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<document id="EFA8F6D5BF9FFAE690FEA0B3A28FFC01" ID-CLB-Dataset="3916" ID-DOI="10.11646/zootaxa.3879.1.1" ID-GBIF-Dataset="4b976ba3-1a7b-4dd5-8afb-630b7cc8a695" ID-ISSN="1175-5326" ID-Zenodo-Dep="4949051" ID-ZooBank="6423894B-97D9-4286-ABB9-D4AF072B57FD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623702926238" checkinUser="felipe" docAuthor="Borkent, Art" docDate="2014" docId="027587C9BD753028FDA21CD24CAEE38C" docLanguage="en" docName="zt03879p327.pdf.imf" docOrigin="Zootaxa 3879 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anebomyia Borkent 2014, new genus" docType="treatment" docVersion="10" lastPageNumber="95" masterDocId="FE4CFFB1BD2E3076FFF81A154D70E204" masterDocTitle="The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera" masterLastPageNumber="327" masterPageNumber="1" pageNumber="92" updateTime="1699015668130" updateUser="ExternalLinkService">
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<mods:title id="8C701D8DF95ED530DCA054BCB5D51074">The Pupae of the Biting Midges of the World (Diptera: Ceratopogonidae), With a Generic Key and Analysis of the Phylogenetic Relationships Between Genera</mods:title>
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<mods:namePart id="0E8AB5201CACB8E30A61B71191AF02A0">Borkent, Art</mods:namePart>
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<mods:date id="1F1965DDC86FD8886C45E646098B9BEB">2014</mods:date>
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<mods:number id="EF4169140F110EC22722D1229FA0F172">2014-10-31</mods:number>
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<treatment id="027587C9BD753028FDA21CD24CAEE38C" ID-DOI="http://doi.org/10.5281/zenodo.5593049" ID-GBIF-Taxon="189161033" ID-Zenodo-Dep="5593049" LSID="urn:lsid:plazi:treatment:027587C9BD753028FDA21CD24CAEE38C" httpUri="http://treatment.plazi.org/id/027587C9BD753028FDA21CD24CAEE38C" lastPageId="94" lastPageNumber="95" pageId="91" pageNumber="92">
<subSubSection id="C2C66554BD75302DFDA21CD24EA9E4E6" box="[602,985,1735,1762]" pageId="91" pageNumber="92" type="nomenclature">
<paragraph id="8A6336DFBD75302DFDA21CD24EA9E4E6" blockId="91.[509,1078,1735,1797]" box="[602,985,1735,1762]" pageId="91" pageNumber="92">
<emphasis id="B8A8EACDBD75302DFDA21CD24EA9E4E6" bold="true" box="[602,985,1735,1762]" italics="true" pageId="91" pageNumber="92">
<taxonomicName id="4DDC4D5CBD75302DFDA21CD24E21E4E6" ID-CoL="89PMM" authority="Borkent" authorityName="Borkent" authorityYear="2014" box="[602,849,1735,1762]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus" status="gen. nov.">Anebomyia Borkent</taxonomicName>
<taxonomicNameLabel id="A39B57B6BD75302DFCA01CDD4EA9E4E6" box="[856,985,1736,1762]" pageId="91" pageNumber="92" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD75302DFE051CF94946E501" box="[509,1078,1772,1797]" pageId="91" pageNumber="92" type="description">
<paragraph id="8A6336DFBD75302DFE051CF94946E501" blockId="91.[509,1078,1735,1797]" box="[509,1078,1772,1797]" pageId="91" pageNumber="92">
(
<figureCitation id="12E72A5ABD75302DFDFD1CF94F04E501" box="[517,628,1772,1797]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="91" pageNumber="92">Figs. 21L</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFD781CF94FDFE501" box="[640,687,1772,1797]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="91" pageNumber="92">27G</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFD431CF94F98E501" box="[699,744,1772,1797]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="91" pageNumber="92">30T</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFD0C1CF94E6FE501" box="[756,799,1772,1797]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="91" pageNumber="92">39F</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFCD31CF94E29E501" box="[811,857,1772,1797]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="91" pageNumber="92">45T</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFC9C1CF94EE3E501" box="[868,915,1772,1797]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="91" pageNumber="92">52E</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFC671CF94EA2E501" box="[927,978,1772,1797]" captionStart="FIGURE 68" captionStartId="313.[151,250,1970,1992]" captionTargetBox="[196,1408,224,1947]" captionTargetId="figure-48@313.[187,1418,218,1957]" captionTargetPageId="313" captionText="FIGURE 68. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Anebomyia atripes. B. Homohelea delanoe. C. Leehelea hispida (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949292" httpUri="https://zenodo.org/record/4949292/files/figure.png" pageId="91" pageNumber="92">68A</figureCitation>
,
<figureCitation id="12E72A5ABD75302DFC251CF9495EE501" box="[989,1070,1772,1797]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="91" pageNumber="92">76K–L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD75302DFF6F1D214F5DE54E" box="[151,557,1844,1866]" pageId="91" pageNumber="92" type="reference_group">
<paragraph id="8A6336DFBD75302DFF6F1D214F5DE54E" blockId="91.[151,1361,1844,1938]" box="[151,557,1844,1866]" pageId="91" pageNumber="92">
<taxonomicName id="4DDC4D5CBD75302DFF6F1D214CD4E54E" ID-CoL="89PMM" authority="Borkent, 2014" authorityName="Borkent" authorityYear="2014" box="[151,420,1844,1866]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD75302DFF6F1D214C79E54E" box="[151,265,1844,1866]" italics="true" pageId="91" pageNumber="92">Anebomyia</emphasis>
<bibRefCitation id="EE4D4B2EBD75302DFEF71D214CD4E54E" author="Borkent" box="[271,420,1844,1866]" firstAuthor="Borkent" pageId="91" pageNumber="92" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent, 2014</bibRefCitation>
</taxonomicName>
: in this work.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD75302DFDCB1D214FAAE56A" pageId="91" pageNumber="92" type="type_taxon">
<paragraph id="8A6336DFBD75302DFDCB1D214FAAE56A" blockId="91.[151,1361,1844,1938]" pageId="91" pageNumber="92">
<typeStatus id="5567887DBD75302DFDCB1D214F15E54D" box="[563,613,1844,1865]" pageId="91" pageNumber="92">Type</typeStatus>
species:
<taxonomicName id="4DDC4D5CBD75302DFD3A1D21492BE54E" ID-CoL="72T6X" authority="Wirth, 1962: 281" authorityName="Wirth" authorityPageNumber="281" authorityYear="1962" box="[706,1115,1844,1866]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD75302DFD3A1D214ED6E54D" box="[706,934,1844,1866]" italics="true" pageId="91" pageNumber="92">Mallochohelea atripes</emphasis>
<bibRefCitation id="EE4D4B2EBD75302DFC541D21492BE54E" author="Wirth" box="[940,1115,1844,1866]" firstAuthor="Wirth" page="281" pageId="91" pageNumber="92" pagination="272 - 287" refId="ref198995" refString="Wirth, W. W. (1962) A reclassification of the Palpomyia - Bezzia-Macropeza groups, and a revision of the North American Sphaeromiini (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 55, 272 - 287." type="journal article" year="1962">Wirth, 1962: 281</bibRefCitation>
</taxonomicName>
, by present designation. Included species and taxonomic discussion: given below.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD75302DFF6F1D69484FE596" box="[151,1343,1916,1938]" pageId="91" pageNumber="92" type="etymology">
<paragraph id="8A6336DFBD75302DFF6F1D69484FE596" blockId="91.[151,1361,1844,1938]" box="[151,1343,1916,1938]" pageId="91" pageNumber="92">Etymology: Anebos-myia (beardless-fly) refers to the lack of sternite 8 tufts on female adults of species in this genus.</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD75302DFF6F1DD04815E5DA" box="[151,1381,1989,2014]" pageId="91" pageNumber="92" type="diagnosis">
<paragraph id="8A6336DFBD75302DFF6F1DD04815E5DA" blockId="91.[151,1381,1989,2014]" box="[151,1381,1989,2014]" pageId="91" pageNumber="92">
<emphasis id="B8A8EACDBD75302DFF6F1DD04C44E5DA" bold="true" box="[151,308,1989,2014]" pageId="91" pageNumber="92">DIAGNOSIS</emphasis>
: Only pupa of
<taxonomicName id="4DDC4D5CBD75302DFE251DD04FECE5DA" authorityName="Borkent" authorityYear="2014" box="[477,668,1989,2014]" class="Insecta" family="Ceratopogonidae" kingdom="Animalia" order="Diptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="family">Ceratopogonidae</taxonomicName>
with the dorsal apotome with two pairs of tubercles (
<figureCitation id="12E72A5ABD75302DFB0B1DD04828E5DA" box="[1267,1368,1989,2014]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="91" pageNumber="92">Fig. 21L</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD72302AFF3F1A824EC2E7C4" pageId="92" pageNumber="93" type="description">
<paragraph id="8A6336DFBD72302AFF3F1A824EC2E7C4" blockId="92.[151,1437,151,1797]" pageId="92" pageNumber="93">
<emphasis id="B8A8EACDBD72302AFF3F1A824CFAE2B4" bold="true" box="[199,394,151,176]" pageId="92" pageNumber="93">DESCRIPTION</emphasis>
: Total length =
<quantity id="4D249B3ABD72302AFDBD1A824F9DE2AB" box="[581,749,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.16" metricValueMax="4.91" metricValueMin="3.41" pageId="92" pageNumber="93" unit="mm" value="4.16" valueMax="4.91" valueMin="3.41">3.41–4.91 mm</quantity>
. Without larval exuviae retained on abdomen. Exuviae with flagellum appressed against lateral margin of midleg, wing (as in
<figureCitation id="12E72A5ABD72302AFC751AA94E8FE2D1" box="[909,1023,188,213]" captionStart="FIGURE 16" captionStartId="261.[151,250,1897,1919]" captionTargetBox="[172,1416,199,1871]" captionTargetId="figure-85@261.[172,1416,193,1876]" captionTargetPageId="261" captionText="FIGURE 16. Anterior portion of heads. A. Parabezzia huberti, in ventral view (distorted slide specimen). B. Parabezzia sp. (5 km. E. Danby, Vermont, USA), ventrolateral view. C. Clinohelea curriei, in ventral view. D. Heteromyia clavata, in ventral view, dorsal apotome not present. E. Heteromyia wokei, in ventral, slightly lateral view. F. Pellucidomyia leei, in ventral view, dorsal apotome not present." figureDoi="http://doi.org/10.5281/zenodo.4949118" httpUri="https://zenodo.org/record/4949118/files/figure.png" pageId="92" pageNumber="93">Figs. 16B</figureCitation>
,
<figureCitation id="12E72A5ABD72302AFBF51AA94930E2D1" box="[1037,1088,188,213]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="92" pageNumber="93">33B</figureCitation>
). Ecdysial tear around base of antenna, along lateral margin of face to palpus (as in
<figureCitation id="12E72A5ABD72302AFCFF1ACA4E09E2FC" box="[775,889,223,248]" captionStart="FIGURE 17" captionStartId="262.[151,250,1910,1932]" captionTargetBox="[158,1430,178,1891]" captionTargetId="figure-91@262.[158,1431,175,1896]" captionTargetPageId="262" captionText="FIGURE 17. Anterior portion of heads. A. Jenkinshelea sp., in ventral view (from Ottawa, ON, Canada). B. Mallochohelea caudellii., in ventral view. C. Sphaeromias longipennis, in ventral view. D. Bezzia sp., in ventral view (6 km E Salmon Arm, BC, Canada). E. Palpomyia sp. (Black Lake, Quebec, Canada), in ventral view. F. Phaenobezzia sp., in ventral view (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949120" httpUri="https://zenodo.org/record/4949120/files/figure.png" pageId="92" pageNumber="93">Figs. 17C</figureCitation>
,
<figureCitation id="12E72A5ABD72302AFC701ACA4ECCE2FC" box="[904,956,223,248]" captionStart="FIGURE 79" captionStartId="324.[151,250,1861,1883]" captionTargetBox="[192,1396,201,1846]" captionTargetId="figure-148@324.[184,1406,195,1847]" captionTargetPageId="324" captionText="FIGURE 79. Evolution of ecdysial sutures and splits in pupal exuviae of Ceratopogonidae, head and anterior portion of thorax in ventral view. Numbers refer to character states of characters 2 and 8 in text. Red indicates ecdysial splits. A. Leptoconopinae. B. Forcipomyiinae. C. Dasyheleinae. D. Most Ceratopogonini. E. Most Ceratopogonini, Clinohelea. F. Allohelea, Atyphohelea, Parabezzia. G. Heteromyia, Pellucidomyia, Hebetula. H. most Sphaeromiini s. lat., most Palpomyiini, Paryphoconus; some Bezzia are nearly completed fused (as in Fig. 79I), with only the dorsal apotome separate, along with an ecdysial split along the base of the antenna and the dorsal thoracic split. I. Jenkinshelea, Macropeza, Mallochohelea, Probezzia, Neobezzia." figureDoi="http://doi.org/10.5281/zenodo.4949336" httpUri="https://zenodo.org/record/4949336/files/figure.png" pageId="92" pageNumber="93">79H</figureCitation>
).
<emphasis id="B8A8EACDBD72302AFC2A1ACA4961E2FC" bold="true" box="[978,1041,223,248]" pageId="92" pageNumber="93">Head</emphasis>
: Dorsal apotome (
<figureCitation id="12E72A5ABD72302AFB131ACA4823E2FC" box="[1259,1363,223,248]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="92" pageNumber="93">Fig. 21L</figureCitation>
), with ventral line of weakness, with additional dorsal pair dorsomedial tubercles, without central dome; dorsolateral cephalic sclerite (as in
<figureCitation id="12E72A5ABD72302AFE621B324C8FE344" box="[410,511,295,320]" captionStart="FIGURE 13" captionStartId="258.[151,250,1897,1919]" captionTargetBox="[172,1416,193,1876]" captionTargetId="figure-99@258.[172,1416,193,1876]" captionTargetPageId="258" captionText="FIGURE 13. Anterior portion of heads and thoraces, in anterior or anterodorsal view. A. Leptoconops kerteszi, ecdysial sutures of head enhanced to show position. B. Culicoides denticulatus. C. Ceratopogon nr. abstrusus. D. Brachypogon sp. (from 6 km E. Falkland, BC, Canada). E. Alluaudomyia bella, dorsal apotome absent. F. Clinohelea curriei. G. Pellucidomyia leei. H. Mallochohelea caudellii. I. Bezzia sp. (from Bolean Lake, BC, Canada)." figureDoi="http://doi.org/10.5281/zenodo.4949104" httpUri="https://zenodo.org/record/4949104/files/figure.png" pageId="92" pageNumber="93">Fig. 13H</figureCitation>
) fused to scutum, each side separated medially by dorsal apotome in whole pupa; mouthparts (
<figureCitation id="12E72A5ABD72302AFEDE1B594CFDE361" box="[294,397,332,357]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="92" pageNumber="93">Fig. 27G</figureCitation>
) with mandible well-developed, lacinia absent; palpus extending posterior to posterolateral margin of labium; labium separated medially by labrum, hypopharynx; apex of antenna (
<figureCitation id="12E72A5ABD72302AFB831B7A49ADE38C" box="[1147,1245,367,392]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="92" pageNumber="93">Fig. 39F</figureCitation>
) barely posterior to posterior extent of midlength portion of midleg (portion lateral to mesosternum), narrowed posteriorly; sensilla: dorsal apotomals (
<figureCitation id="12E72A5ABD72302AFE9E1BA24CBBE3D4" box="[358,459,439,464]" captionStart="FIGURE 21" captionStartId="266.[151,250,1837,1859]" captionTargetBox="[182,1405,192,1792]" captionTargetId="figure-122@266.[160,1419,186,1818]" captionTargetPageId="266" captionText="FIGURE 21. Dorsal apotomes, anterior view. A. Dibezzia prominens (from Mayer 1934c). B. Hebetula tonnoiri (from Elson- Harris 1987). C. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). D. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). E. Mallochohelea termophila (from Spinelli 1984, modified). F. Probezzia pallida (from Wirth &amp; Grogan 1979). G. Neobezzia fittkaui (from Ronderos et al. 2011, modified). H. Nilobezzia basispinigera (from Elson-Harris 1987). I. Nilobezzia virago (from Elson-Harris 1987). J. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). K. Lanatomyia electra (from Elson-Harris 1987). L. Anebomyia atripes (from Wirth &amp; Grogan 1979, modified)." figureDoi="http://doi.org/10.5281/zenodo.4949136" httpUri="https://zenodo.org/record/4949136/files/figure.png" pageId="92" pageNumber="93">Fig. 21L</figureCitation>
)—1 elongate seta, 1 campaniform sensillum; dorsolateral cephalic sclerite sensilla—1 seta, 1 campaniform sensillum; clypeal-labrals (
<figureCitation id="12E72A5ABD72302AFD2B1BC94E4EE3F1" box="[723,830,476,501]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="92" pageNumber="93">Fig. 27G</figureCitation>
)—2 slender setae; oculars (
<figureCitation id="12E72A5ABD72302AFB6C1BC9498FE3F1" box="[1172,1279,476,501]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="92" pageNumber="93">Fig. 27G</figureCitation>
)—2 setae, 1 campaniform sensillum.
<emphasis id="B8A8EACDBD72302AFE341BEA4F55E01C" bold="true" box="[460,549,511,536]" pageId="92" pageNumber="93">Thorax</emphasis>
: Prothoracic extension (
<figureCitation id="12E72A5ABD72302AFC9E1BEA4EACE01C" box="[870,988,511,536]" captionStart="FIGURE 27" captionStartId="272.[151,250,1950,1972]" captionTargetBox="[198,1418,210,1940]" captionTargetId="figure-69@272.[190,1421,205,1944]" captionTargetPageId="272" captionText="FIGURE 27. Posterior portion of heads, in ventral view. A. Mallochohelea caudellii. B. Probezzia sabroskyi. C. Neobezzia fittkaui. D. Nilobezzia curticornis. E. Johannsenomyia argentata. F. Lanatomyia electra. G. Anebomyia atripes. H. Homohelea delanoe." figureDoi="http://doi.org/10.5281/zenodo.4949164" httpUri="https://zenodo.org/record/4949164/files/figure.png" pageId="92" pageNumber="93">Fig. 27G</figureCitation>
) wide, well-developed but narrow dorsolaterally, not extending to antenna; mesonotum without tubercles, not extending posteromedially, not dividing metathorax medially (
<figureCitation id="12E72A5ABD72302AFE6018524F73E064" box="[408,515,583,608]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="92" pageNumber="93">Fig. 52E</figureCitation>
); respiratory organ (
<figureCitation id="12E72A5ABD72302AFD0E18524E2EE064" box="[758,862,583,608]" captionStart="FIGURE 45" captionStartId="290.[151,250,1738,1760]" captionTargetBox="[222,1373,186,1678]" captionTargetId="figure-193@290.[202,1384,184,1720]" captionTargetPageId="290" captionText="FIGURE 45. Respiratory organs, in dorsal view, pattern on tracheal tubes shown only in some. A. Clinohelea tasmaniensis (from Elson-Harris &amp; Kettle 1986). B. Clinohelea bimaculata (from Wirth &amp; Grogan 1979, modified). C. Heteromyia wokei (from Wirth &amp; Grogan 1977). D. Heteromyia clavata (from Wirth &amp; Grogan 1977). E. Pellucidomyia leei (from Elson-Harris &amp; Kettle 1986). F. Pellucidomyia geari (from de Meillon &amp; Wirth 1981). G. Dibezzia prominens (from Mayer 1934c). H. Hebetula tonnoiri (from Debenham 1974). I. Jenkinshelea albaria (from Wirth &amp; Grogan 1979). J. Macropeza albitarsis (from Szadziewski &amp; Dominiak, 2007). K. Mallochohelea albihalter (from Wirth &amp; Grogan 1979). L. Probezzia pallida (from Wirth &amp; Grogan 1979). M. Probezzia xanthogaster (from Wirth &amp; Grogan 1979). N. Neobezzia fittkaui (from Ronderos et al. 2011). O. Nilobezzia aranea (from Elson-Harris 1987). P. Nilobezzia virago (from Elson-Harris 1987). Q. Johannsenomyia argentata (from Wirth &amp; Grogan 1979). R. Lanatomyia electra (from Debenham 1974). S. Lanatomyia miles (from Debenham 1974). T. Anebomyia atripes (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949212" httpUri="https://zenodo.org/record/4949212/files/figure.png" pageId="92" pageNumber="93">Fig. 45T</figureCitation>
) length/width = 2.91–5.40, moderately elongate, somewhat flattened apically, with pores closely abutting at apex of respiratory organ, arranged in single row, outer surface with some wrinkles, with short, wide pedicel, base with elongate posteromedial apodeme, membranous base of respiratory organ short, tracheal tube straight to slightly curved along length, with spirals restricted to base, wrinkles to half length; wing (
<figureCitation id="12E72A5ABD72302AFE0118C24F2DE0F4" box="[505,605,727,752]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="92" pageNumber="93">Fig. 39F</figureCitation>
) without apical tubercle or angle, separated medially by fore-, midlegs; halter apex and hind leg (as in
<figureCitation id="12E72A5ABD72302AFE1618E94F23E111" box="[494,595,764,789]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="92" pageNumber="93">Fig. 33A</figureCitation>
) broadly abutting; halter apex abutting anterolateral knob-like extension of tergite 2; legs (
<figureCitation id="12E72A5ABD72302AFEBB190A4CD6E13C" box="[323,422,799,824]" captionStart="FIGURE 39" captionStartId="284.[151,250,1973,1995]" captionTargetBox="[180,1379,215,1945]" captionTargetId="figure-65@284.[170,1401,210,1949]" captionTargetPageId="284" captionText="FIGURE 39. Legs, wings and apices of the antennae, in ventral view. A. Probezzia sabroskyi. B. Neobezzia fittkaui. C. Nilobezzia sp. (from Australia). D. Johannsenomyia argentata. E. Lanatomyia electra. F. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949198" httpUri="https://zenodo.org/record/4949198/files/figure.png" pageId="92" pageNumber="93">Fig. 39F</figureCitation>
) with lateral margin of foreleg near midlength of wing evenly curved; hind leg visible at lateral margin of wing (as in
<figureCitation id="12E72A5ABD72302AFE2219514F45E159" box="[474,565,836,861]" captionStart="FIGURE 33" captionStartId="278.[151,250,1883,1905]" captionTargetBox="[176,1420,177,1860]" captionTargetId="figure-110@278.[176,1420,177,1860]" captionTargetPageId="278" captionText="FIGURE 33. Cephalothoraces and abdominal segments 1–3, in lateral view. A. Parabezzia sp. (5 km. E. Danby, Vermont, USA). B. Clinohelea curriei. C. Pellucidomyia leei. D. Hebetula tonnoiri. E. Jenkinshelea sp. (from Ottawa, ON, Canada). F. Mallochohelea caudellii. G. Probezzia seminigra. H. Nilobezzia sp. (from Kruger NP, South Africa). I. Sphaeromias longipennis. J. Bezzia sp. (6 km E. Salmon Arm, BC, Canada) K. Palpomyia sp. (Black Lake, Quebec, Canada). L. Phaenobezzia sp. (from Skulsuza, South Africa)." figureDoi="http://doi.org/10.5281/zenodo.4949184" httpUri="https://zenodo.org/record/4949184/files/figure.png" pageId="92" pageNumber="93">Fig. 33I</figureCitation>
); with apex of foreleg moderately anterior to apex of midleg; apex of hind leg abutting apex of midleg laterally; sensilla: anteromedials—2 elongate setae (as in
<figureCitation id="12E72A5ABD72302AFBC8197249B7E184" box="[1072,1223,871,896]" captionStart="FIGURE 31" captionStartId="276.[151,250,1885,1907]" captionTargetBox="[177,1424,178,1864]" captionTargetId="figure-106@276.[177,1424,178,1864]" captionTargetPageId="276" captionText="FIGURE 31. Dorsal setae from left side of thoraces; anteromedial setae. A. Homohelea delanoe. B. Leehelea hispida. C. Sphaeromias longipennis. D. Xenohelea galatea. E. Bezzia nobilis. F. Clastrieromyia dycei. G. Pachyhelea pachymera. H. Palpomyia jonesi. I. Phaenobezzia mashonensis. J. Paryphoconus oliveirai. K. Echinohelea lanei, partial lateral view (distorted slide specimen). L. Palpomyia (JAD685), dorsal view. M. Palpomyia (JAD685), dorsolateral view. N. Probezzia seminigra, dorsal view of left side." figureDoi="http://doi.org/10.5281/zenodo.4949178" httpUri="https://zenodo.org/record/4949178/files/figure.png" pageId="92" pageNumber="93">Figs. 31L–M</figureCitation>
); anterolaterals—1 moderately long seta; dorsal setae (
<figureCitation id="12E72A5ABD72302AFDDF19994FFDE1A1" box="[551,653,908,933]" captionStart="FIGURE 30" captionStartId="275.[151,250,1839,1861]" captionTargetBox="[161,1425,193,1797]" captionTargetId="figure-119@275.[161,1426,193,1813]" captionTargetPageId="275" captionText="FIGURE 30. Dorsal setae from left side of thoraces. A. Serromyia sp. B. Allohelea japonica. C. Monohelea obscura. D. Atyphohelea macroneura. E. Parabezzia huberti. F. Echinohelea lanei. G. Clinohelea curriei. H. Heteromyia clavata. I. Pellucidomyia leei. J. Dibezzia debenhamae (dirty specimen, D-5-T perhaps broken setal socket). K. Hebetula tonnoiri. L. Jenkinshelea magnipennis. M. Macropeza natalensis. N. Mallochohelea caudellii. O. Probezzia sabroskyi. P. Neobezzia fittkaui (from Ronderos et al. 2011, modified). Q. Nilobezzia curticornis. R. Johannsenomyia argentata. S. Lanatomyia miles. T. Anebomyia atripes." figureDoi="http://doi.org/10.5281/zenodo.4949174" httpUri="https://zenodo.org/record/4949174/files/figure.png" pageId="92" pageNumber="93">Fig. 30T</figureCitation>
)—D-1-T, D-2-T, D-4-T, D-5-T setae, D-3-T campaniform sensillum, D-3-T anterolateral to D-4-T; supraalar 2—campaniform sensillum; metathoracics (
<figureCitation id="12E72A5ABD72302AFB9A19BA49BBE1CC" box="[1122,1227,943,968]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="92" pageNumber="93">Fig. 52E</figureCitation>
)—1 campaniform sensillum; M-3-T distant from margin of metathorax (at least 1/3 length of metathorax).
<emphasis id="B8A8EACDBD72302AFB7C19C14988E1E9" bold="true" box="[1156,1272,980,1005]" pageId="92" pageNumber="93">Abdomen</emphasis>
: pigmentation light to dark brown, with tergite 1 with 3 medial spots, tergites 2–7 with medial area with stripe, 2 anterolateral spots, sternites 3–7 with pale to well defined medial stripe, anterolateral spot, segment 2 as wide or slightly wider than segment 3, segments with undivided, thin to thick setae, with rounded to pointed, short tubercles, tergites or sternites entire, each without membranous disc; segment 9 (
<figureCitation id="12E72A5ABD72302AFCC81E714EB0E679" box="[816,960,1124,1149]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="92" pageNumber="93">Figs. 76K–L</figureCitation>
) not strongly modified, terminal processes closely approximated basally, each projecting posterodorsolaterally, tapering to pointed apex; sensilla: tergite 1 (
<figureCitation id="12E72A5ABD72302AFF671EB94C72E6C1" box="[159,258,1196,1221]" captionStart="FIGURE 52" captionStartId="297.[151,250,1942,1964]" captionTargetBox="[190,1402,202,1928]" captionTargetId="figure-58@297.[181,1412,193,1932]" captionTargetPageId="297" captionText="FIGURE 52. Metathoraces and tergites 1, in dorsal view. A. Neobezzia fittkaui. B. Nilobezzia curticornis. C. Johannsenomyia argentata. D. Lanatomyia miles. E. Anebomyia atripes. F. Homohelea delanoe (reconstructed from distorted and dirty specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949234" httpUri="https://zenodo.org/record/4949234/files/figure.png" pageId="92" pageNumber="93">Fig. 52E</figureCitation>
) with 8 setae, 2 campaniform sensilla, including 3 lateral sensilla, D-2-I, D-3-I closely approximated, D- 7-I situated anteriorly near D-3-I; segment 4 (
<figureCitation id="12E72A5ABD72302AFD4F1EDA4E50E6EC" box="[695,800,1231,1256]" captionStart="FIGURE 68" captionStartId="313.[151,250,1970,1992]" captionTargetBox="[196,1408,224,1947]" captionTargetId="figure-48@313.[187,1418,218,1957]" captionTargetPageId="313" captionText="FIGURE 68. Abdominal segments four, in dorsal and ventral view (shagreen not shown). A. Anebomyia atripes. B. Homohelea delanoe. C. Leehelea hispida (reconstructed from distorted specimen)." figureDoi="http://doi.org/10.5281/zenodo.4949292" httpUri="https://zenodo.org/record/4949292/files/figure.png" pageId="92" pageNumber="93">Fig. 68A</figureCitation>
)—D-2-IV, D-3-IV moderately elongate setae on short tubercles; D-5-IV, D-8-IV, D-9-IV moderately elongate setae; D-5-IV on single tubercle, D-8-IV, D-9-IV on separate but closely approximated tubercles, posterior dorsal sensilla in transverse row, arranged medially to laterally: D-5-IV, D-4-IV, D-8-IV, D-9-IV; D-7-IV near D-3-IV; L-1-IV short seta on pointed tubercle, near L-3-IV; L-2-IV, L-3-IV, L-4-IV moderately elongate setae on pointed tubercles, V-5-IV, V-6-IV, V-7-IV moderately elongate setae on short tubercles, all closely approximated; segment 8 without D-3-VIII, without L-1-VIII; segment 9 (
<figureCitation id="12E72A5ABD72302AFEE01FB24C04E7C4" box="[280,372,1447,1472]" captionStart="FIGURE 76" captionStartId="321.[151,250,1765,1787]" captionTargetBox="[225,1365,181,1745]" captionTargetId="figure-185@321.[215,1375,177,1746]" captionTargetPageId="321" captionText="FIGURE 76. Abdominal segments 9. A. Macropeza albitarsis, male, in ventral view (modified from Szadziewski &amp; Dominiak, 2007). B. Macropeza albitarsis, female, in ventral view (from Szadziewski &amp; Dominiak, 2007). C. Mallochohelea termophila, male, in ventral view (from Spinelli 1984). D. Mallochohelea termophila, female, in dorsal view (from Spinelli 1984). E. Probezzia pallida, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). F. Neobezzia fittkaui, male, in dorsal view (from Ronderos, et al. 2011). G. Nilobezzia virago, female, in dorsal view (from Elson-Harris 1987). H. Johannsenomyia argentata, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). I. Johannsenomyia argentata, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979). J. Lanatomyia miles, female, in dorsal view (from Elson- Harris 1987). K. Anebomyia atripes, male, in dorsal and ventral view (from Wirth &amp; Grogan 1979). L. Anebomyia atripes, female, in dorsal and ventral view (from Wirth &amp; Grogan 1979)." figureDoi="http://doi.org/10.5281/zenodo.4949324" httpUri="https://zenodo.org/record/4949324/files/figure.png" pageId="92" pageNumber="93">Figs. 76</figureCitation>
K-L)—with D-5-IX, D-6-IX campaniform sensilla.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD72302AFF3F1FD94C84E471" pageId="92" pageNumber="93" type="distribution">
<paragraph id="8A6336DFBD72302AFF3F1FD94C84E471" blockId="92.[151,1437,151,1797]" pageId="92" pageNumber="93">
<emphasis id="B8A8EACDBD72302AFF3F1FD94F13E7E1" bold="true" box="[199,611,1484,1509]" pageId="92" pageNumber="93">DISTRIBUTION AND HABITAT</emphasis>
: The genus
<taxonomicName id="4DDC4D5CBD72302AFD001FD94E06E7E1" authorityName="Borkent" authorityYear="2014" box="[760,886,1484,1509]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFD001FD94E06E7E1" box="[760,886,1484,1509]" italics="true" pageId="92" pageNumber="93">Anebomyia</emphasis>
</taxonomicName>
is known from five species from the Nearctic, Afrotropical and Oriental Regions (all previously in
<taxonomicName id="4DDC4D5CBD72302AFD0C1FFA4EECE40C" box="[756,924,1519,1544]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFD0C1FFA4EECE40C" box="[756,924,1519,1544]" italics="true" pageId="92" pageNumber="93">Mallochohelea</emphasis>
</taxonomicName>
;
<bibRefCitation id="EE4D4B2EBD72302AFC561FFA493EE40C" author="Borkent" box="[942,1102,1519,1544]" firstAuthor="Borkent" pageId="92" pageNumber="93" refId="ref177229" refString="Borkent, A. (2014) World Species of Biting Midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. illinois. edu / research / FLYTREE / Borkent. html (accessed 20 May 2014)" type="url" year="2014">Borkent 2014</bibRefCitation>
). Immatures are known from stream and river margins, lakes and ponds.
<bibRefCitation id="EE4D4B2EBD72302AFD811C014E03E429" author="Knausenberger" box="[633,883,1556,1581]" firstAuthor="Knausenberger" pageId="92" pageNumber="93" refId="ref188128" refString="Knausenberger, W. I. (1987) Contributions to the autecology and ecosystematics of immature Ceratopogonidae (Diptera), with emphasis on the tribes Heteromyiini and Sphaeromiini in the Middle Atlantic United States. Ph. D. Dissertation, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, xxix + 729 pp." type="book" year="1987">Knausenberger (1987)</bibRefCitation>
discusses the various habitats in which
<taxonomicName id="4DDC4D5CBD72302AFAD71C0148ECE428" authorityName="Wirth" authorityYear="1962" box="[1327,1436,1556,1580]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD72302AFAD71C0148ECE428" box="[1327,1436,1556,1580]" italics="true" pageId="92" pageNumber="93">A. atripes</emphasis>
</taxonomicName>
was collected in the eastern
<collectingCountry id="F2CB764FBD72302AFE241C224F67E454" box="[476,535,1591,1616]" name="United States of America" pageId="92" pageNumber="93">USA</collectingCountry>
. At least some pupae were on emergent vegetation (although they do not have abdominal membranous discs).
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD72302BFF3F1C6A4C27E45A" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="discussion">
<paragraph id="8A6336DFBD72302AFF3F1C6A49CDE501" blockId="92.[151,1437,151,1797]" pageId="92" pageNumber="93">
<emphasis id="B8A8EACDBD72302AFF3F1C6A4F5CE49C" bold="true" box="[199,556,1663,1688]" pageId="92" pageNumber="93">TAXONOMIC DISCUSSION</emphasis>
: Pupae are known from two species (
<tableCitation id="C75E0364BD72302AFC2F1C6A4927E49C" box="[983,1111,1663,1688]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="174.[147,245,486,512]" captionStartId-1="242.[151,239,152,175]" captionTargetBox-0="[150,1771,751,1679]" captionTargetBox-1="[159,1428,341,1978]" captionTargetPageId-0="174" captionTargetPageId-1="242" captionText-0="TABLE 2. References to species of Ceratopogonidae described as eggs, larvae and/or pupae. Species studied as pupae in the current study have their name highlighted in bold font. Stages studied are referred to as 'E' for egg, 'L' for larva and 'P' for pupa. Taxa are arrangement phyletically by subfamily and tribe (some of which are paraphyletic) and within tribes genera are listed alphabetically. Subfamilies are abbreviated as follows: Lept.— Leptoconopinae, Forc.—Forcipomyiinae, Dasy.—Dasyheleinae, Cera.—Ceratopogoninae. References highlighted in bold font are those with SEM photomicrographs. Theses are cited only when their results have not been published. Duplicated figures or repeated descriptions by subsequent authors are not listed here. Fossil taxa are identified with a cross (†). The species name in bold indicates only that named material of the pupa was studied, not that the cited author's specimens were reexamined (although in many instances they were). The larva of Jenkinshelea albaria was tentatively associated by Wirth &amp; Grogan 1979 (not reared); see discussion by Knausenberger, 1987:248." captionText-1="TABLE 3. Number of named extant species with described immatures for genera of Ceratopogonidae (taxa of uncertain identity or identified only as &quot;sp.&quot; are not included). Only genera with at least one immature stage known are listed. Percentages of described species within that genus are those for which those stages are known. Number of named species from Borkent (2014), with a few additions as noted in the Taxonomic Discussion under some genera. Taxa are arranged phyletically by subfamily and tribe and within tribes genera are listed alphabetically." httpUri-0="http://table.plazi.org/id/DEA36657BD8030D8FF6B1BF34923E0EC" httpUri-1="http://table.plazi.org/id/DEA36657BDDC3084FF6F1A8D496DE336" pageId="92" pageNumber="93" tableUuid-0="DEA36657BD8030D8FF6B1BF34923E0EC" tableUuid-1="DEA36657BDDC3084FF6F1A8D496DE336">Tables 2–3</tableCitation>
). All the species placed here in the new genus
<taxonomicName id="4DDC4D5CBD72302AFE981CB14CAEE4B9" authorityName="Borkent" authorityYear="2014" box="[352,478,1700,1725]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFE981CB14CAEE4B9" box="[352,478,1700,1725]" italics="true" pageId="92" pageNumber="93">Anebomyia</emphasis>
</taxonomicName>
were previously in
<taxonomicName id="4DDC4D5CBD72302AFD3A1CB14E1AE4B9" box="[706,874,1700,1725]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFD3A1CB14E1AE4B9" box="[706,874,1700,1725]" italics="true" pageId="92" pageNumber="93">Mallochohelea</emphasis>
</taxonomicName>
. The new genus is recognized because the newly included species are not related to those now restricted to
<taxonomicName id="4DDC4D5CBD72302AFC9C1CD2497CE4E4" box="[868,1036,1735,1760]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFC9C1CD2497CE4E4" box="[868,1036,1735,1760]" italics="true" pageId="92" pageNumber="93">Mallochohelea</emphasis>
</taxonomicName>
, a genus most closely related to
<taxonomicName id="4DDC4D5CBD72302AFF6F1CF94C58E501" box="[151,296,1772,1797]" class="Insecta" family="Ceratopogonidae" genus="Jenkinshelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFF6F1CF94C58E501" box="[151,296,1772,1797]" italics="true" pageId="92" pageNumber="93">Jenkinshelea</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD72302AFECD1CF84CC2E500" box="[309,434,1773,1796]" class="Insecta" family="Ceratopogonidae" genus="Macropeza" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFECD1CF84CC2E500" box="[309,434,1773,1796]" italics="true" pageId="92" pageNumber="93">Macropeza</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD72302AFE381CF94F5FE501" box="[448,559,1772,1797]" class="Insecta" family="Ceratopogonidae" genus="Probezzia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFE381CF94F5FE501" box="[448,559,1772,1797]" italics="true" pageId="92" pageNumber="93">Probezzia</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD72302AFD9E1CF94FA9E501" box="[614,729,1772,1797]" class="Insecta" family="Ceratopogonidae" genus="Neobezzia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFD9E1CF94FA9E501" box="[614,729,1772,1797]" italics="true" pageId="92" pageNumber="93">Neobezzia</emphasis>
</taxonomicName>
(see phylogenetic analysis below,
<figureCitation id="12E72A5ABD72302AFBA51CF949C0E501" box="[1117,1200,1772,1797]" captionStart="FIGURE 81" captionStartId="326.[151,250,1891,1913]" captionTargetBox="[165,1420,194,1823]" captionTargetId="figure-107@326.[139,1458,177,1859]" captionTargetPageId="326" captionText="FIGURE 81. Continuation of Majority rule (50%) cladogram of the genera of derived lineages of Ceratopogonidae. Numbers below each branch represent character state changes as described in the text and the matrix in Table 4. For characters with multiple states the character state changed to is indicated in superscript behind the character number. Bremer support values are shown above the branches. The root of this tree is a continuation from &quot;All remaining Ceratopogonidae&quot; as shown in Figure 80." figureDoi="http://doi.org/10.5281/zenodo.4949344" httpUri="https://zenodo.org/record/4949344/files/figure.png" pageId="92" pageNumber="93">Fig. 81</figureCitation>
).
</paragraph>
<caption id="DEA36657BD72302AFF3F1D21487CE574" ID-Table-UUID="DEA36657BD72302AFF3F1D21487CE574" httpUri="http://table.plazi.org/id/DEA36657BD72302AFF3F1D21487CE574" inLine="true" pageId="92" pageNumber="93" startId="92.[920,978,1879,1904]" targetBox="[159,967,1958,2025]" targetIsTable="true" targetPageId="92">
<paragraph id="8A6336DFBD72302AFF3F1D21487CE574" blockId="92.[151,1436,1844,1904]" pageId="92" pageNumber="93">There are differences between the two genera in both male and female adults and in pupae, with most of the adult features not yet interpreted cladistically (but see character 85). They are summarized as follows:</paragraph>
</caption>
<paragraph id="8A6336DFBD72302AFF671DB34EB7E5ED" pageId="92" pageNumber="93">
<table id="F8DCC47FBD72CF89FF671DB34EB7E5ED" box="[159,967,1958,2025]" gridcols="2" gridrows="2" pageId="92" pageNumber="93">
<tr id="34EC349DBD72CF89FF671DB34EB7E5BB" box="[159,967,1958,1983]" gridrow="0" pageId="92" pageNumber="93">
<td id="773D5DE1BD72CF89FF671DB34CC2E5BB" box="[159,434,1958,1983]" gridcol="0" gridrow="0" pageId="92" pageNumber="93">
<taxonomicName id="4DDC4D5CBD72302AFF671DB34C6DE5BB" authorityName="Borkent" authorityYear="2014" box="[159,285,1958,1983]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">Anebomyia</taxonomicName>
</td>
<td id="773D5DE1BD72CF89FCE81DB34EB7E5BB" box="[784,967,1958,1983]" gridcol="1" gridrow="0" pageId="92" pageNumber="93">
<taxonomicName id="4DDC4D5CBD72302AFCE81DB34EC8E5BB" box="[784,952,1958,1983]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD72302AFCE81DB34EC8E5BB" box="[784,952,1958,1983]" italics="true" pageId="92" pageNumber="93">Mallochohelea</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="34EC349DBD72CF89FF671DC54EB7E5ED" box="[159,967,2000,2025]" gridrow="1" pageId="92" pageNumber="93">
<td id="773D5DE1BD72CF89FF671DC54CC2E5ED" box="[159,434,2000,2025]" gridcol="0" gridrow="1" pageId="92" pageNumber="93">male parameres separate</td>
<td id="773D5DE1BD72CF89FCE81DC54EB7E5ED" box="[784,967,2000,2025]" gridcol="1" gridrow="1" pageId="92" pageNumber="93">parameres fused</td>
</tr>
</table>
</paragraph>
<paragraph id="8A6336DFBD73302BFF671A8B485BE3DC" pageId="93" pageNumber="94">
<table id="F8DCC47FBD73CF89FF671A8B48EEE3DC" box="[159,1438,158,472]" gridcols="2" gridrows="8" pageId="93" pageNumber="94">
<tr id="34EC349DBD73CF89FF671A8B48EEE2B3" box="[159,1438,158,183]" gridrow="0" pageId="93" pageNumber="94">
<th id="773D5DE1BD73CF89FF671A8B4E70E2B3" box="[159,768,158,183]" gridcol="0" gridrow="0" pageId="93" pageNumber="94">female with at least forefemur with one or more spines</th>
<th id="773D5DE1BD73CF89FCE81A8B48EEE2B3" box="[784,1438,158,183]" gridcol="1" gridrow="0" pageId="93" pageNumber="94">femora generally without spines (one species with spines)</th>
</tr>
<tr id="34EC349DBD73CF89FF671AD248EEE2E4" box="[159,1438,199,224]" gridrow="1" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671AD24E70E2E4" box="[159,768,199,224]" gridcol="0" gridrow="1" pageId="93" pageNumber="94">female without strong tufts on sternite 8</td>
<td id="773D5DE1BD73CF89FCE81AD248EEE2E4" box="[784,1438,199,224]" gridcol="1" gridrow="1" pageId="93" pageNumber="94">
<specimenCount id="9CDAFD56BD73302BFCEF1AD24E13E2E4" box="[791,867,199,224]" pageId="93" pageNumber="94" type="female">female</specimenCount>
with strong tufts on sternite 8
</td>
</tr>
<tr id="34EC349DBD73CF89FF671AE448EEE30E" box="[159,1438,241,266]" gridrow="2" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671AE44E70E30E" box="[159,768,241,266]" gridcol="0" gridrow="2" pageId="93" pageNumber="94">pupal dorsal apotome with 4 tubercles</td>
<td id="773D5DE1BD73CF89FCE81AE448EEE30E" box="[784,1438,241,266]" gridcol="1" gridrow="2" pageId="93" pageNumber="94">dorsal apotome with 2 tubercles</td>
</tr>
<tr id="34EC349DBD73CF89FF671B0F48EEE337" box="[159,1438,282,307]" gridrow="3" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671B0F4E70E337" box="[159,768,282,307]" gridcol="0" gridrow="3" pageId="93" pageNumber="94">pupa with two elongate anteromedial setae</td>
<td id="773D5DE1BD73CF89FCE81B0F48EEE337" box="[784,1438,282,307]" gridcol="1" gridrow="3" pageId="93" pageNumber="94">one elongate anteromedial seta</td>
</tr>
<tr id="34EC349DBD73CF89FF671B5148EEE359" box="[159,1438,324,349]" gridrow="4" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671B514E70E359" box="[159,768,324,349]" gridcol="0" gridrow="4" pageId="93" pageNumber="94">pupal ecdysial split between eye and antenna</td>
<td id="773D5DE1BD73CF89FCE81B5148EEE359" box="[784,1438,324,349]" gridcol="1" gridrow="4" pageId="93" pageNumber="94">ecdysial split on eye</td>
</tr>
<tr id="34EC349DBD73CF89FF671B7848EEE382" box="[159,1438,365,390]" gridrow="5" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671B784E70E382" box="[159,768,365,390]" gridcol="0" gridrow="5" pageId="93" pageNumber="94">pupal D-5-T elongate seta</td>
<td id="773D5DE1BD73CF89FCE81B7848EEE382" box="[784,1438,365,390]" gridcol="1" gridrow="5" pageId="93" pageNumber="94">D-5-T short, peg-like</td>
</tr>
<tr id="34EC349DBD73CF89FF671B8348EEE3AB" box="[159,1438,406,431]" gridrow="6" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671B834E70E3AB" box="[159,768,406,431]" gridcol="0" gridrow="6" pageId="93" pageNumber="94">pupal D-2-1 and D-3-1 close together</td>
<td id="773D5DE1BD73CF89FCE81B8348EEE3AB" box="[784,1438,406,431]" gridcol="1" gridrow="6" pageId="93" pageNumber="94">D-2-1 and D-3-1 widely spaced</td>
</tr>
<tr id="34EC349DBD73CF89FF671BAA48EEE3DC" box="[159,1438,447,472]" gridrow="7" pageId="93" pageNumber="94">
<td id="773D5DE1BD73CF89FF671BAA4E70E3DC" box="[159,768,447,472]" gridcol="0" gridrow="7" pageId="93" pageNumber="94">pupal abdominal sternites without membranous discs</td>
<td id="773D5DE1BD73CF89FCE81BAA48EEE3DC" box="[784,1438,447,472]" gridcol="1" gridrow="7" pageId="93" pageNumber="94">abdominal sternites 6, 7 with membranous discs</td>
</tr>
</table>
</paragraph>
<paragraph id="8A6336DFBD73302BFF3F181B4FF1E06B" blockId="93.[151,1437,526,1631]" pageId="93" pageNumber="94">
<bibRefCitation id="EE4D4B2EBD73302BFF3F181B4F5FE023" author="Knausenberger" box="[199,559,526,551]" firstAuthor="Knausenberger" page="267 - 277" pageId="93" pageNumber="94" refId="ref188128" refString="Knausenberger, W. I. (1987) Contributions to the autecology and ecosystematics of immature Ceratopogonidae (Diptera), with emphasis on the tribes Heteromyiini and Sphaeromiini in the Middle Atlantic United States. Ph. D. Dissertation, Virginia Polytechnic Institute and State University, Blacksburg, Virginia, xxix + 729 pp." type="book" year="1987">Knausenberger (1987: 267-277)</bibRefCitation>
provided further differences between the two genera (as
<taxonomicName id="4DDC4D5CBD73302BFB4A181B4867E023" baseAuthorityName="Malloch" baseAuthorityYear="1915" box="[1202,1303,526,551]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="albibasis">
<emphasis id="B8A8EACDBD73302BFB4A181B4867E023" box="[1202,1303,526,551]" italics="true" pageId="93" pageNumber="94">albibasis</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD73302BFAB6181A48ECE022" authorityName="Wirth" authorityYear="1962" box="[1358,1436,527,550]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD73302BFAB6181A48ECE022" box="[1358,1436,527,550]" italics="true" pageId="93" pageNumber="94">atripes</emphasis>
</taxonomicName>
groups), including a larval feature. His information was based on Nearctic species and the character states need further testing with extraterritorial material.
</paragraph>
<paragraph id="8A6336DFBD73302BFF3F186F4DAAE127" blockId="93.[151,1437,526,1631]" pageId="93" pageNumber="94">
As indicated above, members of
<taxonomicName id="4DDC4D5CBD73302BFDC1186F4FC7E097" authorityName="Borkent" authorityYear="2014" box="[569,695,634,659]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFDC1186F4FC7E097" box="[569,695,634,659]" italics="true" pageId="93" pageNumber="94">Anebomyia</emphasis>
</taxonomicName>
differ from true
<taxonomicName id="4DDC4D5CBD73302BFC8C186F496CE097" box="[884,1052,634,659]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFC8C186F496CE097" box="[884,1052,634,659]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
by the absence of distinctive tufts of elongate setae on sternite 8 of female adults. In
<taxonomicName id="4DDC4D5CBD73302BFD19188A4E20E0B2" authorityName="Wirth" authorityYear="1962" box="[737,848,671,694]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD73302BFD19188A4E20E0B2" box="[737,848,671,694]" italics="true" pageId="93" pageNumber="94">A. atripes</emphasis>
</taxonomicName>
, elongate setae are absent. In
<taxonomicName id="4DDC4D5CBD73302BFB54188A485FE0B3" box="[1196,1327,670,695]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fluminea">
<emphasis id="B8A8EACDBD73302BFB54188A485FE0B3" box="[1196,1327,670,695]" italics="true" pageId="93" pageNumber="94">A. fluminea</emphasis>
</taxonomicName>
there are elongate setae on sternite 8 but these are not arranged as a tight tuft (as is true in
<taxonomicName id="4DDC4D5CBD73302BFBD618D749A6E0DF" box="[1070,1238,706,731]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFBD618D749A6E0DF" box="[1070,1238,706,731]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
and other related genera; see character 85). Examination of female adult
<taxonomicName id="4DDC4D5CBD73302BFCFA18F24EF0E0FA" box="[770,896,743,766]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="texensis">
<emphasis id="B8A8EACDBD73302BFCFA18F24EF0E0FA" box="[770,896,743,766]" italics="true" pageId="93" pageNumber="94">M. texensis</emphasis>
</taxonomicName>
indicates that it too lacks a well defined sternite 8 tuft.
</paragraph>
<paragraph id="8A6336DFBD73302BFF3F193B4F23E1D3" blockId="93.[151,1437,526,1631]" pageId="93" pageNumber="94">
<bibRefCitation id="EE4D4B2EBD73302BFF3F193B4C2BE143" author="Wirth" box="[199,347,814,839]" firstAuthor="Wirth" pageId="93" pageNumber="94" pagination="272 - 287" refId="ref198995" refString="Wirth, W. W. (1962) A reclassification of the Palpomyia - Bezzia-Macropeza groups, and a revision of the North American Sphaeromiini (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 55, 272 - 287." type="journal article" year="1962">Wirth (1962)</bibRefCitation>
, in erecting
<taxonomicName id="4DDC4D5CBD73302BFE1F193B4FFFE143" box="[487,655,814,839]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFE1F193B4FFFE143" box="[487,655,814,839]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
, pointed out that some of the included species such as
<taxonomicName id="4DDC4D5CBD73302BFB03193A481FE142" authorityName="Wirth" authorityYear="1962" box="[1275,1391,815,838]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD73302BFB03193A481FE142" box="[1275,1391,815,838]" italics="true" pageId="93" pageNumber="94">M. atripes</emphasis>
</taxonomicName>
,
<taxonomicName id="4DDC4D5CBD73302BFA85193A4D83E16E" authorityName="Wirth" authorityYear="1962" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B8A8EACDBD73302BFA85193A4D83E16E" italics="true" pageId="93" pageNumber="94">M. spinipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4DDC4D5CBD73302BFECA19464CC5E16E" box="[306,437,851,874]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="texensis">
<emphasis id="B8A8EACDBD73302BFECA19464CC5E16E" box="[306,437,851,874]" italics="true" pageId="93" pageNumber="94">M. texensis</emphasis>
</taxonomicName>
differed considerably from the genotype (
<taxonomicName id="4DDC4D5CBD73302BFC5219464949E16F" baseAuthorityName="Malloch" baseAuthorityYear="1915" box="[938,1081,850,875]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="albibasis">
<emphasis id="B8A8EACDBD73302BFC5219464949E16F" box="[938,1081,850,875]" italics="true" pageId="93" pageNumber="94">M. albibasis</emphasis>
</taxonomicName>
) and &quot;may, eventually need a separate category&quot;. He further pointed out in a strikingly prescient statement &quot;The discovery of the pupa of this group should also serve greatly to clarify its relationships&quot;. The pupae have indeed provided the synapomorphies to clarify the relationship of these species.
</paragraph>
<paragraph id="8A6336DFBD73302BFF3F19F74D89E6AB" blockId="93.[151,1437,526,1631]" pageId="93" pageNumber="94">
Although
<bibRefCitation id="EE4D4B2EBD73302BFEC519F74CA4E1FF" author="Wirth" box="[317,468,994,1019]" firstAuthor="Wirth" pageId="93" pageNumber="94" pagination="272 - 287" refId="ref198995" refString="Wirth, W. W. (1962) A reclassification of the Palpomyia - Bezzia-Macropeza groups, and a revision of the North American Sphaeromiini (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 55, 272 - 287." type="journal article" year="1962">Wirth (1962)</bibRefCitation>
considered
<taxonomicName id="4DDC4D5CBD73302BFD9D19F64F9CE1FE" authorityName="Wirth" authorityYear="1962" box="[613,748,995,1018]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="spinipes">
<emphasis id="B8A8EACDBD73302BFD9D19F64F9CE1FE" box="[613,748,995,1018]" italics="true" pageId="93" pageNumber="94">M. spinipes</emphasis>
</taxonomicName>
as closely related to
<taxonomicName id="4DDC4D5CBD73302BFC0A19F64914E1FE" authorityName="Wirth" authorityYear="1962" box="[1010,1124,995,1018]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD73302BFC0A19F64914E1FE" box="[1010,1124,995,1018]" italics="true" pageId="93" pageNumber="94">A. atripes</emphasis>
</taxonomicName>
, I have not included it in
<taxonomicName id="4DDC4D5CBD73302BFF6F1E134C65E61B" authorityName="Borkent" authorityYear="2014" box="[151,277,1030,1055]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFF6F1E134C65E61B" box="[151,277,1030,1055]" italics="true" pageId="93" pageNumber="94">Anebomyia</emphasis>
</taxonomicName>
. I have not examined material of this species and know only that the adult female has spines on its femora; the sternite 8 tufts are not described and the male remains unknown. Like this and a number of other species presently in
<taxonomicName id="4DDC4D5CBD73302BFE741E5B4F44E663" box="[396,564,1102,1127]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFE741E5B4F44E663" box="[396,564,1102,1127]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
, reexamination of specimens is needed before the names are potentially transferred to the new genus. Of course, knowing the pupae of further species will certainly clarify their taxonomic position.
</paragraph>
<paragraph id="8A6336DFBD73302BFF3F1EAF4C27E45A" blockId="93.[151,1437,526,1631]" pageId="93" pageNumber="94">
All European species of
<taxonomicName id="4DDC4D5CBD73302BFE0E1EAF4FEEE6D7" box="[502,670,1210,1235]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFE0E1EAF4FEEE6D7" box="[502,670,1210,1235]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
fit the characterization of this genus (Patrycja Dominiak, pers. comm.). The female of the European/Asian species
<taxonomicName id="4DDC4D5CBD73302BFD1E1ECA4EE0E6F3" authorityName="Remm" authorityYear="1980" box="[742,912,1246,1271]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="tianshanica">
<emphasis id="B8A8EACDBD73302BFD1E1ECA4EE0E6F3" box="[742,912,1246,1271]" italics="true" pageId="93" pageNumber="94">M. tianshanica</emphasis>
</taxonomicName>
, however, has spines on the femora and welldeveloped sternite 8 setal tufts (Patrycja Dominiak, pers. comm.) and the male has basally fused parameres (
<bibRefCitation id="EE4D4B2EBD73302BFAA81F174DA3E73B" author="Remm" firstAuthor="Remm" pageId="93" pageNumber="94" pagination="85 - 128" refId="ref193013" refString="Remm, H. (1980) New species of the family Ceratopogonidae (Diptera) from the Middle Asia. Tartu Riikliku Ulikooli Toimetised, 516, 85 - 128 [in Russian, English summary]" type="journal article" year="1980">Remm 1980</bibRefCitation>
). This suggests that
<taxonomicName id="4DDC4D5CBD73302BFE3E1F324F03E73B" authorityName="Remm" authorityYear="1980" box="[454,627,1318,1343]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="tianshanica">
<emphasis id="B8A8EACDBD73302BFE3E1F324F03E73B" box="[454,627,1318,1343]" italics="true" pageId="93" pageNumber="94">M. tianshanica</emphasis>
</taxonomicName>
is a true
<taxonomicName id="4DDC4D5CBD73302BFD121F334EE2E73B" box="[746,914,1318,1343]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFD121F334EE2E73B" box="[746,914,1318,1343]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
but is the only species known with femoral spines.
<bibRefCitation id="EE4D4B2EBD73302BFF131F5F4CB6E767" author="Debenham's" box="[235,454,1354,1379]" firstAuthor="Debenham's" pageId="93" pageNumber="94" pagination="1 - 92" refId="ref180463" refString="Debenham, M. L. (1974) A revision of the Australian and New Guinea predatory Ceratopogonidae (Diptera: Nematocera) of the tribes Heteromyiini and Sphaeromiini. Australian Journal of Zoology, Supplementary Series, 28, 1 - 92." type="journal article" year="1974">Debenham's (1974)</bibRefCitation>
description of
<taxonomicName id="4DDC4D5CBD73302BFD971F5F4E67E767" box="[623,791,1354,1379]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFD971F5F4E67E767" box="[623,791,1354,1379]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
and the included Australian species suggests that they all actually belong within
<taxonomicName id="4DDC4D5CBD73302BFE5A1F7B4F3AE783" box="[418,586,1390,1415]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFE5A1F7B4F3AE783" box="[418,586,1390,1415]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
. The presence of femoral spines on the Afrotropical species
<taxonomicName id="4DDC4D5CBD73302BFAEA1F7A4CD1E7AF" authority="(de Meillon &amp; Wirth 1981)" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="kirki">
<emphasis id="B8A8EACDBD73302BFAEA1F7A481DE783" box="[1298,1389,1390,1415]" italics="true" pageId="93" pageNumber="94">M. kirki</emphasis>
(
<bibRefCitation id="EE4D4B2EBD73302BFA781F7B4CE9E7AF" author="de Meillon &amp; Wirth" firstAuthor="de Meillon" pageId="93" pageNumber="94" pagination="525 - 561" refId="ref181048" refString="De Meillon, B. &amp; Wirth, W. W. (1981) Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by A. L. Dyce. Annals of the Natal Museum, 24, 525 - 561." type="journal article" year="1981">de Meillon &amp; Wirth 1981</bibRefCitation>
)
</taxonomicName>
may indicate they are members of
<taxonomicName id="4DDC4D5CBD73302BFCC11F874EC7E7AF" authorityName="Borkent" authorityYear="2014" box="[825,951,1426,1451]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFCC11F874EC7E7AF" box="[825,951,1426,1451]" italics="true" pageId="93" pageNumber="94">Anebomyia</emphasis>
</taxonomicName>
but the presence or absence of the female sternite 8 setal tufts is unknown and males are undescribed, making its placement uncertain. The African species
<taxonomicName id="4DDC4D5CBD73302BFF6F1FCE4C70E7F6" box="[151,256,1499,1522]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="siricis">
<emphasis id="B8A8EACDBD73302BFF6F1FCE4C70E7F6" box="[151,256,1499,1522]" italics="true" pageId="93" pageNumber="94">M. siricis</emphasis>
</taxonomicName>
has a female with spines on the forefemur and the male has separate parameres; although the condition of the female sternite 8 seta tufts is uncertain, I consider this sufficient evidence to place it in
<taxonomicName id="4DDC4D5CBD73302BFB7C1FEB4872E413" authorityName="Borkent" authorityYear="2014" box="[1156,1282,1534,1559]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFB7C1FEB4872E413" box="[1156,1282,1534,1559]" italics="true" pageId="93" pageNumber="94">Anebomyia</emphasis>
</taxonomicName>
. The Chinese species
<taxonomicName id="4DDC4D5CBD73302BFF161C364C05E43E" box="[238,373,1571,1594]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B8A8EACDBD73302BFF161C364C05E43E" box="[238,373,1571,1594]" italics="true" pageId="93" pageNumber="94">M. yunnana</emphasis>
</taxonomicName>
, known only as a female, has femora spines and lacks the sternite 8 setal tufts, indicating it too is an
<taxonomicName id="4DDC4D5CBD73302BFF2A1C534C20E45B" authorityName="Borkent" authorityYear="2014" box="[210,336,1606,1631]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFF2A1C534C20E45B" box="[210,336,1606,1631]" italics="true" pageId="93" pageNumber="94">Anebomyia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD73302BFD481CA74E0CE4CE" box="[688,892,1714,1739]" pageId="93" pageNumber="94" type="multiple">
<paragraph id="8A6336DFBD73302BFD481CA74E0CE4CE" blockId="93.[688,892,1714,1739]" box="[688,892,1714,1739]" pageId="93" pageNumber="94">
<emphasis id="B8A8EACDBD73302BFD481CA74E03E4CF" bold="true" box="[688,883,1714,1739]" pageId="93" pageNumber="94">Included species</emphasis>
:
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD73302BFF6F1CEF4C24E5EC" pageId="93" pageNumber="94" type="synonymic_list">
<paragraph id="8A6336DFBD73302BFF6F1CEF48E6E514" blockId="93.[151,1435,1786,2024]" box="[151,1430,1786,1808]" pageId="93" pageNumber="94">
<taxonomicName id="4DDC4D5CBD73302BFF6F1CEF4F69E514" authority="(Wirth), 1962: 281" baseAuthorityName="Wirth" baseAuthorityPageNumber="281" baseAuthorityYear="1962" box="[151,537,1786,1808]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD73302BFF6F1CEF4C25E514" box="[151,341,1786,1808]" italics="true" pageId="93" pageNumber="94">Anebomyia atripes</emphasis>
(Wirth), 1962: 281
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD73302BFDD01CEF4FB0E514" box="[552,704,1786,1808]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFDD01CEF4FB0E514" box="[552,704,1786,1808]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
).
<typeStatus id="5567887DBD73302BFD2D1CEE4E77E514" box="[725,775,1787,1808]" pageId="93" pageNumber="94">Type</typeStatus>
locality: New Brunswick,
<collectingRegion id="4818F83DBD73302BFBEF1CEE49FDE514" box="[1047,1165,1787,1808]" country="United States of America" name="New Jersey" pageId="93" pageNumber="94">New Jersey</collectingRegion>
,
<collectingCountry id="F2CB764FBD73302BFB6F1CEF49BDE514" box="[1175,1229,1786,1808]" name="United States of America" pageId="93" pageNumber="94">USA</collectingCountry>
.
<emphasis id="B8A8EACDBD73302BFB2F1CEE48E6E514" bold="true" box="[1239,1430,1786,1808]" pageId="93" pageNumber="94">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD73302BFF6F1D0B4E62E55C" blockId="93.[151,1435,1786,2024]" pageId="93" pageNumber="94">
<taxonomicName id="4DDC4D5CBD73302BFF6F1D0B4FB6E530" authority="(de Meillon &amp; Wirth), 1981: 550" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityPageNumber="550" baseAuthorityYear="1981" box="[151,710,1822,1844]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fluminea">
<emphasis id="B8A8EACDBD73302BFF6F1D0B4C1AE530" box="[151,362,1822,1844]" italics="true" pageId="93" pageNumber="94">Anebomyia fluminea</emphasis>
(de Meillon &amp; Wirth), 1981: 550
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD73302BFD2E1D0B4E1EE530" box="[726,878,1822,1844]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFD2E1D0B4E1EE530" box="[726,878,1822,1844]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
).
<typeStatus id="5567887DBD73302BFC7D1D0A4EC7E530" box="[901,951,1823,1844]" pageId="93" pageNumber="94">Type</typeStatus>
locality: Sand River between White River and Hazyview, E. Transvaal,
<collectingCountry id="F2CB764FBD73302BFE3C1D574F39E55C" box="[452,585,1858,1880]" name="South Africa" pageId="93" pageNumber="94">South Africa</collectingCountry>
.
<emphasis id="B8A8EACDBD73302BFDAB1D564E62E55C" bold="true" box="[595,786,1858,1880]" pageId="93" pageNumber="94">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD73302BFF6F1D73486BE578" blockId="93.[151,1435,1786,2024]" box="[151,1307,1894,1916]" pageId="93" pageNumber="94">
<taxonomicName id="4DDC4D5CBD73302BFF6F1D734F48E578" authority="(de Meillon), 1961: 50" baseAuthorityName="de Meillon" baseAuthorityPageNumber="50" baseAuthorityYear="1961" box="[151,568,1894,1916]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="siricis">
<emphasis id="B8A8EACDBD73302BFF6F1D734C3DE578" box="[151,333,1894,1916]" italics="true" pageId="93" pageNumber="94">Anebomyia siricis</emphasis>
(de Meillon), 1961: 50
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD73302BFDBF1D734FBAE578" baseAuthorityName="de Meillon &amp; Wirth" baseAuthorityYear="1981" box="[583,714,1894,1916]" class="Insecta" family="Ceratopogonidae" genus="Sphaeromias" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFDBF1D734FBAE578" box="[583,714,1894,1916]" italics="true" pageId="93" pageNumber="94">Sphaeromias</emphasis>
</taxonomicName>
).
<typeStatus id="5567887DBD73302BFD271D724E61E578" box="[735,785,1895,1916]" pageId="93" pageNumber="94">Type</typeStatus>
locality: Fenerive,
<collectingCountry id="F2CB764FBD73302BFC2E1D734922E578" box="[982,1106,1894,1916]" name="Madagascar" pageId="93" pageNumber="94">Madagascar</collectingCountry>
.
<emphasis id="B8A8EACDBD73302BFBA41D72486BE578" bold="true" box="[1116,1307,1894,1916]" pageId="93" pageNumber="94">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD73302BFF6F1D9F486EE5A4" blockId="93.[151,1435,1786,2024]" box="[151,1310,1930,1952]" pageId="93" pageNumber="94">
<taxonomicName id="4DDC4D5CBD73302BFF6F1D9F4F53E5A4" authority="(Wirth), 1962: 283" baseAuthorityName="Wirth" baseAuthorityPageNumber="283" baseAuthorityYear="1962" box="[151,547,1930,1952]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="texensis">
<emphasis id="B8A8EACDBD73302BFF6F1D9F4C2FE5A4" box="[151,351,1930,1952]" italics="true" pageId="93" pageNumber="94">Anebomyia texensis</emphasis>
(Wirth), 1962: 283
</taxonomicName>
(
<taxonomicName id="4DDC4D5CBD73302BFDCA1D9F4FBAE5A4" box="[562,714,1930,1952]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFDCA1D9F4FBAE5A4" box="[562,714,1930,1952]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
).
<typeStatus id="5567887DBD73302BFD271D9E4E61E5A4" box="[735,785,1931,1952]" pageId="93" pageNumber="94">Type</typeStatus>
locality: Kerrville,
<collectingRegion id="4818F83DBD73302BFC211D9E4966E5A4" box="[985,1046,1931,1952]" country="United States of America" name="Texas" pageId="93" pageNumber="94">Texas</collectingRegion>
,
<collectingCountry id="F2CB764FBD73302BFBD81D9F4926E5A4" box="[1056,1110,1930,1952]" name="United States of America" pageId="93" pageNumber="94">USA</collectingCountry>
.
<emphasis id="B8A8EACDBD73302BFBA71D9E486EE5A4" bold="true" box="[1119,1310,1930,1952]" pageId="93" pageNumber="94">new combination.</emphasis>
</paragraph>
<paragraph id="8A6336DFBD73302BFF6F1DBB4C24E5EC" blockId="93.[151,1435,1786,2024]" pageId="93" pageNumber="94">
<taxonomicName id="4DDC4D5CBD73302BFF6F1DBB4F76E5C0" authority="(Yu &amp; Zou)" baseAuthorityName="Yu &amp; Zou" box="[151,518,1966,1988]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="yunnana">
<emphasis id="B8A8EACDBD73302BFF6F1DBB4C1EE5C0" box="[151,366,1966,1988]" italics="true" pageId="93" pageNumber="94">Anebomyia yunnana</emphasis>
(Yu &amp; Zou)
</taxonomicName>
, in
<bibRefCitation id="EE4D4B2EBD73302BFDC31DBA4E5AE5C0" author="Yu" box="[571,810,1966,1988]" etAl="et al." firstAuthor="Yu" page="1503" pageId="93" pageNumber="94" refId="ref200768" refString="Yu, Y. - X., Liu, J. - H., Liu, G. - P., Liu, Z. - J., Hao, B. - S., Yan, G. &amp; Zhao, T. - S. (2005) Ceratopogonidae of China, Insecta, Diptera. Volumes 1 - 2. Military Medical Science Press, Beijing, 1699 pp. [in Chinese]" type="book" year="2005">Yu et al. 2005: 1503</bibRefCitation>
(
<taxonomicName id="4DDC4D5CBD73302BFCB81DBB4EA8E5C0" box="[832,984,1966,1988]" class="Insecta" family="Ceratopogonidae" genus="Mallochohelea" kingdom="Animalia" order="Diptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B8A8EACDBD73302BFCB81DBB4EA8E5C0" box="[832,984,1966,1988]" italics="true" pageId="93" pageNumber="94">Mallochohelea</emphasis>
</taxonomicName>
). Malipo,
<collectingRegion id="4818F83DBD73302BFBAC1DBA487DE5C0" box="[1108,1293,1966,1988]" country="China" name="Yunnan" pageId="93" pageNumber="94">Yunnan Province</collectingRegion>
,
<collectingCountry id="F2CB764FBD73302BFAE71DBB482FE5C0" box="[1311,1375,1966,1988]" name="China" pageId="93" pageNumber="94">China</collectingCountry>
.
<emphasis id="B8A8EACDBD73302BFA881DBA4C24E5EC" bold="true" pageId="93" pageNumber="94">new combination.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C2C66554BD703028FF3F1A824CAEE38C" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="8A6336DFBD703028FF3F1A824CAEE38C" blockId="94.[151,1436,151,392]" pageId="94" pageNumber="95">
<emphasis id="B8A8EACDBD703028FF3F1A824F76E2B4" bold="true" box="[199,518,151,176]" pageId="94" pageNumber="95">MATERIAL EXAMINED</emphasis>
:
<taxonomicName id="4DDC4D5CBD703028FDE31A8D4FFDE2AB" baseAuthorityName="Wirth" baseAuthorityYear="1962" box="[539,653,152,175]" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="atripes">
<emphasis id="B8A8EACDBD703028FDE31A8D4FFDE2AB" box="[539,653,152,175]" italics="true" pageId="94" pageNumber="95">A. atripes</emphasis>
</taxonomicName>
: 1 pupal exuviae, Black Lake, North Burgess Township,
<collectingRegion id="4818F83DBD703028FAB81A8248E8E2B4" box="[1344,1432,151,176]" country="Canada" name="Ontario" pageId="94" pageNumber="95">Ontario</collectingRegion>
,
<collectingCountry id="F2CB764FBD703028FF6F1AA94D9EE2D1" box="[151,238,188,213]" name="Canada" pageId="94" pageNumber="95">Canada</collectingCountry>
,
<date id="FE62101FBD703028FEF91AA94CF5E2D1" box="[257,389,188,213]" pageId="94" pageNumber="95" value="1971-06-30">30-VI-1971</date>
(CNCI); 1 pupal exuviae, Patuxent Wildlife Research Center, Prince George County,
<collectingRegion id="4818F83DBD703028FF6F1ACA4C77E2FC" box="[151,263,223,248]" country="United States of America" name="Maryland" pageId="94" pageNumber="95">Maryland</collectingRegion>
,
<collectingCountry id="F2CB764FBD703028FEEA1ACA4C3DE2FC" box="[274,333,223,248]" name="United States of America" pageId="94" pageNumber="95">USA</collectingCountry>
,
<date id="FE62101FBD703028FEA01ACA4CBFE2FC" box="[344,463,223,248]" pageId="94" pageNumber="95" value="1976-06-04">4-VI-1976</date>
(USNM); 2 pupal exuviae, as previous locality,
<date id="FE62101FBD703028FC081ACA4919E2FC" box="[1008,1129,223,248]" pageId="94" pageNumber="95" value="1977-05-24">24-V-1977</date>
(USNM); 3 pupal exuviae, as previous locality,
<date id="FE62101FBD703028FE791B114F75E319" box="[385,517,260,285]" pageId="94" pageNumber="95" value="1977-06-13">13-VI-1977</date>
(USNM); 2 pupal exuviae, as previous locality, 1977 (USNM); 1 pupal exuviae, Lower 3 Runs Creek, Savannah R. Plantation, Barnwell County,
<collectingRegion id="4818F83DBD703028FC7E1B324944E344" box="[902,1076,295,320]" country="United States of America" name="South Carolina" pageId="94" pageNumber="95">South Carolina</collectingRegion>
,
<collectingCountry id="F2CB764FBD703028FBBA1B32490DE344" box="[1090,1149,295,320]" name="United States of America" pageId="94" pageNumber="95">USA</collectingCountry>
,
<date id="FE62101FBD703028FB731B324874E344" box="[1163,1284,295,320]" pageId="94" pageNumber="95" value="1979-02-16">16-II-1979</date>
(WLGC).
<taxonomicName id="4DDC4D5CBD703028FA7C1B3D4D89E361" class="Insecta" family="Ceratopogonidae" genus="Anebomyia" kingdom="Animalia" order="Diptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="fluminea">
<emphasis id="B8A8EACDBD703028FA7C1B3D4D89E361" italics="true" pageId="94" pageNumber="95">A. fluminea</emphasis>
</taxonomicName>
: 1 pupal exuviae (of
<typeStatus id="5567887DBD703028FE0B1B594F29E361" box="[499,601,332,357]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
), Sand River between White River and Hazyview, East Transvaal,
<collectingCountry id="F2CB764FBD703028FAA31B594D92E38C" name="South Africa" pageId="94" pageNumber="95">South Africa</collectingCountry>
,
<date id="FE62101FBD703028FF141B7A4C1CE38C" box="[236,364,367,392]" pageId="94" pageNumber="95" value="1973-12-02">2-XII-1973</date>
(NMSA).
</paragraph>
</subSubSection>
</treatment>
</document>
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