis>
Profemoral anterior surface deeply furrowed, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (e.g.
<figureCitation id="131B2A7BFFFE6E12FAA112B6FF0FFD41" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="9" pageNumber="96">Fig. 12D</figureCitation>
).
<emphasis id="B954EAECFFFE6E12FF0E12D3FE9AFD40" bold="true" box="[223,345,726,752]" pageId="9" pageNumber="96">Pygidium.</emphasis>
Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex.
<emphasis id="B954EAECFFFE6E12FB9012FFFB78FCA4" bold="true" box="[1089,1211,762,788]" pageId="9" pageNumber="96">Aedeagus.</emphasis>
Left paramere apex produced into a rounded oblique projection (
<figureCitation id="131B2A7BFFFE6E12FD5A131AFD26FC89" box="[651,741,799,825]" captionStart="FIGURE 9" captionStartId="12.[152,255,1959,1983]" captionTargetBox="[155,1431,238,1929]" captionTargetId="figure-53@12.[151,1435,234,1933]" captionTargetPageId="12" captionText="FIGURE 9. Male aedeagus of Rhytomus species, left lateral, dorsal, right lateral.A, R. elongatus; B, R. fervidus; C, R. howdeni; D, R. larseni. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14746462" httpUri="https://zenodo.org/record/14746462/files/figure.png" pageId="9" pageNumber="96">Fig. 9A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33A6575FFFE6E12FF161346FDC2FC31" pageId="9" pageNumber="96" type="discussion">
<paragraph id="8B9F36FEFFFE6E12FF161346FDC2FC31" blockId="9.[151,1437,150,897]" pageId="9" pageNumber="96">
<emphasis id="B954EAECFFFE6E12FF161346FEF8FCED" bold="true" box="[199,315,835,861]" pageId="9" pageNumber="96">Remarks.</emphasis>
This species was described based on
<specimenCount id="9D26FD77FFFE6E12FD211346FC4AFCED" box="[752,905,835,861]" count="2" pageId="9" pageNumber="96" type="generic" typeStatus="syntype">two syntypes</specimenCount>
. In 2015, Génier and Larsen labeled
<specimenCount id="9D26FD77FFFE6E12FAE51346FA5FFCED" box="[1332,1436,835,861]" count="1" pageId="9" pageNumber="96" type="male">one male</specimenCount>
from this series as the
<typeStatus id="549B885CFFFE6E12FE431362FE3FFC31" box="[402,508,871,897]" pageId="9" pageNumber="96" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>