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<document id="FCF7AB59972DD701B50B37A37CD2758A" ID-CLB-Dataset="31408" ID-DOI="10.11646/zootaxa.4350.2.9" ID-GBIF-Dataset="5d01d0ba-9009-45b4-aef1-bea31993f0d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053222" ID-ZooBank="CB65B034-1CFD-475B-B4FE-1BCF7F0A388D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510901782709" checkinUser="plazi" docAuthor="Gordon, Dennis P., Voje, Kjetil L. &amp; Taylor, Paul D." docDate="2017" docId="027D87EBFFF5260AFCC1D56ED982FD5A" docLanguage="en" docName="zootaxa.4350.2.9.pdf" docOrigin="Zootaxa 4350 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Steginoporella modesta Gordon &amp; Voje &amp; Taylor 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="356" masterDocId="FE44FF93FFFF2601FC56D37ADC18FFFD" masterDocTitle="Living and fossil Steginoporellidae (Bryozoa: Cheilostomata) from New Zealand" masterLastPageNumber="362" masterPageNumber="345" pageNumber="355" updateTime="1698500382477" updateUser="ExternalLinkService">
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<mods:title id="1ABB8AD8ED9CF214C4CE5B5DED8A716D">Living and fossil Steginoporellidae (Bryozoa: Cheilostomata) from New Zealand</mods:title>
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<mods:namePart id="4E1E31C897BB0D3C2F445A9F09D2D295">Gordon, Dennis P.</mods:namePart>
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<mods:namePart id="5E9D375E1298175B9B39B4F8D6B9D093">Voje, Kjetil L.</mods:namePart>
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<mods:namePart id="A24401EF900C8B1579AB7ECBEA1C0A07">Taylor, Paul D.</mods:namePart>
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<mods:title id="400212D544C0CAC8FBE8CD5E69CBFBF7">Zootaxa</mods:title>
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<mods:date id="CC828B03DD6926A7913DC46885E3DEE8">2017</mods:date>
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<mods:number id="70D08155C6645B45FD59BDE3D35FF5A3">2017-11-17</mods:number>
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<mods:number id="97EFBE05673A3F879B19EF3261C36AD4">4350</mods:number>
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<mods:identifier id="4823670EC68071E37E6D404CED479BA5" type="DOI">10.11646/zootaxa.4350.2.9</mods:identifier>
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<treatment id="027D87EBFFF5260AFCC1D56ED982FD5A" ID-DOI="http://doi.org/10.5281/zenodo.6042322" ID-GBIF-Taxon="137309550" ID-Zenodo-Dep="6042322" LSID="urn:lsid:plazi:treatment:027D87EBFFF5260AFCC1D56ED982FD5A" httpUri="http://treatment.plazi.org/id/027D87EBFFF5260AFCC1D56ED982FD5A" lastPageId="11" lastPageNumber="356" pageId="10" pageNumber="355">
<subSubSection id="C2CE6576FFF5260BFCC1D56EDD4CF9B2" pageId="10" pageNumber="355" type="nomenclature">
<paragraph id="8A6B36FDFFF5260BFCC1D56EDDE1F9D3" blockId="10.[151,505,1556,1615]" box="[151,505,1556,1582]" pageId="10" pageNumber="355">
<heading id="D1238191FFF5260BFCC1D56EDDE1F9D3" bold="true" box="[151,505,1556,1582]" fontSize="11" level="1" pageId="10" pageNumber="355" reason="1">
<emphasis id="B8A0EAEFFFF5260BFCC1D56EDDE1F9D3" bold="true" box="[151,505,1556,1582]" pageId="10" pageNumber="355">
<taxonomicName id="4DD44D7EFFF5260BFCC1D56EDDA8F9D3" authority="Gordon &amp; Voje &amp; Taylor, 2017" authorityName="Gordon &amp; Voje &amp; Taylor" authorityYear="2017" box="[151,432,1556,1582]" class="Gymnolaemata" family="Steginoporellidae" genus="Steginoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="10" pageNumber="355" phylum="Bryozoa" rank="species" species="modesta" status="sp. nov.">
<emphasis id="B8A0EAEFFFF5260BFCC1D56EDDA8F9D3" bold="true" box="[151,432,1556,1582]" italics="true" pageId="10" pageNumber="355">Steginoporella modesta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3935794FFF5260BFDE0D56EDDE1F9D3" box="[438,505,1556,1582]" pageId="10" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8A6B36FDFFF5260BFCC1D54CDD4CF9B2" blockId="10.[151,505,1556,1615]" box="[151,340,1590,1615]" pageId="10" pageNumber="355">
(
<figureCitation id="12EF2A78FFF5260BFCC9D54CDCE0F9B2" box="[159,248,1590,1615]" captionStart="FIGURE 1" captionStartId="2.[151,250,963,985]" captionTargetBox="[253,1336,518,942]" captionTargetId="figure@2.[252,1336,517,942]" captionTargetPageId="2" captionText="FIGURE 1. Six species of Recent New Zealand Steginoporella (NIWA stations in parentheses). A, S. neozelanica (TAN 1105 / 27); B, S. marcusi (respectively from three localities, left to right, D 896, TAN 1105 / 42, TAN 1105 / 60); C, S. perplexa (small colony from Z 15617); D, S. discors n. sp. (TAN 0906); E, S. modesta n. sp. (TAN 0413 / 109); F, S. lineata n. sp. (TAN 0205 / 64). All to same scale (scalebar 25 mm)." httpUri="https://zenodo.org/record/1053224/files/figure.png" pageId="10" pageNumber="355">Figs 1E</figureCitation>
,
<figureCitation id="12EF2A78FFF5260BFD52D54DDD53F9B2" box="[260,331,1591,1615]" captionStart="FIGURES 20 25" captionStartId="9.[151,264,1182,1205]" captionTargetBox="[250,1338,194,1161]" captionTargetId="figure@9.[249,1338,193,1161]" captionTargetPageId="9" captionText="FIGURES 20 25. Zooids of New Zealand Steginoporella (NIWA stations in parentheses). 20, S. discors n. sp. (Z 9743); 21, 23, S. modesta n. sp., respectively showing autozooids and an ancestrula and daughter zooid (TAN 0413 / 109); 22, 24 25, S. lineata n. sp., showing autozooids and larger avicularian B-zooids (TAN 0205 / 64). Figs 20 22 at same scale (scalebar 1 mm); scalebars 0.5 mm in figs 23 25." httpUri="https://zenodo.org/record/1053234/files/figure.png" pageId="10" pageNumber="355">21, 23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2CE6576FFF5260BFCC1D504D890F96A" box="[151,1160,1662,1687]" pageId="10" pageNumber="355" type="etymology">
<paragraph id="8A6B36FDFFF5260BFCC1D504D890F96A" blockId="10.[151,1436,1662,2011]" box="[151,1160,1662,1687]" pageId="10" pageNumber="355">
<emphasis id="B8A0EAEFFFF5260BFCC1D504DD06F96A" bold="true" box="[151,286,1662,1687]" pageId="10" pageNumber="355">Etymology.</emphasis>
Latin
<emphasis id="B8A0EAEFFFF5260BFD30D504DDD6F96A" box="[358,462,1662,1687]" italics="true" pageId="10" pageNumber="355">modestus</emphasis>
, modest, unassuming, alluding to the small size of the colony.
</paragraph>
</subSubSection>
<subSubSection id="C2CE6576FFF5260BFC91D5D8D8D8F922" pageId="10" pageNumber="355" type="materials_examined">
<paragraph id="8A6B36FDFFF5260BFC91D5D8D8D8F922" blockId="10.[151,1436,1662,2011]" pageId="10" pageNumber="355">
<materialsCitation id="3ABC3CA0FFF5260BFC91D5D8D8A4F922" ID-GBIF-Occurrence="1701201433" collectingDate="2004-11-13" collectionCode="NIWA, TAN" latitude="-37.549" location="White Island" longLatPrecision="70" longitude="176.988" municipality="Material" pageId="10" pageNumber="355" specimenCode="NIWA23304, TAN0413, NIWA122576" specimenCount="144" stateProvince="Bay of Plenty" typeStatus="holotype">
<emphasis id="B8A0EAEFFFF5260BFC91D5D8DDACF946" bold="true" box="[199,436,1698,1723]" pageId="10" pageNumber="355">
<collectingMunicipality id="6A0FAC87FFF5260BFC91D5D8DD37F946" box="[199,303,1698,1723]" pageId="10" pageNumber="355">Material</collectingMunicipality>
examined.
</emphasis>
<emphasis id="B8A0EAEFFFF5260BFDE8D5D8DE36F946" box="[446,558,1698,1723]" italics="true" pageId="10" pageNumber="355">
<typeStatus id="556F885FFFF5260BFDE8D5D8DE30F946" box="[446,552,1698,1723]" pageId="10" pageNumber="355" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DA729E86FFF5260BFE6ED5D9DED3F946" box="[568,715,1699,1723]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="10" pageNumber="355">NIWA23304</specimenCode>
,
<specimenCode id="DA729E86FFF5260BFE8FD5D9DF54F946" box="[729,844,1699,1723]" collectionCode="TAN" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="10" pageNumber="355">TAN0413</specimenCode>
/109,
<geoCoordinate id="EFE0503AFFF5260BFFDDD5D8DFE7F946" box="[907,1023,1698,1723]" direction="south" orientation="latitude" pageId="10" pageNumber="355" precision="55" value="-37.549">37.549° S</geoCoordinate>
,
<geoCoordinate id="EFE0503AFFF5260BF859D5D8D88BF947" box="[1039,1171,1698,1723]" direction="east" orientation="longitude" pageId="10" pageNumber="355" precision="55" value="176.988">176.988° E</geoCoordinate>
, west of
<location id="8F0B6026FFF5260BF954D5D8D98FF946" LSID="urn:lsid:plazi:treatment:027D87EBFFF5260AFCC1D56ED982FD5A:8F0B6026FFF5260BF954D5D8D98FF946" box="[1282,1431,1698,1723]" latitude="-37.549" longLatPrecision="70" longitude="176.988" municipality="Material" name="White Island" pageId="10" pageNumber="355" stateProvince="Bay of Plenty">White Island</location>
,
<collectingRegion id="4810F81FFFF5260BFCC1D5BDDD2AF922" box="[151,306,1734,1759]" country="New Zealand" name="Bay of Plenty" pageId="10" pageNumber="355">Bay of Plenty</collectingRegion>
,
<collectingDate id="EE2EE9D5FFF5260BFD6BD5BDDE00F922" box="[317,536,1734,1759]" pageId="10" pageNumber="355" value="2004-11-13">
<date id="FE6A103DFFF5260BFD6BD5BDDE00F922" box="[317,536,1734,1759]" pageId="10" pageNumber="355" value="2004-11-13">
13 November
<quantity id="4D2C9B18FFF5260BFD8BD5BDDE00F922" box="[477,536,1735,1759]" metricMagnitude="3" metricUnit="m" metricValue="2.004142" pageId="10" pageNumber="355" unit="m" value="2004.142">2004</quantity>
</date>
</collectingDate>
,
<specimenCount id="9CD2FD74FFF5260BFE75D5BDDE69F923" box="[547,625,1735,1759]" pageId="10" pageNumber="355" type="generic">142 m.</specimenCount>
<emphasis id="B8A0EAEFFFF5260BFE2ED5BDDEFFF923" box="[632,743,1735,1758]" italics="true" pageId="10" pageNumber="355">
<typeStatus id="556F885FFFF5260BFE2ED5BDDEF9F923" box="[632,737,1735,1758]" pageId="10" pageNumber="355" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCode id="DA729E86FFF5260BFEB9D5BCDF97F922" box="[751,911,1734,1759]" collectionCode="NIWA" country="New Zealand" httpUri="http://grbio.org/cool/nyq3-3g6s" name="National Institute of Water and Atmospheric Research" pageId="10" pageNumber="355">
NIWA
<date id="FE6A103DFFF5260BFF6AD5BCDF97F922" box="[828,911,1734,1759]" pageId="10" pageNumber="355" value="76-12-25">122576</date>
</specimenCode>
, same data as for
<typeStatus id="556F885FFFF5260BF800D5BCD8A4F922" box="[1110,1212,1734,1759]" pageId="10" pageNumber="355" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2CE6576FFF5260AFC91D590DFBBFE51" lastPageId="11" lastPageNumber="356" pageId="10" pageNumber="355" type="description">
<paragraph id="8A6B36FDFFF5260BFC91D590D8D0F8DA" blockId="10.[151,1436,1662,2011]" pageId="10" pageNumber="355">
<emphasis id="B8A0EAEFFFF5260BFC91D590DD41F8FE" bold="true" box="[199,345,1770,1795]" pageId="10" pageNumber="355">Description.</emphasis>
Colony encrusting, comprising small lobate to sublinear pluriserial growths not more than six zooids across, up to
<quantity id="4D2C9B18FFF5260BFD2DD475DDD2F8DB" box="[379,458,1807,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="355" unit="mm" value="19.0">19 mm</quantity>
long and
<quantity id="4D2C9B18FFF5260BFE6ED474DE62F8DB" box="[568,634,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="355" unit="mm" value="5.0">5 mm</quantity>
wide. Colour of dried colonies pale brownish-beige.
</paragraph>
<paragraph id="8A6B36FDFFF5260BFC91D448D8ADF84B" blockId="10.[151,1436,1662,2011]" pageId="10" pageNumber="355">Zooids monomorphic, roundly hexagonal, with one of the pair of daughter zooids at the bifurcation of a zooid row more or less parallel-sided, with mostly arcuate distal and proximal margins (mean L/W = 1.45); widest mostly about mid-length. Cryptocystal area more than half zooid length, extensive shelf sunken, weakly concave, mostly granular-tubercular, with sloping pseudoporous semicircular area proximal to median process.</paragraph>
<paragraph id="8A6B36FDFFF5260AFC91D4B8DDACFF28" blockId="10.[151,1436,1662,2011]" lastBlockId="11.[151,1436,151,680]" lastPageId="11" lastPageNumber="356" pageId="10" pageNumber="355">Opercular area delimited proximally by stout triangular pivots; median process with distolateral wings that do not reach pivots or lateral margins; proximomedial foramen of small to moderate size. Inner face of operculum smooth, lacking sclerites.</paragraph>
<paragraph id="8A6B36FDFFF4260AFC91D3A5DF62FE98" blockId="11.[151,1436,151,680]" pageId="11" pageNumber="356">Polypide not seen. Ancestrula smaller than other zooids but resembling them, deep-bodied, the sides interiorwalled and granular; smooth thin margins of frontal cryptocyst on both sides converging without meeting proximally, descending to substratum, with narrow granular strip between. First daughter zooid budded laterally, followed near-simultaneously by distolateral pair from ancestrula.</paragraph>
<paragraph id="8A6B36FDFFF4260AFC91D215DFBBFE51" blockId="11.[151,1436,151,680]" pageId="11" pageNumber="356">
<emphasis id="B8A0EAEFFFF4260AFC91D215DD65FE75" bold="true" box="[199,381,367,392]" pageId="11" pageNumber="356">Measurements.</emphasis>
In micrometres: ZL 1092±173, 9321389 (4, 10); ZW 751±86, 643912 (4, 10); OpL 310±43, 256367 (1, 10);OpW 513±33, 456557 (1, 10). AnL 978, AnW 400.
</paragraph>
</subSubSection>
<subSubSection id="C2CE6576FFF4260AFC91D2CDDCDCFD78" pageId="11" pageNumber="356" type="discussion">
<paragraph id="8A6B36FDFFF4260AFC91D2CDDCDCFD78" blockId="11.[151,1436,151,680]" pageId="11" pageNumber="356">
<emphasis id="B8A0EAEFFFF4260AFC91D2CDDD23FE2D" bold="true" box="[199,315,439,464]" pageId="11" pageNumber="356">Remarks.</emphasis>
Colonies of
<taxonomicName id="4DD44D7EFFF4260AFD98D2C2DE50FE2D" authority="Gordon &amp; Voje &amp; Taylor, 2017" authorityName="Gordon &amp; Voje &amp; Taylor" authorityYear="2017" box="[462,584,439,464]" class="Gymnolaemata" family="Steginoporellidae" genus="Steginoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="11" pageNumber="356" phylum="Bryozoa" rank="species" species="modesta" status="sp. nov.">
<emphasis id="B8A0EAEFFFF4260AFD98D2C2DE50FE2D" box="[462,584,439,464]" italics="true" pageId="11" pageNumber="356">S. modesta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3935794FFF4260AFE06D2CDDE88FE2D" box="[592,656,439,464]" pageId="11" pageNumber="356" rank="species">
<emphasis id="B8A0EAEFFFF4260AFE06D2CDDE88FE2D" bold="true" box="[592,656,439,464]" pageId="11" pageNumber="356">n. sp.</emphasis>
</taxonomicNameLabel>
, like those of
<taxonomicName id="4DD44D7EFFF4260AFF62D2CDD836FE2D" authority="Gordon &amp; Voje &amp; Taylor, 2017" authorityName="Gordon &amp; Voje &amp; Taylor" authorityYear="2017" box="[820,1070,439,464]" class="Gymnolaemata" family="Steginoporellidae" genus="Steginoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="11" pageNumber="356" phylum="Bryozoa" rank="species" species="lineata" status="sp. nov.">
<emphasis id="B8A0EAEFFFF4260AFF62D2CDD836FE2D" box="[820,1070,439,464]" italics="true" pageId="11" pageNumber="356">Steginoporella lineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3935794FFF4260AF863D2CDD86DFE2D" box="[1077,1141,439,464]" pageId="11" pageNumber="356" rank="species">
<emphasis id="B8A0EAEFFFF4260AF863D2CDD86DFE2D" bold="true" box="[1077,1141,439,464]" pageId="11" pageNumber="356">n. sp.</emphasis>
</taxonomicNameLabel>
, another deep-sea species (see below), are very small and somewhat narrow. Unlike
<taxonomicName id="4DD44D7EFFF4260AFF04D2A7DFDAFE08" authority="Gordon &amp; Voje &amp; Taylor, 2017" authorityName="Gordon &amp; Voje &amp; Taylor" authorityYear="2017" box="[850,962,476,501]" class="Gymnolaemata" family="Steginoporellidae" genus="Steginoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="11" pageNumber="356" phylum="Bryozoa" rank="species" species="lineata" status="sp. nov.">
<emphasis id="B8A0EAEFFFF4260AFF04D2A7DFDAFE08" box="[850,962,476,501]" italics="true" pageId="11" pageNumber="356">S. lineata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3935794FFF4260AFF98D2A6D80AFE08" box="[974,1042,476,501]" pageId="11" pageNumber="356" rank="species">
<emphasis id="B8A0EAEFFFF4260AFF98D2A6D80AFE08" bold="true" box="[974,1042,476,501]" pageId="11" pageNumber="356">n. sp.</emphasis>
</taxonomicNameLabel>
, zooids of
<taxonomicName id="4DD44D7EFFF4260AF8C8D2A7D904FE08" authority="Gordon &amp; Voje &amp; Taylor, 2017" authorityName="Gordon &amp; Voje &amp; Taylor" authorityYear="2017" box="[1182,1308,476,501]" class="Gymnolaemata" family="Steginoporellidae" genus="Steginoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomata" pageId="11" pageNumber="356" phylum="Bryozoa" rank="species" species="modesta" status="sp. nov.">
<emphasis id="B8A0EAEFFFF4260AF8C8D2A7D904FE08" box="[1182,1308,476,501]" italics="true" pageId="11" pageNumber="356">S. modesta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3935794FFF4260AF97FD2A6D975FE08" box="[1321,1389,476,501]" pageId="11" pageNumber="356" rank="species">
<emphasis id="B8A0EAEFFFF4260AF97FD2A6D975FE08" bold="true" box="[1321,1389,476,501]" pageId="11" pageNumber="356">n. sp.</emphasis>
</taxonomicNameLabel>
are monomorphic. The species is evidently pigmented in life—while a specimen was being prepared for SEM the bleaching solution turned pale purple. All specimens, including a small ancestrulate colony, were loose in the sample but their undersides are carpeted by a thin layer of sponge spicules, indicating the apparent substratum in life.
</paragraph>
</subSubSection>
<subSubSection id="C2CE6576FFF4260AFC91D1F5D982FD5A" box="[199,1434,655,680]" pageId="11" pageNumber="356" type="distribution">
<paragraph id="8A6B36FDFFF4260AFC91D1F5D982FD5A" blockId="11.[151,1436,151,680]" box="[199,1434,655,680]" pageId="11" pageNumber="356">
<emphasis id="B8A0EAEFFFF4260AFC91D1F5DD78FD55" bold="true" box="[199,352,655,680]" pageId="11" pageNumber="356">Distribution.</emphasis>
Endemic to
<collectingCountry id="F2C3766DFFF4260AFDB8D1EADE9FFD55" box="[494,647,655,680]" name="New Zealand" pageId="11" pageNumber="356">New Zealand</collectingCountry>
, being known only from the
<typeStatus id="556F885FFFF4260AFF9CD1EADFE2FD55" box="[970,1018,656,680]" pageId="11" pageNumber="356">type</typeStatus>
locality in the
<collectingRegion id="4810F81FFFF4260AF8F5D1EAD958FD55" box="[1187,1344,655,680]" country="New Zealand" name="Bay of Plenty" pageId="11" pageNumber="356">Bay of Plenty</collectingRegion>
;
<specimenCount id="9CD2FD74FFF4260AF91AD1EAD982FD5A" box="[1356,1434,656,680]" pageId="11" pageNumber="356" type="generic">
<quantity id="4D2C9B18FFF4260AF91AD1EAD98EFD5A" box="[1356,1430,656,680]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="11" pageNumber="356" unit="m" value="142.0">142 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>