treatments-xml/data/92/71/38/9271386EB246FFBC42979157FE46FAD1.xml

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<mods:title id="AECEE767F41EC89005CF914A3159027D">Redescription and new records of a poorly known leucosiid crab, Pseudophilyra punctulata Chen &amp; Ng, 2003, and description of a new species of Pseudophilyra from Japan (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="EF812DFD4B6332C33D9195BB165FF8E5">Komai, Tomoyuki</mods:namePart>
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<taxonomicName id="DDD8F2FBB246FFB942979157FD55FAA2" ID-CoL="8YYN7" authority="Chen &amp; Ng, 2003" authorityName="Chen &amp; Ng" authorityYear="2003" box="[151,685,1292,1318]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="253" phylum="Arthropoda" rank="species" species="punctulata">
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<emphasis id="28AC556AB246FFB942979157FE36FAA2" bold="true" box="[151,462,1292,1318]" italics="true" pageId="2" pageNumber="253">Pseudophilyra punctulata</emphasis>
<bibRefCitation id="7E49F489B246FFB943D59157FD55FAA2" author="Chen, H. - L. &amp; Ng, P. K. L." box="[469,685,1292,1318]" pageId="2" pageNumber="253" pagination="61 - 69" refId="ref6186" refString="Chen, H. - L. &amp; Ng, P. K. L. (2003) On new species of Leucosiidae (Crustacea: Decapoda: Brachyura) from Singapore and the South China Sea. Raffles Bulletin of Zoology, 51, 61 - 69." type="journal article" year="2003">Chen &amp; Ng, 2003</bibRefCitation>
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<paragraph id="1A678978B246FFB942979174FD4DFAC3" blockId="2.[151,693,1292,1387]" box="[151,693,1326,1351]" pageId="2" pageNumber="253">[New Japanese name: Minami-kobushi-modoki]</paragraph>
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<subSubSection id="52C2DAF3B246FFB942979109FEEAFAEF" box="[151,274,1362,1387]" pageId="2" pageNumber="253" type="description">
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(
<figureCitation id="82E395FDB246FFB9429F9109FEF1FAEF" box="[159,265,1362,1387]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1727,1749]" captionStartId-1="4.[151,250,1946,1968]" captionStartId-2="5.[151,250,1869,1891]" captionStartId-3="6.[151,250,1588,1610]" captionTargetBox-0="[151,1435,483,1702]" captionTargetBox-1="[151,1435,913,1925]" captionTargetBox-2="[223,1347,233,1832]" captionTargetBox-3="[187,1409,226,1539]" captionTargetId-0="figure@3.[151,1435,483,1702]" captionTargetId-1="figure@4.[151,1435,913,1925]" captionTargetId-2="figure@5.[204,1387,193,1853]" captionTargetId-3="figure@6.[150,1436,193,1567]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." captionText-1="FIGURE 2. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." captionText-2="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." captionText-3="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri-0="https://zenodo.org/record/2625333/files/figure.png" httpUri-1="https://zenodo.org/record/2625335/files/figure.png" httpUri-2="https://zenodo.org/record/2625337/files/figure.png" httpUri-3="https://zenodo.org/record/2625339/files/figure.png" pageId="2" pageNumber="253">Figs. 1–4</figureCitation>
)
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<emphasis id="28AC556AB246FFB9429791C1FE66FA34" box="[151,414,1434,1456]" italics="true" pageId="2" pageNumber="253">Pseudophilyra punctulata</emphasis>
<treatmentCitation id="9B79AF69B246FFB943A391C1FD79FA34" author="Chen, H. - L. &amp; Ng, P. K. L." box="[419,641,1434,1456]" page="64" pageId="2" pageNumber="253" year="2003">
<bibRefCitation id="7E49F489B246FFB943A391C1FDA3FA34" author="Chen, H. - L. &amp; Ng, P. K. L." box="[419,603,1434,1456]" pageId="2" pageNumber="253" pagination="61 - 69" refId="ref6186" refString="Chen, H. - L. &amp; Ng, P. K. L. (2003) On new species of Leucosiidae (Crustacea: Decapoda: Brachyura) from Singapore and the South China Sea. Raffles Bulletin of Zoology, 51, 61 - 69." type="journal article" year="2003">Chen &amp; Ng, 2003</bibRefCitation>
: 64
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, figs. 1A, 2 (
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locality: Siloso Beach, Sentosa, 
<collectingCountry id="62CFC9E8B246FFB9467C91C1FB1FFA34" box="[1148,1255,1434,1456]" name="Singapore" pageId="2" pageNumber="253">Singapore</collectingCountry>
).
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<treatmentCitationGroup id="3AC8AE56B246FFB9429791E2FC83FA4B" box="[151,891,1465,1487]" pageId="2" pageNumber="253">
<taxonomicName id="DDD8F2FBB246FFB9429791E2FE72FA4B" box="[151,394,1465,1487]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="253" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="28AC556AB246FFB9429791E2FE72FA4B" box="[151,394,1465,1487]" italics="true" pageId="2" pageNumber="253">Pseudophilyra punctata</emphasis>
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(sic).— 
<treatmentCitation id="9B79AF69B246FFB943DA91E2FD6CFA4B" author="Ng, P. K. L. &amp; Guinot, D &amp; Davie, P. J. F." box="[474,660,1465,1487]" page="93" pageId="2" pageNumber="253" year="2008">
<bibRefCitation id="7E49F489B246FFB943DA91E2FD89FA4B" author="Ng, P. K. L. &amp; Guinot, D &amp; Davie, P. J. F." box="[474,625,1465,1487]" pageId="2" pageNumber="253" pagination="1 - 286" refId="ref6937" refString="Ng, P. K. L., Guinot, D &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="28AC556AB246FFB9400291E1FDCDFA4B" box="[514,565,1465,1487]" italics="true" pageId="2" pageNumber="253">et al.</emphasis>
2008
</bibRefCitation>
: 93
</treatmentCitation>
(list; no new locality).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="28AC556AB246FFB9429791A7FE84F991" bold="true" box="[151,380,1532,1557]" pageId="2" pageNumber="253">Material examined</emphasis>
. 
<materialsCitation id="AAB08325B246FFB9438C91A7FD52F9BC" ID-GBIF-Occurrence="2234227844" collectingDate="1987-12-21" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; S. L. Tay" country="Singapore" location="Singapore" pageId="2" pageNumber="253" specimenCode="ZRC 1993.92" specimenCount="1" typeStatus="holotype">
<typeStatus id="C56337DAB246FFB9438C91A7FE01F991" box="[396,505,1532,1557]" pageId="2" pageNumber="253" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="4A7E2103B246FFB9400691A7FD5CF990" box="[518,676,1532,1556]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="253">ZRC 1993.92</specimenCode>
, immature male (5.8×5.0 mm), Siloso Beach, Sentosa, 
<collectingCountry id="62CFC9E8B246FFB9472391A7FA60F991" box="[1315,1432,1532,1557]" name="Singapore" pageId="2" pageNumber="253">Singapore</collectingCountry>
, coll. 
<collectorName id="B72DECAEB246FFB942CF927BFEC6F9BC" box="[207,318,1568,1592]" pageId="2" pageNumber="253">P.K.L. Ng</collectorName>
&amp; 
<collectorName id="B72DECAEB246FFB94363927BFE3BF9BC" box="[355,451,1568,1592]" pageId="2" pageNumber="253">S.L. Tay</collectorName>
, 
<date id="6E66AFB8B246FFB943CE927BFD5EF9BC" box="[462,678,1567,1592]" pageId="2" pageNumber="253" value="1987-12-21">
<collectingDate id="7E225650B246FFB943CE927BFD5EF9BC" box="[462,678,1567,1592]" pageId="2" pageNumber="253" value="1987-12-21">21 December 1987</collectingDate>
</date>
.
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</paragraph>
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<materialsCitation id="AAB08325B246FFB942C7921EFE7AF904" ID-GBIF-Occurrence="2234228077" collectingDate="1992-08-01" collectionCode="ZRC" collectorName="E. Low &amp; Others" country="Japan" location="Changi Point" municipality="Singapore" pageId="2" pageNumber="253" specimenCode="ZRC 1995.550, 1" specimenCount="1" typeStatus="paratype">
<typeStatus id="C56337DAB246FFB942C7921EFED6F9D9" box="[199,302,1605,1629]" pageId="2" pageNumber="253" type="paratype">Paratype</typeStatus>
: 
<specimenCode id="4A7E2103B246FFB94339921FFE1BF9D9" box="[313,483,1604,1629]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="253">ZRC 1995.550</specimenCode>
, 
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immature female (6.0× 
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), 
<location id="1F07DFA3B246FFB94178921FFBF7F9D9" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB94178921FFBF7F9D9" box="[888,1039,1604,1629]" country="Japan" municipality="Singapore" name="Changi Point" pageId="2" pageNumber="253">Changi Point</location>
, 
<collectingMunicipality id="FA031302B246FFB9461B921FFB77F9D9" box="[1051,1167,1604,1629]" pageId="2" pageNumber="253">Singapore</collectingMunicipality>
, coll. 
<collectorName id="B72DECAEB246FFB946D4921EFAD1F9D8" box="[1236,1321,1605,1628]" pageId="2" pageNumber="253">E. Low</collectorName>
, 
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<collectingDate id="7E225650B246FFB94735921FFF2BF904" pageId="2" pageNumber="253" value="1992-08-01">1 August 1992</collectingDate>
</date>
. 
<collectorName id="B72DECAEB246FFB942DE923CFED5F904" box="[222,301,1639,1664]" pageId="2" pageNumber="253">Others</collectorName>
: 
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<collectingCountry id="62CFC9E8B246FFB94339923CFE7AF904" box="[313,386,1639,1664]" name="Japan" pageId="2" pageNumber="253">Japan</collectingCountry>
</emphasis>
</materialsCitation>
: 
<materialsCitation id="AAB08325B246FFB943939233FADCF921" ID-GBIF-Occurrence="2234227853" collectingDate="2001-07-08" collectionCode="CBM-ZC" collectorName="T. Komai" country="Japan" county="Ryukyu Islands" location="Uehara" municipality="Yaeyama Islands" pageId="2" pageNumber="253" specimenCode="CBM-ZC 7024, 1" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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, 
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immature male (6.1× 
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), 
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(8.2× 
<quantity id="DD20249DB246FFB946699233FB3FF904" box="[1129,1223,1640,1664]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="253" unit="mm" value="7.4">7.4 mm</quantity>
), 
<location id="1F07DFA3B246FFB946DB923CFAD7F904" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB946DB923CFAD7F904" box="[1243,1327,1639,1664]" country="Japan" county="Ryukyu Islands" municipality="Yaeyama Islands" name="Uehara" pageId="2" pageNumber="253">Uehara</location>
, 
<location id="1F07DFA3B246FFB9473B923CFF26F921" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB9473B923CFF26F921" country="Japan" county="Ryukyu Islands" municipality="Yaeyama Islands" name="Iriomote Island" pageId="2" pageNumber="253">Iriomote Island</location>
, 
<collectingMunicipality id="FA031302B246FFB942EB92D6FE54F921" box="[235,428,1676,1701]" pageId="2" pageNumber="253">Yaeyama Islands</collectingMunicipality>
, 
<collectingCounty id="F306F1F4B246FFB943B992D7FD93F921" box="[441,619,1676,1701]" pageId="2" pageNumber="253">Ryukyu Islands</collectingCounty>
, subtidal sea grass beds, 
<date id="6E66AFB8B246FFB9418992D7FBEAF920" box="[905,1042,1676,1701]" pageId="2" pageNumber="253" value="2001-07-08">
<collectingDate id="7E225650B246FFB9418992D7FBEAF920" box="[905,1042,1676,1701]" pageId="2" pageNumber="253" value="2001-07-08">8 July 2001</collectingDate>
</date>
, 
<collectingMethod id="C399F16FB246FFB9461F92D7FB88F920" box="[1055,1136,1676,1701]" pageId="2" pageNumber="253">dip net</collectingMethod>
, coll. 
<collectorName id="B72DECAEB246FFB946B692D6FADCF921" box="[1206,1316,1676,1701]" pageId="2" pageNumber="253">T. Komai</collectorName>
</materialsCitation>
; 
<materialsCitation id="AAB08325B246FFB9473192D7FD7AF94C" ID-GBIF-Occurrence="2234228086" collectingDate="2001-07-08" collectionCode="CBM-ZC" collectorName="T. Komai" country="Japan" county="Ryukyu Islands" location="Uehara" municipality="Yaeyama Islands" pageId="2" pageNumber="253" specimenCode="CBM-ZC 7074" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCode id="4A7E2103B246FFB9473192D7FF2AF94C" collectionCode="CBM-ZC" pageId="2" pageNumber="253">CBM-ZC 7074</specimenCode>
, 
<specimenCount id="0CDE42F1B246FFB942E392EBFEB1F94C" box="[227,329,1711,1736]" pageId="2" pageNumber="253" type="female">1 female</specimenCount>
(8.7× 
<quantity id="DD20249DB246FFB9439192EBFE0AF94C" box="[401,498,1712,1736]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="253" unit="mm" value="7.7">7.7 mm</quantity>
), same data
</materialsCitation>
; 
<materialsCitation id="AAB08325B246FFB9409492EBFB4DF969" ID-GBIF-Occurrence="2234227857" collectingDate="2001-07-01" collectionCode="CBM-ZC" collectorName="T. Komai." country="Philippines" county="Yaeyama Islands" location="Shiraho" municipality="Ishigaki Island" pageId="2" pageNumber="253" specimenCode="CBM-ZC 7053" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCode id="4A7E2103B246FFB9409492EBFCBFF94C" box="[660,839,1711,1736]" collectionCode="CBM-ZC" pageId="2" pageNumber="253">CBM-ZC 7053</specimenCode>
, 
<specimenCount id="0CDE42F1B246FFB9415792EBFC46F94C" box="[855,958,1711,1736]" pageId="2" pageNumber="253" type="female">1 female</specimenCount>
(8.2× 
<quantity id="DD20249DB246FFB9460592EBFB9FF94C" box="[1029,1127,1712,1736]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="253" unit="mm" value="7.4">7.4 mm</quantity>
), 
<location id="1F07DFA3B246FFB9467F92F4FB21F94C" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB9467F92F4FB21F94C" box="[1151,1241,1711,1736]" country="Philippines" county="Yaeyama Islands" municipality="Ishigaki Island" name="Shiraho" pageId="2" pageNumber="253">Shiraho</location>
, 
<collectingMunicipality id="FA031302B246FFB946EA92F4FA60F94C" box="[1258,1432,1711,1736]" pageId="2" pageNumber="253">Ishigaki Island</collectingMunicipality>
, 
<collectingCounty id="F306F1F4B246FFB94297928EFEA1F969" box="[151,345,1748,1773]" pageId="2" pageNumber="253">Yaeyama Islands</collectingCounty>
, subtidal sea grass beds, 
<date id="6E66AFB8B246FFB9407F928FFCF2F968" box="[639,778,1748,1773]" pageId="2" pageNumber="253" value="2001-07-01">
<collectingDate id="7E225650B246FFB9407F928FFCF2F968" box="[639,778,1748,1773]" pageId="2" pageNumber="253" value="2001-07-01">1 July 2001</collectingDate>
</date>
, 
<collectingMethod id="C399F16FB246FFB94117928FFC92F968" box="[791,874,1748,1773]" pageId="2" pageNumber="253">dip net</collectingMethod>
, coll. 
<collectorName id="B72DECAEB246FFB941B2928EFBDEF969" box="[946,1062,1748,1773]" pageId="2" pageNumber="253">T. Komai.</collectorName>
<emphasis id="28AC556AB246FFB9462F928FFB4DF969" bold="true" box="[1071,1205,1748,1773]" pageId="2" pageNumber="253">
<collectingCountry id="62CFC9E8B246FFB9462F928FFB4DF969" box="[1071,1205,1748,1773]" name="Philippines" pageId="2" pageNumber="253">Philippines</collectingCountry>
</emphasis>
</materialsCitation>
: 
<materialsCitation id="AAB08325B246FFB946C6928FFA6CF8B1" ID-GBIF-Occurrence="2234227942" collectingDate="2004-06-11" collectionCode="ZRC" collectorName="Panglao Marine Biodiversity Project" country="Thailand" county="Visayas" elevation="2" latitude="9.56" location="Poblacion" longLatPrecision="129" longitude="123.72667" municipality="Panglao" pageId="2" pageNumber="253" specimenCode="ZRC 2018.0257, 1" specimenCount="1" stateProvince="Bohol" typeStatus="paratype">
<specimenCode id="4A7E2103B246FFB946C6928FFA78F969" box="[1222,1408,1748,1773]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="253">ZRC 2018.0257</specimenCode>
, 
<specimenCode id="4A7E2103B246FFB9478E928FFA64F968" box="[1422,1436,1748,1772]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="253">1</specimenCode>
immature male (4.4× 
<quantity id="DD20249DB246FFB9438992ACFE1FF894" box="[393,487,1783,1808]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="253" unit="mm" value="5.2">5.2 mm</quantity>
), station S11, in fine sand and seagrass bed, lagoon off 
<location id="1F07DFA3B246FFB9467992ACFB14F894" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB9467992ACFB14F894" box="[1145,1260,1783,1808]" country="Thailand" county="Visayas" latitude="9.56" longLatPrecision="129" longitude="123.72667" municipality="Panglao" name="Poblacion" pageId="2" pageNumber="253" stateProvince="Bohol">Poblacion</location>
, 
<quantity id="DD20249DB246FFB946FB92A3FAD3F88B" box="[1275,1323,1784,1808]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="253" unit="m" value="2.0">
<elevation id="91F56E4BB246FFB946FB92A3FAD3F88B" box="[1275,1323,1784,1808]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="253" unit="m" value="2.0">2 m</elevation>
</quantity>
, 
<collectingMunicipality id="FA031302B246FFB9473B92ACFA60F894" box="[1339,1432,1783,1808]" pageId="2" pageNumber="253">Panglao</collectingMunicipality>
, 
<collectingRegion id="D81C479AB246FFB942979347FF26F8B1" box="[151,222,1820,1845]" country="Philippines" name="Bohol" pageId="2" pageNumber="253">Bohol</collectingRegion>
, 
<collectingCounty id="F306F1F4B246FFB942EB9347FEBDF8B1" box="[235,325,1820,1845]" pageId="2" pageNumber="253">Visayas</collectingCounty>
, 
<geoCoordinate id="7FECEFBFB246FFB943529347FE3BF8B1" box="[338,451,1820,1845]" direction="north" orientation="latitude" pageId="2" pageNumber="253" precision="92" value="9.56">09°33.6'N</geoCoordinate>
<geoCoordinate id="7FECEFBFB246FFB943CC9347FDB3F8B1" box="[460,587,1820,1845]" direction="east" orientation="longitude" pageId="2" pageNumber="253" precision="92" value="123.72667">123°43.6'E</geoCoordinate>
, 
<date id="6E66AFB8B246FFB940579347FD0AF8B0" box="[599,754,1820,1844]" pageId="2" pageNumber="253" value="2004-06-11">
<collectingDate id="7E225650B246FFB940579347FD0AF8B0" box="[599,754,1820,1844]" pageId="2" pageNumber="253" value="2004-06-11">11 June 2004</collectingDate>
</date>
, coll. 
<collectorName id="B72DECAEB246FFB941389347FB2FF8B1" box="[824,1239,1820,1845]" pageId="2" pageNumber="253">Panglao Marine Biodiversity Project</collectorName>
2004. 
<emphasis id="28AC556AB246FFB947279347FA6CF8B1" bold="true" box="[1319,1428,1820,1845]" pageId="2" pageNumber="253">
<collectingCountry id="62CFC9E8B246FFB947279347FA6CF8B1" box="[1319,1428,1820,1845]" name="Thailand" pageId="2" pageNumber="253">Thailand</collectingCountry>
</emphasis>
</materialsCitation>
: 
<materialsCitation id="AAB08325B246FFB94297931BFE7BF8F9" ID-GBIF-Occurrence="2234227862" collectingDate="1990-10-05" collectionCode="CBM-ZC" collectorName="T. Komai" country="Thailand" location="Ao Tang Khen" pageId="2" pageNumber="253" specimenCode="CBM-ZC 12912" specimenCount="1" specimenCount-male="1" stateProvince="Phuket" typeStatus="paratype">
<specimenCode id="4A7E2103B246FFB94297931BFEABF8DC" box="[151,339,1856,1880]" collectionCode="CBM-ZC" pageId="2" pageNumber="253">CBM-ZC 12912</specimenCode>
, 
<specimenCount id="0CDE42F1B246FFB9435E931BFE51F8DC" box="[350,425,1855,1880]" pageId="2" pageNumber="253" type="male">1 male</specimenCount>
(6.7×6.0 mm), 
<location id="1F07DFA3B246FFB9405B931BFCFBF8DC" LSID="urn:lsid:plazi:treatment:9271386EB246FFBC42979157FE46FAD1:1F07DFA3B246FFB9405B931BFCFBF8DC" box="[603,771,1855,1880]" country="Thailand" name="Ao Tang Khen" pageId="2" pageNumber="253" stateProvince="Phuket">Ao Tang Khen</location>
, 
<collectingRegion id="D81C479AB246FFB9410E9364FCA7F8DC" box="[782,863,1855,1880]" country="Thailand" name="Phuket" pageId="2" pageNumber="253">Phuket</collectingRegion>
, intertidal, sand flat with sea grass beds, 
<date id="6E66AFB8B246FFB9472D9364FF2AF8F8" pageId="2" pageNumber="253" value="1990-10-05">
<collectingDate id="7E225650B246FFB9472D9364FF2AF8F8" pageId="2" pageNumber="253" value="1990-10-05">5 October 1990</collectingDate>
</date>
, coll. 
<collectorName id="B72DECAEB246FFB94315933EFE7BF8F9" box="[277,387,1892,1917]" pageId="2" pageNumber="253">T. Komai</collectorName>
</materialsCitation>
; 
<materialsCitation id="AAB08325B246FFB9438E933FFC7CF824" ID-GBIF-Occurrence="2234227847" collectingDate="1995-10-24" collectionCode="CBM-ZC" collectorName="T. Komai." country="Thailand" location="Ao Tang Khen" pageId="2" pageNumber="253" specimenCode="CBM-ZC 8323, 2" specimenCount="1" stateProvince="Phuket" typeStatus="paratype">
<specimenCode id="4A7E2103B246FFB9438E933FFDC2F8F8" box="[398,570,1892,1916]" collectionCode="CBM-ZC" pageId="2" pageNumber="253">CBM-ZC 8323</specimenCode>
, 
<specimenCode id="4A7E2103B246FFB94046933FFDACF8F8" box="[582,596,1892,1916]" collectionCode="CBM-ZC" pageId="2" pageNumber="253">2</specimenCode>
immature males (4.4× 
<quantity id="DD20249DB246FFB94152933FFC53F8F8" box="[850,939,1892,1916]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="253" unit="mm" value="3.9">3.9 mm</quantity>
, 4.6× 
<quantity id="DD20249DB246FFB941EA933FFBBCF8F8" box="[1002,1092,1892,1917]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="253" unit="mm" value="4.1">4.1 mm</quantity>
), 2 immature females (4.6× 
<quantity id="DD20249DB246FFB94777933FFF3DF81B" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="253" unit="mm" value="4.2">4.2 mm</quantity>
, 4.9× 
<quantity id="DD20249DB246FFB9430293D3FEA5F824" box="[258,349,1928,1952]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="2" pageNumber="253" unit="mm" value="4.4">4.4 mm</quantity>
), same locality, 
<date id="6E66AFB8B246FFB9401193D3FD28F824" box="[529,720,1927,1952]" pageId="2" pageNumber="253" value="1995-10-24">
<collectingDate id="7E225650B246FFB9401193D3FD28F824" box="[529,720,1927,1952]" pageId="2" pageNumber="253" value="1995-10-24">24 October 1995</collectingDate>
</date>
, coll. 
<collectorName id="B72DECAEB246FFB9411393D3FC7CF824" box="[787,900,1927,1952]" pageId="2" pageNumber="253">T. Komai.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B246FFBC42C793F7FE00FD01" lastPageId="7" lastPageNumber="258" pageId="2" pageNumber="253" type="description">
<paragraph id="1A678978B246FFB842C793F7FEC7FE29" blockId="2.[151,1436,1532,2025]" lastBlockId="3.[151,1436,151,429]" lastPageId="3" lastPageNumber="254" pageId="2" pageNumber="253">
<emphasis id="28AC556AB246FFB942C793F7FE8CF841" bold="true" box="[199,372,1964,1989]" pageId="2" pageNumber="253">Redescription.</emphasis>
<emphasis id="28AC556AB246FFB9437F93F7FE04F840" box="[383,508,1964,1988]" italics="true" pageId="2" pageNumber="253">Adult male</emphasis>
. Carapace (
<figureCitation id="82E395FDB246FFB9408A93F7FD25F840" box="[650,733,1964,1989]" captionStart="FIGURE 2" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[151,1435,913,1925]" captionTargetId="figure@4.[151,1435,913,1925]" captionTargetPageId="4" captionText="FIGURE 2. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625335/files/figure.png" pageId="2" pageNumber="253">Figs. 2</figureCitation>
, 
<figureCitation id="82E395FDB246FFB940ED93F7FCE9F840" box="[749,785,1964,1988]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="2" pageNumber="253">3A</figureCitation>
) subpyriform in general outline, 1.12 times longer than broad; dorsal surface convex, glabrous, but coarsely punctate. Front (
<figureCitation id="82E395FDB246FFB941AB938BFC17F86C" box="[939,1007,2000,2025]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="2" pageNumber="253">Fig. 3</figureCitation>
) weakly produced anteriorly, anterior margin trilobate, median lobe subtriangular with rounded apex projecting anteriorly, distinctly exceeding lateral teeth; lateral lobe (inner orbital lobe) blunt, not particularly produced. Orbits small; supraorbital margin strongly oblique, with short, blunt lateral tooth demarcated mesially by deep, narrow longitudinal suture; bilobed anterior margin of efferent branchial channel forming lower orbital margin, inner angle acutely pointed. Hepatic region slightly inflated, forming low, dome-like elevation. Mesogastric region with obsolescent median ridge, shallowly depressed on both sides of ridge. Lateral to posterior margins keel-like, bordered with relatively coarse granules (
<figureCitation id="82E395FDB247FFB8429F9534FF0FFE0C" box="[159,247,367,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="3" pageNumber="254">Fig. 3C</figureCitation>
), with low convexity just below hepatic region (pterygostomial margin); posterior margin bi-keeled, both nearly straight.
</paragraph>
<caption id="4EA7D9F0B247FFB8429792E4FA6DF896" httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="3" pageNumber="254" startId="3.[151,250,1727,1749]" targetBox="[151,1435,483,1702]" targetPageId="3">
<paragraph id="1A678978B247FFB8429792E4FA6DF896" blockId="3.[151,1435,1727,1810]" pageId="3" pageNumber="254">
<emphasis id="28AC556AB247FFB8429792E4FEF4F950" bold="true" box="[151,268,1727,1749]" pageId="3" pageNumber="254">FIGURE 1</emphasis>
. 
<taxonomicName id="DDD8F2FBB247FFB8431992E4FD23F951" authority="Chen &amp; Ng, 2003" authorityName="Chen &amp; Ng" authorityYear="2003" box="[281,731,1727,1749]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="254" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB247FFB8431992E4FDD8F951" box="[281,544,1727,1749]" italics="true" pageId="3" pageNumber="254">Pseudophilyra punctulata</emphasis>
Chen &amp; Ng, 2003
</taxonomicName>
, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view.
</paragraph>
</caption>
<paragraph id="1A678978B247FFB842C79362FAE6F81E" blockId="3.[151,1436,1849,2018]" pageId="3" pageNumber="254">
Eyestalks (
<figureCitation id="82E395FDB247FFB843499362FE51F8D6" box="[329,425,1849,1874]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="3" pageNumber="254">Fig. 3A</figureCitation>
) retractable, subcylindrical, mesial side protruding into minute blunt distal tubercle exceeding cornea. Antennular fossa continuous with orbit, partially sealed by basal plate on antennule; antennules folded slightly obliquely within fossa. Antenna very small, inserted between antennular fossa and orbit.
</paragraph>
<paragraph id="1A678978B247FFBF42C793FEFAE7FE98" blockId="3.[151,1436,1849,2018]" lastBlockId="4.[151,1437,151,861]" lastPageId="4" lastPageNumber="255" pageId="3" pageNumber="254">
Thoracic sternum glabrous. Sternite 3 (
<figureCitation id="82E395FDB247FFB8408E93FEFD11F83A" box="[654,745,1957,1982]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="3" pageNumber="254">Fig. 3D</figureCitation>
) subtriangular, anterior margin produced anteriorly, surface slightly depressed medially. Sternite 4 (
<figureCitation id="82E395FDB247FFB840589392FD49F866" box="[600,689,1993,2018]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="3" pageNumber="254">Fig. 3D</figureCitation>
) largest, anterior and lateral margins coarsely granular, otherwise weakly granular with scattered minute punctae. Sternites 5–8 becoming smaller toward posterior, surfaces generally smooth but with few punctae. Episternite 4 partially fused with sternite 4; episternites 5–8 each with shallowly concave outer margin. Sternopleonal cavity deep, extending onto posterior part of thoracic sternite 3, margins on sternite 4, corresponding to pleomere 6 and telson, bordered with coarse granules (
<figureCitation id="82E395FDB240FFBF46B9955FFAEAFE98" box="[1209,1298,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="4" pageNumber="255">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="1A678978B240FFBF42C7957CFE57FE29" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="255">
Maxilliped 3 (
<figureCitation id="82E395FDB240FFBF4365957CFE41FEC4" box="[357,441,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="4" pageNumber="255">Fig. 3E</figureCitation>
) almost flat, smooth, polished on outer surface. Ischium subrectangular, distinctly longer than wide; merus triangular, tapering distally to subacute tip, subequal in length to ischium measured along mesial margin; lateral margin of ischium and merus minutely beaded. Exopod with distal article spatuliform, lateral margin minutely beaded.
</paragraph>
<paragraph id="1A678978B240FFBF42C795ECFAB0FD49" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="255">
Chelipeds (
<figureCitation id="82E395FDB240FFBF434895ECFE61FE54" box="[328,409,439,464]" captionStart="FIGURE 2" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[151,1435,913,1925]" captionTargetId="figure@4.[151,1435,913,1925]" captionTargetPageId="4" captionText="FIGURE 2. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625335/files/figure.png" pageId="4" pageNumber="255">Figs. 2</figureCitation>
, 
<figureCitation id="82E395FDB240FFBF43A695E3FE1FFE54" box="[422,487,440,464]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="4" pageNumber="255">3F, G</figureCitation>
) symmetrical, 1.4 times as long as carapace, glabrous. Merus subcylindrical, about 3.3 times as long as wide, covered with pearly granules of various sizes except for distal one-third of upper surface and distal two-thirds of lower surfaces where granules almost absent. Carpus short, cup-shaped, inner surface concave; upper inner margin with row of small granules, but otherwise almost smooth. Palm compressed, 2.0 times as long as wide, slightly arcuate on outer (extensor) margin, inner (flexor) margin slightly sinuous; upper surface almost smooth, lower surface with scattered punctae; fixed finger slightly deflexed, nearly straight, terminating in slightly curved, blunt tip; dactylus 0.8 times as long as palm, terminating in slightly curved, blunt tip; both occlusal margins meeting in distal 0.4 and forming oval hiatus in proximal 0.6, each with few, low teeth proximally.
</paragraph>
<paragraph id="1A678978B240FFBF42C7968CFE84FCD9" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="255">
Ambulatory legs (
<figureCitation id="82E395FDB240FFBF4391968CFE2CFD74" box="[401,468,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[151,1435,913,1925]" captionTargetId="figure@4.[151,1435,913,1925]" captionTargetPageId="4" captionText="FIGURE 2. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625335/files/figure.png" pageId="4" pageNumber="255">Fig. 2</figureCitation>
) slender, glabrous, similar in shape, gradually decreasing in length from first to fourth; each merus cylindrical; each carpus as long as or shorter than propodus; propodus somewhat compressed; each dactylus flattened, slightly curved, tapering distally to minute corneous claw, subequal in length to carpus and propodus combined.
</paragraph>
<caption id="4EA7D9F0B240FFBF429793C1FE53F86A" httpUri="https://zenodo.org/record/2625335/files/figure.png" pageId="4" pageNumber="255" startId="4.[151,250,1946,1968]" targetBox="[151,1435,913,1925]" targetPageId="4">
<paragraph id="1A678978B240FFBF429793C1FE53F86A" blockId="4.[151,1436,1946,2030]" pageId="4" pageNumber="255">
<emphasis id="28AC556AB240FFBF429793C1FEF6F82B" bold="true" box="[151,270,1946,1968]" pageId="4" pageNumber="255">FIGURE 2</emphasis>
. 
<taxonomicName id="DDD8F2FBB240FFBF431C93C1FD1DF834" authority="Chen &amp; Ng, 2003" authorityName="Chen &amp; Ng" authorityYear="2003" box="[284,741,1946,1968]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="255" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB240FFBF431C93C1FDDCF834" box="[284,548,1946,1968]" italics="true" pageId="4" pageNumber="255">Pseudophilyra punctulata</emphasis>
Chen &amp; Ng, 2003
</taxonomicName>
, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace.
</paragraph>
</caption>
<caption id="4EA7D9F0B241FFBE42979316FD13F85A" httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="5" pageNumber="256" startId="5.[151,250,1869,1891]" targetBox="[223,1347,233,1832]" targetPageId="5">
<paragraph id="1A678978B241FFBE42979316FD13F85A" blockId="5.[151,1436,1869,2014]" pageId="5" pageNumber="256">
<emphasis id="28AC556AB241FFBE42979316FEF4F8E6" bold="true" box="[151,268,1869,1891]" pageId="5" pageNumber="256">FIGURE 3</emphasis>
. 
<taxonomicName id="DDD8F2FBB241FFBE43189316FD21F8E7" authority="Chen &amp; Ng, 2003" authorityName="Chen &amp; Ng" authorityYear="2003" box="[280,729,1869,1891]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="256" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB241FFBE43189316FDE7F8E7" box="[280,543,1869,1891]" italics="true" pageId="5" pageNumber="256">Pseudophilyra punctulata</emphasis>
Chen &amp; Ng, 2003
</taxonomicName>
, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view.
</paragraph>
</caption>
<caption id="4EA7D9F0B242FFBD4297926FFBF1F903" httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="6" pageNumber="257" startId="6.[151,250,1588,1610]" targetBox="[187,1409,226,1539]" targetPageId="6">
<paragraph id="1A678978B242FFBD4297926FFBF1F903" blockId="6.[151,1435,1588,1671]" pageId="6" pageNumber="257">
<emphasis id="28AC556AB242FFBD4297926FFEE8F9CD" bold="true" box="[151,272,1588,1610]" pageId="6" pageNumber="257">FIGURE 4</emphasis>
. 
<taxonomicName id="DDD8F2FBB242FFBD431F926FFD08F9CE" authority="Chen &amp; Ng, 2003" authorityName="Chen &amp; Ng" authorityYear="2003" box="[287,752,1588,1610]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="257" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB242FFBD431F926FFDD1F9CE" box="[287,553,1588,1610]" italics="true" pageId="6" pageNumber="257">Pseudophilyra punctulata</emphasis>
Chen &amp; Ng, 2003
</taxonomicName>
, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view.
</paragraph>
</caption>
<paragraph id="1A678978B242FFBD42C792F5FA6EF8B0" blockId="6.[151,1437,1710,2023]" pageId="6" pageNumber="257">
Pleon (
<figureCitation id="82E395FDB242FFBD431892F5FE8AF943" box="[280,370,1710,1735]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="6" pageNumber="257">Fig. 3H</figureCitation>
) elongate-triangular, gently curved sternally. Somites 1 and 2 very short, transversely linear. Main fused section consisting of somites 3–6 elongate trapezoidal, slightly constricted at proximal one third; trace of sutures between somites 3/4 and 5/6 still discernible along lateral margin; surface with scattered punctae. Somite 6 with trace of tubercle somewhat proximal to distal margin. Telson elongate-triangular, 1.7 times as long as wide.
</paragraph>
<paragraph id="1A678978B242FFBD42C79365FA6FF840" blockId="6.[151,1437,1710,2023]" pageId="6" pageNumber="257">
Gonopod 1 (
<figureCitation id="82E395FDB242FFBD43539365FE41F8D3" box="[339,441,1854,1879]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="6" pageNumber="257">
Fig. 
<date id="6E66AFB8B242FFBD43889364FE59F8D3" box="[392,417,1855,1879]" pageId="6" pageNumber="257">3I</date>
, J
</figureCitation>
) slender, sinuously curved, gradually tapering, slightly twisted, reaching nearly to suture between thoracic sternites 3/4, deeply bifurcate at distal 0.3; proximal part well calcified, distal branches chitinous; inner branch slightly longer than outer branch, each branch terminating in acute tip. Gonopod 2 (
<figureCitation id="82E395FDB242FFBD46CF93DDFADEF81B" box="[1231,1318,1926,1951]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="6" pageNumber="257">Fig. 3K</figureCitation>
) 0.4 times as long as gonopod 1, slightly curved; proximal part expanded; distal 0.8 slender, not tapering, tip simple, rounded.
</paragraph>
<paragraph id="1A678978B242FFBD42C7938BFAC7F863" blockId="6.[151,1437,1710,2023]" box="[199,1343,1998,2023]" pageId="6" pageNumber="257">
<emphasis id="28AC556AB242FFBD42C7938BFE83F863" box="[199,379,1999,2023]" italics="true" pageId="6" pageNumber="257">Immature males</emphasis>
. Generally similar to adult male. Carapace (
<figureCitation id="82E395FDB242FFBD416B9395FC39F863" box="[875,961,1998,2023]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1749]" captionTargetBox="[151,1435,483,1702]" captionTargetId="figure@3.[151,1435,483,1702]" captionTargetPageId="3" captionText="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="6" pageNumber="257">Fig. 1A</figureCitation>
) 1.11–1.16 times as long as wide.
</paragraph>
<paragraph id="1A678978B243FFBC42C794C3FA6EFF34" blockId="7.[151,1436,151,1365]" box="[199,1430,151,176]" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC42C794C3FEDEFF34" box="[199,294,152,176]" italics="true" pageId="7" pageNumber="258">Females</emphasis>
. Carapace (
<figureCitation id="82E395FDB243FFBC43AA94CCFDF3FF34" box="[426,523,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1749]" captionTargetBox="[151,1435,483,1702]" captionTargetId="figure@3.[151,1435,483,1702]" captionTargetPageId="3" captionText="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="7" pageNumber="258">Figs. 1C</figureCitation>
, 
<figureCitation id="82E395FDB243FFBC401794C3FDC3FF34" box="[535,571,152,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="7" pageNumber="258">4A</figureCitation>
) 1.10–1.12 times as long as wide. Front slightly less produced than in males.
</paragraph>
<paragraph id="1A678978B243FFBC42C794E7FD36FE29" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
Exposed lateral part of thoracic sternum very narrow, smooth, forming outer wall of deeply excavated sternopleonal cavity; sutures between sternites 4/5, 5/6, 6/7 and 7/8 distinct; sternites 1–3 fused, deeply depressed below to accommodate telson. Sternopleonal cavity (
<figureCitation id="82E395FDB243FFBC4102955FFCA5FE98" box="[770,861,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="7" pageNumber="258">Fig. 4B</figureCitation>
) circular in outline, margins distinctly delimited; sutures between sternites 4/5, 5/6, 6/7, and 7/8 widely interrupted medially; median suture absent; sternite 4 large, occupying anterior half of sternopleonal cavity, with scattered short, occasionally feathered, setae. Vulvae (
<figureCitation id="82E395FDB243FFBC47419517FF53FE0C" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="7" pageNumber="258">Fig. 4B, C</figureCitation>
) small, located just posterior to mesial end of suture between sternites 5/6, outline generally semicircular, outer margin produced toward mesial into rounded lobe.
</paragraph>
<paragraph id="1A678978B243FFBC42C795ECFC3AFE71" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
Cheliped (
<figureCitation id="82E395FDB243FFBC433D95ECFE40FE54" box="[317,440,439,464]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1749]" captionTargetBox="[151,1435,483,1702]" captionTargetId="figure@3.[151,1435,483,1702]" captionTargetPageId="3" captionText="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="7" pageNumber="258">Fig. 1C, D</figureCitation>
) slightly slender than that of males; palm (
<figureCitation id="82E395FDB243FFBC41A595ECFC06FE54" box="[933,1022,439,464]" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="7" pageNumber="258">Fig. 4D</figureCitation>
) 2.0 times as long as wide; occlusal margins of fingers without conspicuous teeth proximal to meeting point.
</paragraph>
<paragraph id="1A678978B243FFBC42C795A4FC31FDE4" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
Pleon of adults (
<figureCitation id="82E395FDB243FFBC438495A4FE24FD9C" box="[388,476,511,536]" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="7" pageNumber="258">Fig. 4C</figureCitation>
) broad, dome-like; somites 1 and 2 short, transversely band-shaped, somite 2 about twice longer than somite 1; main fused section (somites 3–6) ovoid, no trace of sutures. Telson small, subsemicircular, exceeding beyond anterior margin of thoracic sternite 3.
</paragraph>
<paragraph id="1A678978B243FFBC42C79637FE00FD01" blockId="7.[151,1436,151,1365]" box="[199,504,620,645]" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC42C79637FE95FD01" bold="true" box="[199,365,620,645]" pageId="7" pageNumber="258">Colour in life.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B243FFBC42C796D4FB26FC91" pageId="7" pageNumber="258" type="distribution">
<paragraph id="1A678978B243FFBC42C796D4FB26FC91" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC42C796D4FE09FD2C" bold="true" box="[199,497,655,680]" pageId="7" pageNumber="258">Distribution and habitat.</emphasis>
Originally described from 
<collectingCountry id="62CFC9E8B243FFBC412196D4FC6EFD2C" box="[801,918,655,680]" name="Singapore" pageId="7" pageNumber="258">Singapore</collectingCountry>
; now newly recorded from 
<collectingRegion id="D81C479AB243FFBC46CA96D4FAE0FD2C" box="[1226,1304,655,680]" country="Thailand" name="Phuket" pageId="7" pageNumber="258">Phuket</collectingRegion>
(
<collectingCountry id="62CFC9E8B243FFBC472796D4FA68FD2C" box="[1319,1424,655,680]" name="Thailand" pageId="7" pageNumber="258">Thailand</collectingCountry>
), 
<collectingRegion id="D81C479AB243FFBC429796EFFF23FD49" box="[151,219,692,717]" country="Philippines" name="Bohol" pageId="7" pageNumber="258">Bohol</collectingRegion>
(
<collectingCountry id="62CFC9E8B243FFBC42EB96EFFE95FD49" box="[235,365,692,717]" name="Philippines" pageId="7" pageNumber="258">Philippines</collectingCountry>
) and Yaeyama Islands, Ryukyu Islands (
<collectingCountry id="62CFC9E8B243FFBC413896EFFC86FD48" box="[824,894,692,716]" name="Japan" pageId="7" pageNumber="258">Japan</collectingCountry>
); intertidal to shallow subtidal. Specimens from the Ryukyu Islands and Phuket were collected from intertidal sea grass beds formed on sand flats. The 
<typeStatus id="C56337DAB243FFBC473A968CFA64FD74" box="[1338,1436,727,752]" pageId="7" pageNumber="258" type="holotype">holotype</typeStatus>
male was collected from sandy intertidal areas at the edge of a coral reef near patches of seagrass.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B243FFBC42C79744FE46FAD1" pageId="7" pageNumber="258" type="discussion">
<paragraph id="1A678978B243FFBC42C79744FC0CFC04" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC42C79744FEC3FCBC" bold="true" box="[199,315,799,824]" pageId="7" pageNumber="258">Remarks.</emphasis>
As noted above, the type series of 
<taxonomicName id="DDD8F2FBB243FFBC40F8977BFBD9FCBC" authorityName="Chen &amp; Ng" authorityYear="2003" box="[760,1057,800,824]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB243FFBC40F8977BFBD9FCBC" box="[760,1057,800,824]" italics="true" pageId="7" pageNumber="258">Pseudophilyra punctulata</emphasis>
</taxonomicName>
consists only of two immature specimens (
<figureCitation id="82E395FDB243FFBC431B971FFEA4FCD8" box="[283,348,836,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1749]" captionTargetBox="[151,1435,483,1702]" captionTargetId="figure@3.[151,1435,483,1702]" captionTargetPageId="3" captionText="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="7" pageNumber="258">Fig. 1</figureCitation>
). The 
<typeStatus id="C56337DAB243FFBC43A6971FFDF0FCD9" box="[422,520,836,861]" pageId="7" pageNumber="258" type="holotype">holotype</typeStatus>
is a male, but the gonopods are not fully developed. The 
<typeStatus id="C56337DAB243FFBC468A971EFB12FCD9" box="[1162,1258,837,861]" pageId="7" pageNumber="258" type="paratype">paratype</typeStatus>
is a female, but the pleon is narrowly triangular (
<figureCitation id="82E395FDB243FFBC4007973CFDA7FC04" box="[519,607,871,896]" captionStart="FIGURE 1" captionStartId="3.[151,250,1727,1749]" captionTargetBox="[151,1435,483,1702]" captionTargetId="figure@3.[151,1435,483,1702]" captionTargetPageId="3" captionText="FIGURE 1. Pseudophilyra punctulata Chen &amp; Ng, 2003, type specimens. A, ZRC 1993.92, holotype, immature male (5.8×5.0 mm), Singapore, carapace in dorsal view (pereopods all detached); B, same, thoracic sternum and pleon in ventral view; C, ZRC 1995.550, paratype, immature female (6.0×5.4 mm), Singapore, habitus in dorsal view; D, same, habitus in ventral view." httpUri="https://zenodo.org/record/2625333/files/figure.png" pageId="7" pageNumber="258">Fig. 1D</figureCitation>
), indicating that it is also immature.
</paragraph>
<paragraph id="1A678978B243FFBC42C797D7FA6DFBB0" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
The immature specimens from 
<collectingRegion id="D81C479AB243FFBC402797D7FD8DFC21" box="[551,629,908,933]" country="Thailand" name="Phuket" pageId="7" pageNumber="258">Phuket</collectingRegion>
(CBM-ZC 8 323) and Panglao (ZRC 2018.0257) are very similar to the 
<typeStatus id="C56337DAB243FFBC429797EBFF3FFC4C" box="[151,199,944,968]" pageId="7" pageNumber="258">type</typeStatus>
specimens. The adult male specimen from 
<collectingRegion id="D81C479AB243FFBC40CF97F4FCD8FC4C" box="[719,800,943,968]" country="Thailand" name="Phuket" pageId="7" pageNumber="258">Phuket</collectingRegion>
, 
<collectingCountry id="62CFC9E8B243FFBC413097F4FC6CFC4C" box="[816,916,943,968]" name="Thailand" pageId="7" pageNumber="258">Thailand</collectingCountry>
(CBM-ZC 12912), collected from the same locality as the four immature specimens (CBM-ZC 8323) enables us to assess adult male characters, including the gonopod 1 morphology. This species characteristically has a deeply bifurcate distal part of the gonopod 1, and this character is also seen in 
<taxonomicName id="DDD8F2FBB243FFBC43A49046FDF0FBB0" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[420,520,1052,1076]" class="Crustacea" genus="Pseudopilyra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB243FFBC43A49046FDF0FBB0" box="[420,520,1052,1076]" italics="true" pageId="7" pageNumber="258">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB243FFBC400E9047FDB5FBB1" bold="true" box="[526,589,1052,1077]" pageId="7" pageNumber="258">
<taxonomicNameLabel id="339FE811B243FFBC400E9047FDB5FBB1" box="[526,589,1052,1077]" pageId="7" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="DDD8F2FBB243FFBC40829046FCF3FBB0" authorityName="Miers" authorityYear="1879" box="[642,779,1052,1076]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB243FFBC40829046FCF3FBB0" box="[642,779,1052,1076]" italics="true" pageId="7" pageNumber="258">P. tridentata</emphasis>
</taxonomicName>
among known congeners (see account of 
<taxonomicName id="DDD8F2FBB243FFBC46DC9046FAB8FBB0" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[1244,1344,1052,1076]" class="Crustacea" genus="Pseudopilyra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB243FFBC46DC9046FAB8FBB0" box="[1244,1344,1052,1076]" italics="true" pageId="7" pageNumber="258">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB243FFBC47479047FA7DFBB1" bold="true" box="[1351,1413,1052,1077]" pageId="7" pageNumber="258">
<taxonomicNameLabel id="339FE811B243FFBC47479047FA7DFBB1" box="[1351,1413,1052,1077]" pageId="7" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="1A678978B243FFBC42C79064FCF7FA89" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
<bibRefCitation id="7E49F489B243FFBC42C79064FE60FBDC" author="Chen, H. - L. &amp; Ng, P. K. L." box="[199,408,1087,1112]" pageId="7" pageNumber="258" pagination="61 - 69" refId="ref6186" refString="Chen, H. - L. &amp; Ng, P. K. L. (2003) On new species of Leucosiidae (Crustacea: Decapoda: Brachyura) from Singapore and the South China Sea. Raffles Bulletin of Zoology, 51, 61 - 69." type="journal article" year="2003">Chen &amp; Ng (2003)</bibRefCitation>
compared 
<taxonomicName id="DDD8F2FBB243FFBC4014901AFD5FFBDC" authorityName="Chen &amp; Ng" authorityYear="2003" box="[532,679,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB243FFBC4014901AFD5FFBDC" box="[532,679,1088,1112]" italics="true" pageId="7" pageNumber="258">P. punctulata</emphasis>
</taxonomicName>
mainly with 
<taxonomicName id="DDD8F2FBB243FFBC413A901AFC37FBDC" authorityName="Ihle" authorityYear="1918" box="[826,975,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="28AC556AB243FFBC413A901AFC37FBDC" box="[826,975,1088,1112]" italics="true" pageId="7" pageNumber="258">P. intermedia</emphasis>
</taxonomicName>
but the male gonopod 1 structure is quite different between the two species. In 
<taxonomicName id="DDD8F2FBB243FFBC4041903EFD2FFBF8" authorityName="Ihle" authorityYear="1918" box="[577,727,1124,1148]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="28AC556AB243FFBC4041903EFD2FFBF8" box="[577,727,1124,1148]" italics="true" pageId="7" pageNumber="258">P. intermedia</emphasis>
</taxonomicName>
, the gonopod 1 is strongly twisted and terminates in a simple, slender process, and is never as deeply bifurcate as in 
<taxonomicName id="DDD8F2FBB243FFBC411790D2FC56FB24" authorityName="Chen &amp; Ng" authorityYear="2003" box="[791,942,1160,1184]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB243FFBC411790D2FC56FB24" box="[791,942,1160,1184]" italics="true" pageId="7" pageNumber="258">P. punctulata</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="7E49F489B243FFBC41E890D2FAF7FB24" author="Komatsu, H. &amp; Takeda, M." box="[1000,1295,1159,1184]" pageId="7" pageNumber="258" pagination="267 - 283" refId="ref6441" refString="Komatsu, H. &amp; Takeda, M. (2000) Leucosiid crabs (Crustacea: Decapoda: Brachyura) from the Osumi Islands, southwest Japan, with description of a new species of Cryptocnemus. Species Diversity, 5, 267 - 283. https: // doi. org / 10.12782 / specdiv. 5.267" type="journal article" year="2000">Komatsu &amp; Takeda 2000</bibRefCitation>
: fig. 2k). In addition, 
<taxonomicName id="DDD8F2FBB243FFBC430190F6FE6DFB40" authorityName="Chen &amp; Ng" authorityYear="2003" box="[257,405,1196,1220]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB243FFBC430190F6FE6DFB40" box="[257,405,1196,1220]" italics="true" pageId="7" pageNumber="258">P. punctulata</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="DDD8F2FBB243FFBC402A90F6FD46FB40" authorityName="Ihle" authorityYear="1918" box="[554,702,1196,1220]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="28AC556AB243FFBC402A90F6FD46FB40" box="[554,702,1196,1220]" italics="true" pageId="7" pageNumber="258">P. intermedia</emphasis>
</taxonomicName>
in the better developed frontal median tooth of the carapace, and the median tubercle on the male pleomere 6 is reduced to a trace. In male 
<taxonomicName id="DDD8F2FBB243FFBC41D7908AFB94FB6C" authorityName="Ihle" authorityYear="1918" box="[983,1132,1232,1256]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="28AC556AB243FFBC41D7908AFB94FB6C" box="[983,1132,1232,1256]" italics="true" pageId="7" pageNumber="258">P. intermedia</emphasis>
</taxonomicName>
, there is a distinct tubercle on the pleomere 6 (cf. 
<bibRefCitation id="7E49F489B243FFBC439490AEFD49FA88" author="Komatsu, H. &amp; Takeda, M." box="[404,689,1268,1293]" pageId="7" pageNumber="258" pagination="267 - 283" refId="ref6441" refString="Komatsu, H. &amp; Takeda, M. (2000) Leucosiid crabs (Crustacea: Decapoda: Brachyura) from the Osumi Islands, southwest Japan, with description of a new species of Cryptocnemus. Species Diversity, 5, 267 - 283. https: // doi. org / 10.12782 / specdiv. 5.267" type="journal article" year="2000">Komatsu &amp; Takeda 2000</bibRefCitation>
: fig. 4j).
</paragraph>
<paragraph id="1A678978B243FFBC42C7914CFE46FAD1" blockId="7.[151,1436,151,1365]" pageId="7" pageNumber="258">
Differentiating characters between 
<taxonomicName id="DDD8F2FBB243FFBC405B9142FD09FAB4" authorityName="Chen &amp; Ng" authorityYear="2003" box="[603,753,1304,1328]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB243FFBC405B9142FD09FAB4" box="[603,753,1304,1328]" italics="true" pageId="7" pageNumber="258">P. punctulata</emphasis>
</taxonomicName>
, 
<taxonomicName id="DDD8F2FBB243FFBC41039142FC68FAB4" authorityName="Miers" authorityYear="1879" box="[771,912,1304,1328]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB243FFBC41039142FC68FAB4" box="[771,912,1304,1328]" italics="true" pageId="7" pageNumber="258">P. tridentata</emphasis>
</taxonomicName>
and 
<taxonomicName id="DDD8F2FBB243FFBC41CD9142FBCDFAB4" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[973,1077,1304,1328]" class="Crustacea" genus="Pseudopilyra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB243FFBC41CD9142FBCDFAB4" box="[973,1077,1304,1328]" italics="true" pageId="7" pageNumber="258">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB243FFBC4640914CFB7BFAB4" bold="true" box="[1088,1155,1303,1328]" pageId="7" pageNumber="258">
<taxonomicNameLabel id="339FE811B243FFBC4640914CFB7BFAB4" box="[1088,1155,1303,1328]" pageId="7" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are discussed under the account of 
<taxonomicName id="DDD8F2FBB243FFBC43139166FE80FAD0" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[275,376,1340,1364]" class="Crustacea" genus="Pseudopilyra" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB243FFBC43139166FE80FAD0" box="[275,376,1340,1364]" italics="true" pageId="7" pageNumber="258">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB243FFBC437F9167FE46FAD1" bold="true" box="[383,446,1340,1365]" pageId="7" pageNumber="258">
<taxonomicNameLabel id="339FE811B243FFBC437F9167FE46FAD1" box="[383,446,1340,1365]" pageId="7" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>