treatments-xml/data/92/71/38/9271386EB243FFB4429791FCFE38FE4B.xml
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<document id="9CF0BA4AD80A7838C91F435A77BE2EAE" ID-CLB-Dataset="27488" ID-DOI="10.11646/zootaxa.4550.2.6" ID-GBIF-Dataset="361ee2d4-577a-41e7-a953-fe213bca92dd" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625331" ID-ZooBank="34D31A7C-D135-4CD2-9882-8AACFDBEF5E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554276958880" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Shimetsugu, Miho &amp; Ng, Peter K. L." docDate="2019" docId="9271386EB243FFB4429791FCFE38FE4B" docLanguage="en" docName="zootaxa.4550.2.6.pdf" docOrigin="Zootaxa 4550 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudophilyra parilis Komai &amp; Shimetsugu &amp; Ng 2019, n. sp." docType="treatment" docUuid="6C966DBD-79D5-4C66-B2BC-FF38C412953D" docUuidSource="ZooBank" docVersion="10" lastPageNumber="266" masterDocId="6E484016B244FFBB4200945BFFF8FF84" masterDocTitle="Redescription and new records of a poorly known leucosiid crab, Pseudophilyra punctulata Chen &amp; Ng, 2003, and description of a new species of Pseudophilyra from Japan (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="267" masterPageNumber="251" pageNumber="258" updateTime="1701998482300" updateUser="plazi">
<mods:mods id="96B35F9EDB0E2FF47C4FD30CB0D01B61" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="603C79FD1C272E0FE5E67D35FFFBC4B9">Redescription and new records of a poorly known leucosiid crab, Pseudophilyra punctulata Chen &amp; Ng, 2003, and description of a new species of Pseudophilyra from Japan (Crustacea: Decapoda: Brachyura)</mods:title>
</mods:titleInfo>
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<mods:namePart id="822A6F158CEF12876355D255BB39636C">Komai, Tomoyuki</mods:namePart>
</mods:name>
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<mods:namePart id="0A08A564BB6477D9EF7DA254DD8567A7">Shimetsugu, Miho</mods:namePart>
</mods:name>
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<mods:namePart id="D38E9E874525CD2B7DE52BB125FD4784">Ng, Peter K. L.</mods:namePart>
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<mods:title id="9ECA477739ADC991E6D4080160944622">Zootaxa</mods:title>
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<mods:part id="F27DCBA7A5B5AAF104B9B6DAE1A2613F">
<mods:date id="55C7B81F2E88C877F4DE29B65BE1B548">2019</mods:date>
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<mods:number id="94D797C1CEA3AB91BDB54C44D4F3E941">2019-01-24</mods:number>
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<mods:number id="6A01F0DF6D3E2A1F8200994997686DF6">4550</mods:number>
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<mods:identifier id="5F404B747F7A99F9F84C839821A34B6B" type="DOI">10.11646/zootaxa.4550.2.6</mods:identifier>
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<treatment id="9271386EB243FFB4429791FCFE38FE4B" ID-DOI="http://doi.org/10.5281/zenodo.5934161" ID-GBIF-Taxon="154535020" ID-Zenodo-Dep="5934161" LSID="urn:lsid:zoobank.org:act:6C966DBD-79D5-4C66-B2BC-FF38C412953D" httpUri="http://treatment.plazi.org/id/9271386EB243FFB4429791FCFE38FE4B" lastPageId="15" lastPageNumber="266" pageId="7" pageNumber="258">
<subSubSection id="52C2DAF3B243FFBC429791FCFD41FA67" pageId="7" pageNumber="258" type="nomenclature">
<paragraph id="1A678978B243FFBC429791FCFE1AFA46" blockId="7.[151,697,1447,1543]" box="[151,482,1447,1474]" pageId="7" pageNumber="258">
<heading id="412F3E14B243FFBC429791FCFE1AFA46" bold="true" box="[151,482,1447,1474]" fontSize="11" level="1" pageId="7" pageNumber="258" reason="1">
<emphasis id="28AC556AB243FFBC429791FCFE1AFA46" bold="true" box="[151,482,1447,1474]" pageId="7" pageNumber="258">
<taxonomicName id="DDD8F2FBB243FFBC429791FCFE61FA45" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[151,409,1447,1473]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="258" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB243FFBC429791FCFE61FA45" bold="true" box="[151,409,1447,1473]" italics="true" pageId="7" pageNumber="258">Pseudophilyra parilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="339FE811B243FFBC43A091F3FE1AFA46" box="[416,482,1448,1474]" pageId="7" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1A678978B243FFBC42979190FD41FA67" blockId="7.[151,697,1447,1543]" box="[151,697,1482,1507]" pageId="7" pageNumber="258">[New Japanese name: Hamabe-kobushi-modoki]</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B243FFBC429791B5FEE9F983" box="[151,273,1518,1543]" pageId="7" pageNumber="258" type="description">
<paragraph id="1A678978B243FFBC429791B5FEE9F983" blockId="7.[151,697,1447,1543]" box="[151,273,1518,1543]" pageId="7" pageNumber="258">
(
<figureCitation id="82E395FDB243FFBC429F91B5FEF1F983" box="[159,265,1518,1543]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="8.[151,250,1941,1963]" captionStartId-1="9.[151,250,1877,1899]" captionStartId-2="10.[151,250,1575,1597]" captionStartId-3="11.[151,250,1966,1988]" captionTargetBox-0="[151,1435,828,1919]" captionTargetBox-1="[214,1360,231,1827]" captionTargetBox-2="[188,1389,280,1509]" captionTargetBox-3="[207,1373,193,1942]" captionTargetId-0="figure@8.[151,1435,828,1919]" captionTargetId-1="figure@9.[196,1379,193,1858]" captionTargetId-2="figure@10.[151,1435,215,1543]" captionTargetId-3="figure@11.[207,1373,193,1942]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="FIGURE 5. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." captionText-1="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." captionText-2="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." captionText-3="FIGURE 8. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratypes, male (7.3×6.3 mm) (A, B), female (7.9×7.3 mm) (C, D), showing colour in life. A, C, habitus, dorsal view; B, D, habitus, ventral view." httpUri-0="https://zenodo.org/record/2625341/files/figure.png" httpUri-1="https://zenodo.org/record/2625343/files/figure.png" httpUri-2="https://zenodo.org/record/2625345/files/figure.png" httpUri-3="https://zenodo.org/record/2625349/files/figure.png" pageId="7" pageNumber="258">Figs. 58</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B243FFBC4297926DFEB2F8CF" pageId="7" pageNumber="258" type="materials_examined">
<paragraph id="1A678978B243FFBC4297926DFB2CF9F7" blockId="7.[151,1437,1590,2011]" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC4297926DFE83F9CB" bold="true" box="[151,379,1590,1615]" pageId="7" pageNumber="258">Material examined</emphasis>
.
<materialsCitation id="AAB08325B243FFBC438A926DFB2CF9F7" ID-GBIF-Occurrence="2234227975" collectingDate="2013-07-23" collectionCode="CBM-ZC" collectorName="S. Kimura." county="Maehama Beach" latitude="34.2685" location="Daiou-cho" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" pageId="7" pageNumber="258" specimenCode="CBM-ZC 14485" specimenCount="1" specimenCount-male="1" stateProvince="Mie" typeStatus="holotype">
<typeStatus id="C56337DAB243FFBC438A926DFE0FF9CB" box="[394,503,1590,1615]" pageId="7" pageNumber="258" type="holotype">Holotype</typeStatus>
:
<specimenCode id="4A7E2103B243FFBC4003926CFD47F9CB" box="[515,703,1590,1615]" collectionCode="CBM-ZC" pageId="7" pageNumber="258">CBM-ZC 14485</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC40CC926CFCE1F9CB" box="[716,793,1590,1615]" pageId="7" pageNumber="258" type="male">1 male</specimenCount>
(7.2×
<quantity id="DD20249DB243FFBC415D926DFC42F9CB" box="[861,954,1590,1615]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="258" unit="mm" value="6.2">6.2 mm</quantity>
),
<collectingCounty id="F306F1F4B243FFBC41CE926DFB76F9CB" box="[974,1166,1590,1615]" pageId="7" pageNumber="258">Maehama Beach</collectingCounty>
,
<collectingMunicipality id="FA031302B243FFBC469B926DFAEBF9CB" box="[1179,1299,1590,1615]" pageId="7" pageNumber="258">Funakoshi</collectingMunicipality>
,
<location id="1F07DFA3B243FFBC471F926DFA60F9CB" LSID="urn:lsid:plazi:treatment:9271386EB243FFB4429791FCFE38FE4B:1F07DFA3B243FFBC471F926DFA60F9CB" box="[1311,1432,1590,1615]" county="Maehama Beach" latitude="34.2685" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" name="Daiou-cho" pageId="7" pageNumber="258" stateProvince="Mie">Daiou-cho</location>
,
<location id="1F07DFA3B243FFBC42979201FF1AF9F7" LSID="urn:lsid:plazi:treatment:9271386EB243FFB4429791FCFE38FE4B:1F07DFA3B243FFBC42979201FF1AF9F7" box="[151,226,1626,1651]" county="Maehama Beach" latitude="34.2685" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" name="Shima" pageId="7" pageNumber="258" stateProvince="Mie">Shima</location>
,
<collectingRegion id="D81C479AB243FFBC42ED9201FE6EF9F7" box="[237,406,1626,1651]" country="Japan" name="Mie" pageId="7" pageNumber="258">Mie Prefecture</collectingRegion>
,
<geoCoordinate id="7FECEFBFB243FFBC43A29201FDDFF9F7" box="[418,551,1626,1651]" direction="north" orientation="latitude" pageId="7" pageNumber="258" precision="9" value="34.2685">34°16.11N</geoCoordinate>
,
<geoCoordinate id="7FECEFBFB243FFBC40329201FD3BF9F7" box="[562,707,1626,1651]" direction="east" orientation="longitude" pageId="7" pageNumber="258" precision="9" value="136.8685">136°52.11E</geoCoordinate>
, intertidal, sand,
<date id="6E66AFB8B243FFBC41829200FBEEF9F7" box="[898,1046,1626,1651]" pageId="7" pageNumber="258" value="2013-07-23">
<collectingDate id="7E225650B243FFBC41829200FBEEF9F7" box="[898,1046,1626,1651]" pageId="7" pageNumber="258" value="2013-07-23">23 July 2013</collectingDate>
</date>
, coll.
<collectorName id="B72DECAEB243FFBC46599200FB2CF9F7" box="[1113,1236,1626,1651]" pageId="7" pageNumber="258">S. Kimura.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="1A678978B243FFBC42C79224FEB2F8CF" blockId="7.[151,1437,1590,2011]" pageId="7" pageNumber="258">
<materialsCitation id="AAB08325B243FFBC42C79224FC07F913" ID-GBIF-Occurrence="2234227824" box="[199,1023,1662,1687]" collectingDate="2013-07-23" collectionCode="ZRC" collectorName="S. Kimura." county="Maehama Beach" latitude="34.2685" location="Daiou-cho" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" pageId="7" pageNumber="258" specimenCode="ZRC 2018.0258" specimenCount="1" specimenCount-male="1" stateProvince="Mie" typeStatus="paratype">
<typeStatus id="C56337DAB243FFBC42C79224FED6F913" box="[199,302,1663,1687]" pageId="7" pageNumber="258" type="paratype">Paratype</typeStatus>
:
<specimenCode id="4A7E2103B243FFBC43399224FE0AF913" box="[313,498,1662,1687]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="258">ZRC 2018.0258</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC43FF9224FDB3F913" box="[511,587,1662,1687]" pageId="7" pageNumber="258" type="male">1 male</specimenCount>
(6.5×
<quantity id="DD20249DB243FFBC408F9225FD14F913" box="[655,748,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="258" unit="mm" value="5.6">5.6 mm</quantity>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="AAB08325B243FFBC460C9224FB83F93F" ID-GBIF-Occurrence="2234227997" collectingDate="2013-08-21" collectionCode="ZRC" collectorName="S. Kimura" county="Maehama Beach" latitude="34.2685" location="Daiou-cho" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" pageId="7" pageNumber="258" specimenCode="ZRC 2018.0259" specimenCount="1" specimenCount-female="1" stateProvince="Mie" typeStatus="paratype">
<specimenCode id="4A7E2103B243FFBC460C9224FB3DF913" box="[1036,1221,1662,1687]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="258">ZRC 2018.0259</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC46D29224FACCF913" box="[1234,1332,1662,1687]" pageId="7" pageNumber="258" type="female">1 female</specimenCount>
(8.1×
<quantity id="DD20249DB243FFBC47779224FF3FF93F" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="258" unit="mm" value="7.4">7.4 mm</quantity>
), 1 immature female (6.0×
<quantity id="DD20249DB243FFBC43F392F9FDB7F93F" box="[499,591,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="7" pageNumber="258" unit="mm" value="5.4">5.4 mm</quantity>
), same locality,
<date id="6E66AFB8B243FFBC410692F8FC45F93F" box="[774,957,1699,1723]" pageId="7" pageNumber="258" value="2013-08-21">
<collectingDate id="7E225650B243FFBC410692F8FC45F93F" box="[774,957,1699,1723]" pageId="7" pageNumber="258" value="2013-08-21">21 August 2013</collectingDate>
</date>
, coll.
<collectorName id="B72DECAEB243FFBC460392F8FB83F93F" box="[1027,1147,1698,1723]" pageId="7" pageNumber="258">S. Kimura</collectorName>
</materialsCitation>
;
<materialsCitation id="AAB08325B243FFBC468792F8FB4EF95B" ID-GBIF-Occurrence="2234228104" collectingDate="2017-08-26" collectionCode="ZRC" collectorName="M. Shimetsugu" county="Maehama Beach" latitude="34.2685" location="Daiou-cho" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" pageId="7" pageNumber="258" specimenCode="CBM-ZC 14486" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mie" typeStatus="paratype">
<specimenCode id="4A7E2103B243FFBC468792F8FABCF93F" box="[1159,1348,1698,1723]" collectionCode="CBM-ZC" pageId="7" pageNumber="258">CBM-ZC 14486</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC475092F8FA64F93F" box="[1360,1436,1698,1723]" pageId="7" pageNumber="258" type="male">1 male</specimenCount>
(7.3×
<quantity id="DD20249DB243FFBC42D2929DFED7F95B" box="[210,303,1734,1759]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="258" unit="mm" value="6.3">6.3 mm</quantity>
),
<specimenCount id="0CDE42F1B243FFBC4344929CFE50F95B" box="[324,424,1734,1759]" pageId="7" pageNumber="258" type="female">1 female</specimenCount>
(7.9×
<quantity id="DD20249DB243FFBC43EC929CFDB1F95B" box="[492,585,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="258" unit="mm" value="7.3">7.3 mm</quantity>
), same locality,
<date id="6E66AFB8B243FFBC4103929DFC46F95B" box="[771,958,1734,1759]" pageId="7" pageNumber="258" value="2017-08-26">
<collectingDate id="7E225650B243FFBC4103929DFC46F95B" box="[771,958,1734,1759]" pageId="7" pageNumber="258" value="2017-08-26">26 August 2017</collectingDate>
</date>
, coll.
<collectorName id="B72DECAEB243FFBC4605929CFB4EF95B" box="[1029,1206,1734,1759]" pageId="7" pageNumber="258">M. Shimetsugu</collectorName>
</materialsCitation>
;
<materialsCitation id="AAB08325B243FFBC46C3929CFB44F887" ID-GBIF-Occurrence="2234228016" collectingDate="2017-09-08" collectionCode="ZRC" collectorName="S. Kimura" county="Maehama Beach" latitude="34.2685" location="Daiou-cho" longLatPrecision="11" longitude="136.8685" municipality="Funakoshi" pageId="7" pageNumber="258" specimenCode="CBM-ZC 14487" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mie" typeStatus="paratype">
<specimenCode id="4A7E2103B243FFBC46C3929CFA79F95B" box="[1219,1409,1735,1759]" collectionCode="CBM-ZC" pageId="7" pageNumber="258">CBM-ZC 14487</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC478E929CFF35F887" pageId="7" pageNumber="258" type="male">1 male</specimenCount>
(7.0×
<quantity id="DD20249DB243FFBC431092B1FE93F887" box="[272,363,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="258" unit="mm" value="6.2">6.2 mm</quantity>
),
<specimenCount id="0CDE42F1B243FFBC437E92B0FE18F887" box="[382,480,1770,1795]" pageId="7" pageNumber="258" type="female">1 female</specimenCount>
(9.5×
<quantity id="DD20249DB243FFBC402292B1FD85F887" box="[546,637,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="258" unit="mm" value="8.8">8.8 mm</quantity>
), same locality,
<date id="6E66AFB8B243FFBC413292B0FBF8F887" box="[818,1024,1770,1795]" pageId="7" pageNumber="258" value="2017-09-08">
<collectingDate id="7E225650B243FFBC413292B0FBF8F887" box="[818,1024,1770,1795]" pageId="7" pageNumber="258" value="2017-09-08">8 September 2017</collectingDate>
</date>
, coll.
<collectorName id="B72DECAEB243FFBC464592B0FB44F887" box="[1093,1212,1770,1795]" pageId="7" pageNumber="258">S. Kimura</collectorName>
</materialsCitation>
;
<materialsCitation id="AAB08325B243FFBC46C792B0FEB2F8CF" ID-GBIF-Occurrence="2234228023" collectingDate="2018-06-01" collectionCode="CBM-ZC" collectorName="S. Kimura." latitude="33.6535" location="Shirahama" longLatPrecision="11" longitude="135.35817" municipality="Saino" pageId="7" pageNumber="258" specimenCode="CBM-ZC 15184" specimenCount="1" specimenCount-female="1" stateProvince="Wakayama" typeStatus="paratype">
<specimenCode id="4A7E2103B243FFBC46C792B0FA7BF887" box="[1223,1411,1770,1795]" collectionCode="CBM-ZC" pageId="7" pageNumber="258">CBM-ZC 15184</specimenCode>
,
<specimenCount id="0CDE42F1B243FFBC478E92B0FF1BF8A3" pageId="7" pageNumber="258" type="female">1 female</specimenCount>
(7.7×
<quantity id="DD20249DB243FFBC43259355FE78F8A3" box="[293,384,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="7" pageNumber="258" unit="mm" value="6.9">6.9 mm</quantity>
),
<collectingMunicipality id="FA031302B243FFBC43949355FE2EF8A3" box="[404,470,1806,1831]" pageId="7" pageNumber="258">Saino</collectingMunicipality>
,
<location id="1F07DFA3B243FFBC43E19355FDA4F8A3" LSID="urn:lsid:plazi:treatment:9271386EB243FFB4429791FCFE38FE4B:1F07DFA3B243FFBC43E19355FDA4F8A3" box="[481,604,1806,1831]" latitude="33.6535" longLatPrecision="11" longitude="135.35817" municipality="Saino" name="Shirahama" pageId="7" pageNumber="258" stateProvince="Wakayama">Shirahama</location>
,
<collectingRegion id="D81C479AB243FFBC40679355FC98F8A3" box="[615,864,1806,1831]" country="Japan" name="Wakayama" pageId="7" pageNumber="258">Wakayama Prefecture</collectingRegion>
,
<geoCoordinate id="7FECEFBFB243FFBC416B9354FC0AF8A3" box="[875,1010,1807,1831]" direction="north" orientation="latitude" pageId="7" pageNumber="258" precision="9" value="33.6535">33°39.21N</geoCoordinate>
,
<geoCoordinate id="7FECEFBFB243FFBC41FD9355FB77F8A3" box="[1021,1167,1806,1831]" direction="east" orientation="longitude" pageId="7" pageNumber="258" precision="9" value="135.35817">135°21.49E</geoCoordinate>
, intertidal,
<date id="6E66AFB8B243FFBC470D9354FA60F8A3" box="[1293,1432,1807,1831]" pageId="7" pageNumber="258" value="2018-06-01">
<collectingDate id="7E225650B243FFBC470D9354FA60F8A3" box="[1293,1432,1807,1831]" pageId="7" pageNumber="258" value="2018-06-01">1 June 2018</collectingDate>
</date>
, coll.
<collectorName id="B72DECAEB243FFBC42CF9368FEB2F8CF" box="[207,330,1842,1867]" pageId="7" pageNumber="258">S. Kimura.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B243FFB542C7930DFC39FC91" lastPageId="14" lastPageNumber="265" pageId="7" pageNumber="258" type="description">
<paragraph id="1A678978B243FFB342C7930DFA9DFEC4" blockId="7.[151,1437,1590,2011]" lastBlockId="8.[151,1437,151,789]" lastPageId="8" lastPageNumber="259" pageId="7" pageNumber="258">
<emphasis id="28AC556AB243FFBC42C7930DFEA1F8EB" bold="true" box="[199,345,1878,1903]" pageId="7" pageNumber="258">Description.</emphasis>
<emphasis id="28AC556AB243FFBC4363930DFDEAF8EA" box="[355,530,1878,1902]" italics="true" pageId="7" pageNumber="258">
<typeStatus id="C56337DAB243FFBC4363930DFE32F8EA" box="[355,458,1878,1902]" pageId="7" pageNumber="258" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Carapace (
<figureCitation id="82E395FDB243FFBC4098930DFD13F8EB" box="[664,747,1878,1903]" captionStart="FIGURE 5" captionStartId="8.[151,250,1941,1963]" captionTargetBox="[151,1435,828,1919]" captionTargetId="figure@8.[151,1435,828,1919]" captionTargetPageId="8" captionText="FIGURE 5. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625341/files/figure.png" pageId="7" pageNumber="258">Figs. 5</figureCitation>
,
<figureCitation id="82E395FDB243FFBC40FA930DFCE6F8EB" box="[762,798,1878,1903]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="7" pageNumber="258">6A</figureCitation>
) subpyriform in general outline, 1.15 times as long as wide; dorsal surface convex, glabrous, coarsely punctate. Front (
<figureCitation id="82E395FDB243FFBC418B9321FC1DF817" box="[907,997,1914,1939]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="7" pageNumber="258">Fig. 6B</figureCitation>
) weakly produced anteriorly, anterior margin trilobate, median lobe subsemicircular, projecting anteriorly, distinctly exceeding lateral teeth; lateral lobe (inner orbital lobe) blunt, not particularly produced. Orbits small; supraorbital margin strongly oblique, with short, blunt lateral tooth demarcated mesially by deep, narrow longitudinal suture; bilobed anterior margin of efferent branchial channel forming lower orbital margin, inner angle acutely pointed. Hepatic region slightly inflated, forming low, dome-like elevation. Mesogastric region with obsolescent median ridge, shallowly depressed on both sides of ridge. Lateral to posterior margins keel-like, bordered with relatively coarse granules (
<figureCitation id="82E395FDB24CFFB346C9955FFADBFE99" box="[1225,1315,260,285]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="8" pageNumber="259">Fig. 6C</figureCitation>
), with low convexity just below hepatic region (pterygostomial margin); posterior margin bi-keeled, both nearly straight.
</paragraph>
<paragraph id="1A678978B24CFFB342C79517FAE6FE29" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="259">
Eyestalks (
<figureCitation id="82E395FDB24CFFB343499517FE5FFEE1" box="[329,423,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="8" pageNumber="259">Fig. 6B</figureCitation>
) retractable, subcylindrical, mesial side protruding into minute blunt distal tubercle exceeding cornea. Antennular fossa continuous with orbit, partially sealed by basal plate on antennule; antennules folded slightly obliquely within fossa. Antenna very small, inserted between antennular fossa and orbit.
</paragraph>
<paragraph id="1A678978B24CFFB342C795ECFB51FD01" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="259">
Thoracic sternum glabrous. Sternite 3 (
<figureCitation id="82E395FDB24CFFB3409D95ECFD03FE54" box="[669,763,439,464]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="8" pageNumber="259">Fig. 6D</figureCitation>
) subtriangular, anterior margin produced anteriorly with subacute median point, surface slightly depressed medially. Sternite 4 (
<figureCitation id="82E395FDB24CFFB341AD9587FBFBFE71" box="[941,1027,476,501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="8" pageNumber="259">Fig. 6D</figureCitation>
) largest, anterior and lateral margins coarsely granular, otherwise with sparse granules and few punctae. Sternites 58 becoming smaller toward posterior, surfaces with few punctae. Episternite 4 clearly demarcated from sternite 4; episternites 58 each with shallowly concave outer margin. Sternopleonal cavity deep, extending onto posterior part of thoracic sternite 3, margins on sternite 4, corresponding to pleomere 6 and telson, bordered with coarse granules
</paragraph>
<paragraph id="1A678978B24CFFB342C796D4FE57FC91" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="259">
Maxilliped 3 (
<figureCitation id="82E395FDB24CFFB3436E96D4FE3EFD2C" box="[366,454,655,680]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="8" pageNumber="259">Fig. 6E</figureCitation>
) almost flat, outer surface with scattered punctae. Ischium subrectangular, distinctly longer than wide; merus triangular, tapering distally to subacute tip, subequal in length to ischium measured along mesial margin; lateral margin of ischium and merus minutely beaded. Exopod with distal article spatuliform, lateral margin minutely beaded.
</paragraph>
<caption id="4EA7D9F0B24CFFB3429793CEFAABF84E" httpUri="https://zenodo.org/record/2625341/files/figure.png" pageId="8" pageNumber="259" startId="8.[151,250,1941,1963]" targetBox="[151,1435,828,1919]" targetPageId="8">
<paragraph id="1A678978B24CFFB3429793CEFAABF84E" blockId="8.[151,1436,1941,1994]" pageId="8" pageNumber="259">
<emphasis id="28AC556AB24CFFB3429793CEFEF7F82E" bold="true" box="[151,271,1941,1963]" pageId="8" pageNumber="259">FIGURE 5</emphasis>
.
<taxonomicName id="DDD8F2FBB24CFFB3431D93CEFE04F82F" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[285,508,1941,1963]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24CFFB3431D93CEFE04F82F" box="[285,508,1941,1963]" italics="true" pageId="8" pageNumber="259">Pseudophilyra parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24CFFB3400493CEFDB8F82E" bold="true" box="[516,576,1941,1962]" pageId="8" pageNumber="259">
<taxonomicNameLabel id="339FE811B24CFFB3400493CEFDB8F82E" box="[516,576,1941,1962]" pageId="8" pageNumber="259" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace.
</paragraph>
</caption>
<caption id="4EA7D9F0B24DFFB24297930EFBCAF861" httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="9" pageNumber="260" startId="9.[151,250,1877,1899]" targetBox="[214,1360,231,1827]" targetPageId="9">
<paragraph id="1A678978B24DFFB24297930EFBCAF861" blockId="9.[151,1436,1876,2021]" pageId="9" pageNumber="260">
<emphasis id="28AC556AB24DFFB24297930EFEF5F8EE" bold="true" box="[151,269,1877,1899]" pageId="9" pageNumber="260">FIGURE 6</emphasis>
.
<taxonomicName id="DDD8F2FBB24DFFB2431A930FFE0FF8EE" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[282,503,1876,1898]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24DFFB2431A930FFE0FF8EE" box="[282,503,1876,1898]" italics="true" pageId="9" pageNumber="260">Pseudophilyra parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24DFFB243FE930EFDC0F8EE" bold="true" box="[510,568,1877,1898]" pageId="9" pageNumber="260">
<taxonomicNameLabel id="339FE811B24DFFB243FE930EFDC0F8EE" box="[510,568,1877,1898]" pageId="9" pageNumber="260" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view.
</paragraph>
</caption>
<caption id="4EA7D9F0B24EFFB14297927CFD3BF9FE" httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="10" pageNumber="261" startId="10.[151,250,1575,1597]" targetBox="[188,1389,280,1509]" targetPageId="10">
<paragraph id="1A678978B24EFFB14297927CFD3BF9FE" blockId="10.[151,1436,1575,1658]" pageId="10" pageNumber="261">
<emphasis id="28AC556AB24EFFB14297927CFEF4F9B8" bold="true" box="[151,268,1575,1597]" pageId="10" pageNumber="261">FIGURE 7</emphasis>
.
<taxonomicName id="DDD8F2FBB24EFFB14318927CFE0BF9B9" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[280,499,1575,1597]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24EFFB14318927CFE0BF9B9" box="[280,499,1575,1597]" italics="true" pageId="10" pageNumber="261">Pseudophilyra parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24EFFB143F9927CFDCAF9B8" bold="true" box="[505,562,1575,1596]" pageId="10" pageNumber="261">
<taxonomicNameLabel id="339FE811B24EFFB143F9927CFDCAF9B8" box="[505,562,1575,1596]" pageId="10" pageNumber="261" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off).
</paragraph>
</caption>
<paragraph id="1A678978B24EFFB142C792FAFE9EF85E" blockId="10.[151,1437,1697,2010]" pageId="10" pageNumber="261">
Chelipeds (
<figureCitation id="82E395FDB24EFFB1434892FAFE61F93E" box="[328,409,1697,1722]" captionStart="FIGURE 5" captionStartId="8.[151,250,1941,1963]" captionTargetBox="[151,1435,828,1919]" captionTargetId="figure@8.[151,1435,828,1919]" captionTargetPageId="8" captionText="FIGURE 5. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625341/files/figure.png" pageId="10" pageNumber="261">Figs. 5</figureCitation>
,
<figureCitation id="82E395FDB24EFFB143A692FAFE1FF93E" box="[422,487,1697,1722]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="10" pageNumber="261">6F, G</figureCitation>
) symmetrical, 1.5 times as long as carapace, glabrous. Merus subcylindrical, about 3.0 times as long as wide, covered with pearly granules of various sizes except for distal one-third of upper surface and distal two-thirds of lower surfaces where granules almost absent carpus. Carpus short, cup-shaped, inner surface concave; upper inner margin with row of small granules, but otherwise almost smooth. Palm compressed, 1.9 times as long as wide, slightly sinuous on outer (extensor) margin, inner (flexor) margin also slightly sinuous; upper surface almost smooth, lower surface with scattered punctae; fixed finger weakly deflexed, nearly straight, terminating in slightly curved, blunt tip; dactylus 0.7 times as long as palm, terminating in slightly curved, blunt tip; both occlusal margins meeting in distal half and forming oval hiatus in proximal half, each without conspicuous teeth.
</paragraph>
<caption id="4EA7D9F0B24FFFB0429793F5FC55F866" httpUri="https://zenodo.org/record/2625349/files/figure.png" pageId="11" pageNumber="262" startId="11.[151,250,1966,1988]" targetBox="[207,1373,193,1942]" targetPageId="11">
<paragraph id="1A678978B24FFFB0429793F5FC55F866" blockId="11.[151,1435,1966,2018]" pageId="11" pageNumber="262">
<emphasis id="28AC556AB24FFFB0429793F5FEF4F847" bold="true" box="[151,268,1966,1988]" pageId="11" pageNumber="262">FIGURE 8</emphasis>
.
<taxonomicName id="DDD8F2FBB24FFFB0431993F5FE0DF840" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[281,501,1966,1988]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="262" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24FFFB0431993F5FE0DF840" box="[281,501,1966,1988]" italics="true" pageId="11" pageNumber="262">Pseudophilyra parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24FFFB043FB93F5FDCCF847" bold="true" box="[507,564,1966,1987]" pageId="11" pageNumber="262">
<taxonomicNameLabel id="339FE811B24FFFB043FB93F5FDCCF847" box="[507,564,1966,1987]" pageId="11" pageNumber="262" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, CBM-ZC 14486, paratypes, male (7.3×6.3 mm) (A, B), female (7.9×7.3 mm) (C, D), showing colour in life. A, C, habitus, dorsal view; B, D, habitus, ventral view.
</paragraph>
</caption>
<paragraph id="1A678978B248FFB742C794CCFE84FE99" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="263">
Ambulatory legs (
<figureCitation id="82E395FDB248FFB7439494CCFE1CFF34" box="[404,484,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1941,1963]" captionTargetBox="[151,1435,828,1919]" captionTargetId="figure@8.[151,1435,828,1919]" captionTargetPageId="8" captionText="FIGURE 5. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625341/files/figure.png" pageId="12" pageNumber="263">Figs. 5</figureCitation>
,
<figureCitation id="82E395FDB248FFB743F194CCFDEDFF34" box="[497,533,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="12" pageNumber="263">6H</figureCitation>
) slender, glabrous, similar in shape, gradually decreasing in length from first to fourth; each merus cylindrical; each carpus as long as or shorter than propodus; propodus somewhat compressed; each dactylus flattened, slightly curved, tapering distally to minute corneous claw, subequal in length to carpus and propodus combined.
</paragraph>
<paragraph id="1A678978B248FFB742C7957CFA7BFE54" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="263">
Pleon (
<figureCitation id="82E395FDB248FFB74319957CFE91FEC4" box="[281,361,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="12" pageNumber="263">
Fig.
<date id="6E66AFB8B248FFB74351957CFE91FEC4" box="[337,361,295,320]" pageId="12" pageNumber="263">6I</date>
</figureCitation>
) elongate-triangular, gently curved sternally. Somites 1 and 2 very short, transversely linear. Main fused section consisting of somites 36 elongate trapezoidal, outer surface with scattered punctae; trace of suture between somites 4 and 5 still discernible; proximolateral margin of fused somite 3 and 4 slightly expanded, lateral margin with low convexity medially. Somite 6 with trace of median tubercle somewhat proximal to distal margin; lateral margin with low convexity at proximal 0.2. Telson elongate-triangular, 1.9 times as long as wide.
</paragraph>
<paragraph id="1A678978B248FFB742C79587FF0BFD01" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="263">
Gonopod 1 (
<figureCitation id="82E395FDB248FFB7435B9587FE2BFE70" box="[347,467,476,501]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="12" pageNumber="263">Fig. 6J, K</figureCitation>
) slender, slightly sinuously curved (distal half nearly straight except for outwardly curved distal part), gradually tapering, reaching nearly to suture between thoracic sternites 3/4, deeply bifurcate at distal 0.3; branches subequal in length, each terminating in acute tip. Gonopod 2 (
<figureCitation id="82E395FDB248FFB74655967FFB55FDB9" box="[1109,1197,548,573]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="12" pageNumber="263">Fig. 6L</figureCitation>
) about 0.3 length of gonopod 1, gently curved, proximal part slightly expanded; distal part unequally bilobed, terminal lobe bluntly pointed.
</paragraph>
<paragraph id="1A678978B248FFB742C796CAFE42FD48" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="263">
<emphasis id="28AC556AB248FFB742C796CAFE8DFD2C" box="[199,373,656,680]" italics="true" pageId="12" pageNumber="263">
<typeStatus id="C56337DAB248FFB742C796CAFED6FD2C" box="[199,302,657,680]" pageId="12" pageNumber="263" type="paratype">Paratype</typeStatus>
males
</emphasis>
. Agree well with
<typeStatus id="C56337DAB248FFB7403F96D4FD59FD2C" box="[575,673,655,680]" pageId="12" pageNumber="263" type="holotype">holotype</typeStatus>
male. Carapace 1.131.15 times as long as wide. Chelipeds 1.41.5 times as long as carapace.
</paragraph>
<caption id="4EA7D9F0B248FFB7429793EDFB69F86F" httpUri="https://zenodo.org/record/2625351/files/figure.png" pageId="12" pageNumber="263" startId="12.[151,250,1974,1996]" targetBox="[151,1435,769,1953]" targetPageId="12">
<paragraph id="1A678978B248FFB7429793EDFB69F86F" blockId="12.[151,1436,1974,2027]" pageId="12" pageNumber="263">
<emphasis id="28AC556AB248FFB7429793EDFEF7F848" bold="true" box="[151,271,1974,1996]" pageId="12" pageNumber="263">FIGURE 9</emphasis>
.
<taxonomicName id="DDD8F2FBB248FFB7431E93EDFD50F848" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[286,680,1974,1996]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="263" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB248FFB7431E93EDFDE7F848" box="[286,543,1974,1996]" italics="true" pageId="12" pageNumber="263">Pseudophilyra tridentata</emphasis>
Miers, 1879
</taxonomicName>
, CBM-ZC 7846, male (6.1×5.3 mm), Okinawa Islands, Ryukyu Islands, Japan, carapace in dorsal view, stained with methylene blue to show details of dorsal ornamentation.
</paragraph>
</caption>
<caption id="4EA7D9F0B249FFB64297932BFC8BF866" httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="13" pageNumber="264" startId="13.[151,250,1904,1926]" targetBox="[229,1373,222,1867]" targetPageId="13">
<paragraph id="1A678978B249FFB64297932BFC8BF866" blockId="13.[151,1436,1904,2018]" pageId="13" pageNumber="264">
<emphasis id="28AC556AB249FFB64297932BFEE0F801" bold="true" box="[151,280,1904,1926]" pageId="13" pageNumber="264">FIGURE 10</emphasis>
.
<taxonomicName id="DDD8F2FBB249FFB64325932BFD5EF802" authority="Miers, 1879" authorityName="Miers" authorityYear="1879" box="[293,678,1904,1926]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="264" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB249FFB64325932BFDDBF802" box="[293,547,1904,1926]" italics="true" pageId="13" pageNumber="264">Pseudophilyra tridentata</emphasis>
Miers, 1879
</taxonomicName>
, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view.
</paragraph>
</caption>
<paragraph id="1A678978B24AFFB542C794C3FF18FF51" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
<emphasis id="28AC556AB24AFFB542C794C3FEE3FF34" box="[199,283,152,176]" italics="true" pageId="14" pageNumber="265">Female</emphasis>
. Carapace (
<figureCitation id="82E395FDB24AFFB543A594CCFE06FF34" box="[421,510,151,176]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="14" pageNumber="265">Fig. 7A</figureCitation>
) 1.081.10 times as long as wide. Front relatively slightly less produced than in males.
</paragraph>
<paragraph id="1A678978B24AFFB542C79484FD72FE54" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
Exposed lateral part of thoracic sternum very narrow (
<figureCitation id="82E395FDB24AFFB5415C9484FC44FF7C" box="[860,956,223,248]" captionStart="FIGURE 8" captionStartId="11.[151,250,1966,1988]" captionTargetBox="[207,1373,193,1942]" captionTargetId="figure@11.[207,1373,193,1942]" captionTargetPageId="11" captionText="FIGURE 8. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratypes, male (7.3×6.3 mm) (A, B), female (7.9×7.3 mm) (C, D), showing colour in life. A, C, habitus, dorsal view; B, D, habitus, ventral view." httpUri="https://zenodo.org/record/2625349/files/figure.png" pageId="14" pageNumber="265">Fig. 8D</figureCitation>
), smooth, forming outer wall of deeply excavated sternopleonal cavity; sutures between sternites 4/5, 5/6, 6/7 and 7/8 distinct; sternites 13 fused, deeply depressed below to accommodate telson. Sternopleonal cavity (
<figureCitation id="82E395FDB24AFFB5417E957CFC20FEC4" box="[894,984,295,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="14" pageNumber="265">Fig. 7B</figureCitation>
) circular in outline, margins distinctly delimited; sutures between sternites 4/5, 5/6, 6/7, and 7/8 widely interrupted medially; median suture absent; sternite 4 large, occupying anterior half of sternopleonal concavity, with scattered short, occasionally feathered, setae. Vulvae (
<figureCitation id="82E395FDB24AFFB5434595CFFE3DFE28" box="[325,453,404,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="14" pageNumber="265">Fig. 7B, C</figureCitation>
) small, located just posterior to mesial end of suture between sternites 5/6, outline circular, without protrusion on outer margin.
</paragraph>
<paragraph id="1A678978B24AFFB542C79587FAEFFD9C" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
Cheliped slightly slender than that of males, 1.3 times as long as carapace; palm 2.2 times as long as wide; occlusal margins of fingers of cheliped without conspicuous teeth proximal to meeting point (
<figureCitation id="82E395FDB24AFFB546B095A4FAF2FD9C" box="[1200,1290,511,536]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="14" pageNumber="265">Fig. 7D</figureCitation>
).
</paragraph>
<paragraph id="1A678978B24AFFB542C7967FFB76FD00" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
Pleon (
<figureCitation id="82E395FDB24AFFB54315967FFE92FDB8" box="[277,362,548,573]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="14" pageNumber="265">Fig. 7E</figureCitation>
) broad, dome-like; somites 1 and 2 short, transversely band-shaped, somite 2 about twice longer than somite 1; main fused section (somites 36) ovoid, no trace of sutures. Telson (damaged in illustrated specimen) small, subsemicircular, exceeding beyond anterior margin of thoracic sternite 3.
</paragraph>
<paragraph id="1A678978B24AFFB542C796D4FC39FC91" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
<emphasis id="28AC556AB24AFFB542C796D4FE94FD2C" bold="true" box="[199,364,655,680]" pageId="14" pageNumber="265">Colour in life.</emphasis>
See
<figureCitation id="82E395FDB24AFFB543A296D4FE1DFD2C" box="[418,485,655,680]" captionStart="FIGURE 8" captionStartId="11.[151,250,1966,1988]" captionTargetBox="[207,1373,193,1942]" captionTargetId="figure@11.[207,1373,193,1942]" captionTargetPageId="11" captionText="FIGURE 8. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratypes, male (7.3×6.3 mm) (A, B), female (7.9×7.3 mm) (C, D), showing colour in life. A, C, habitus, dorsal view; B, D, habitus, ventral view." httpUri="https://zenodo.org/record/2625349/files/figure.png" pageId="14" pageNumber="265">Fig. 8</figureCitation>
. Carapace generally gray or gray-brown, sometimes with irregular blotches of white. Thoracic sternum and pleon entirely white. Cheliped with large gray or gray-brown patches on dorsal surface of merus and palm; fingers with tint of light brown on dorsal surface; ventral surface overall white. Ambulatory legs generally white, with spots of gray or brown on meri, carpi and propodi.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B24AFFB542C79744FD46FCD9" pageId="14" pageNumber="265" type="distribution">
<paragraph id="1A678978B24AFFB542C79744FD46FCD9" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
<emphasis id="28AC556AB24AFFB542C79744FE98FCBC" bold="true" box="[199,352,799,824]" pageId="14" pageNumber="265">Distribution.</emphasis>
Presently known only from the
<typeStatus id="C56337DAB24AFFB540E1977BFCE9FCBC" box="[737,785,800,824]" pageId="14" pageNumber="265">type</typeStatus>
locality, Funakoshi, Daiou-cho, Shima,
<collectingRegion id="D81C479AB24AFFB546EA9744FA60FCBC" box="[1258,1432,799,824]" country="Japan" name="Mie" pageId="14" pageNumber="265">Mie Prefecture</collectingRegion>
,
<collectingCountry id="62CFC9E8B24AFFB54297971EFF22FCD8" box="[151,218,837,860]" name="Japan" pageId="14" pageNumber="265">Japan</collectingCountry>
; intertidal to shallow subtidal sand bottom.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B24AFFB542C7973CFD2EFBF8" pageId="14" pageNumber="265" type="discussion">
<paragraph id="1A678978B24AFFB542C7973CFD2EFBF8" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="265">
<emphasis id="28AC556AB24AFFB542C7973CFEC3FC04" bold="true" box="[199,315,871,896]" pageId="14" pageNumber="265">Remarks.</emphasis>
<taxonomicName id="DDD8F2FBB24AFFB543419733FDCCFC04" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[321,564,872,896]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB543419733FDCCFC04" box="[321,564,872,896]" italics="true" pageId="14" pageNumber="265">Pseudophilyra parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5403B973CFD82FC04" bold="true" box="[571,634,871,896]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5403B973CFD82FC04" box="[571,634,871,896]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
closely resembles
<taxonomicName id="DDD8F2FBB24AFFB5414D9732FC18FC04" authorityName="Chen &amp; Ng" authorityYear="2003" box="[845,992,872,896]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB5414D9732FC18FC04" box="[845,992,872,896]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
in the general shape and ornamentation of the carapace, the structure of the cheliped and the deeply bifurcate distal part of the male gonopod 1. The new species differs from
<taxonomicName id="DDD8F2FBB24AFFB5438297EAFDE0FC4C" authorityName="Chen &amp; Ng" authorityYear="2003" box="[386,536,944,968]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB5438297EAFDE0FC4C" box="[386,536,944,968]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
in the coarser granules bordering the carapace lateral margins (
<figureCitation id="82E395FDB24AFFB546F497F4FAB3FC4C" box="[1268,1355,943,968]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6C</figureCitation>
versus
<figureCitation id="82E395FDB24AFFB54297978FFF14FC68" box="[151,236,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3C</figureCitation>
) and the less curved distal part of the male gonopod 1 (
<figureCitation id="82E395FDB24AFFB54168978FFC21FC68" box="[872,985,980,1005]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6J, K</figureCitation>
versus
<figureCitation id="82E395FDB24AFFB54631978FFB61FC68" box="[1073,1177,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">
Fig.
<date id="6E66AFB8B24AFFB54665978FFB87FC68" box="[1125,1151,980,1004]" pageId="14" pageNumber="265">3I</date>
, J
</figureCitation>
). The other potentially useful characters are: in males, the sternite 4 is more strongly granular in
<taxonomicName id="DDD8F2FBB24AFFB541C997A2FBD5FB94" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[969,1069,1016,1040]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB541C997A2FBD5FB94" box="[969,1069,1016,1040]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5463497ACFB8BFB94" bold="true" box="[1076,1139,1015,1040]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5463497ACFB8BFB94" box="[1076,1139,1015,1040]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="DDD8F2FBB24AFFB546CC97A2FA98FB94" authorityName="Chen &amp; Ng" authorityYear="2003" box="[1228,1376,1016,1040]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB546CC97A2FA98FB94" box="[1228,1376,1016,1040]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5476E97ACFF41FBB1" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6D</figureCitation>
versus
<figureCitation id="82E395FDB24AFFB5430E9047FE9FFBB0" box="[270,359,1052,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3D</figureCitation>
); the surface of the fused part of the pleon is more strongly punctate in
<taxonomicName id="DDD8F2FBB24AFFB546839046FB1FFBB0" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[1155,1255,1052,1076]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB546839046FB1FFBB0" box="[1155,1255,1052,1076]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB546EE9047FAD4FBB1" bold="true" box="[1262,1324,1052,1077]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB546EE9047FAD4FBB1" box="[1262,1324,1052,1077]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in
<taxonomicName id="DDD8F2FBB24AFFB547879046FEF7FBDC" authorityName="Chen &amp; Ng" authorityYear="2003" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB547879046FEF7FBDC" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5431F9064FE93FBDC" box="[287,363,1087,1112]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">
Fig.
<date id="6E66AFB8B24AFFB543549064FE93FBDC" box="[340,363,1087,1112]" pageId="14" pageNumber="265">6I</date>
</figureCitation>
versus
<figureCitation id="82E395FDB24AFFB543C39064FDE5FBDC" box="[451,541,1087,1112]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3H</figureCitation>
); and the outer margin of the female vulva is straight in
<taxonomicName id="DDD8F2FBB24AFFB5469E901AFAFCFBDC" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[1182,1284,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB5469E901AFAFCFBDC" box="[1182,1284,1088,1112]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5470C9064FAB3FBDC" bold="true" box="[1292,1355,1087,1112]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5470C9064FAB3FBDC" box="[1292,1355,1087,1112]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, rather than forming a rounded protrusion in
<taxonomicName id="DDD8F2FBB24AFFB5403C903EFD28FBF8" authorityName="Chen &amp; Ng" authorityYear="2003" box="[572,720,1124,1148]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB5403C903EFD28FBF8" box="[572,720,1124,1148]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B24AFFB442C790D3FDE3FE0C" lastPageId="15" lastPageNumber="266" pageId="14" pageNumber="265" type="reference_group">
<paragraph id="1A678978B24AFFB442C790D3FAF8FF7C" blockId="14.[151,1437,151,2013]" lastBlockId="15.[151,1437,151,464]" lastPageId="15" lastPageNumber="266" pageId="14" pageNumber="265">
<taxonomicName id="DDD8F2FBB24AFFB542C790D3FE18FB24" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[199,480,1160,1184]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB542C790D3FE18FB24" box="[199,480,1160,1184]" italics="true" pageId="14" pageNumber="265">Pseudophilyra tridentata</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="62CFC9E8B24AFFB5414090D2FC78FB24" box="[832,896,1161,1184]" name="Japan" pageId="14" pageNumber="265">Japan</collectingCountry>
on the basis of a female
<typeStatus id="C56337DAB24AFFB5469A90DCFB07FB24" box="[1178,1279,1159,1184]" pageId="14" pageNumber="265" type="holotype">holotype</typeStatus>
, and has been recorded from various Indo-West Pacific localities (
<bibRefCitation id="7E49F489B24AFFB540D890F7FC95FB41" author="Alcock, A." box="[728,877,1196,1221]" pageId="14" pageNumber="265" pagination="134 - 296" refId="ref5841" refString="Alcock, A. (1896) Materials for carcinological fauna of India. No. 2. The Brachyura Oxystomata. Journal of the Asiatic Society of Bengal, 65, 134 - 296, pls. 6 - 8." type="journal article" year="1896">Alcock 1896</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB5417890F7FBEAFB40" author="Calman, W. T." box="[888,1042,1196,1221]" pageId="14" pageNumber="265" pagination="1 - 50" refId="ref6029" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society, Series 2, Zoology, 3, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman 1900</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB5461D90F7FB50FB41" author="Laurie, D." box="[1053,1192,1196,1221]" pageId="14" pageNumber="265" pagination="1" refId="ref6553" refString="Laurie, D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. Report of Ceylon Pearl Oyster Fisheries, Part V (Supplementary Reports 40), 349 - 432, pls. 1, 2." type="journal article" year="1906">Laurie 1906</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB546B390F7FAACFB40" author="Rathbun, M. J." box="[1203,1364,1196,1221]" pageId="14" pageNumber="265" pagination="301 - 367" refId="ref7059" refString="Rathbun, M. J. (1910) The Danish Expedition to Siam 1899 - 1900. V. Brachyura. Det Kongelige Danske Uidenskabernes Selskabs Skrifter, Series 7, 4, 301 - 367, pls. 1, 2." type="book chapter" year="1910">Rathbun 1910</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB5476090F7FF2AFB6C" author="Balss, H." pageId="14" pageNumber="265" pagination="1 - 20" refId="ref5927" refString="Balss, H. (1916) Expeditionen S. M. Schiff &quot; Pola &quot; in das Rote Meer. Nordliche und Sudliche Halfte 1895 / 96 - 1897 / 98. Zoologische Ergebnisse. XXXI. Die Decapoden des Roten Meeres. II. Anomuren, Dromiaceen und Oxystomen. Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 92, 1 - 20." type="journal article" year="1916">Balss 1916</bibRefCitation>
,
<bibRefCitation id="7E49F489B24AFFB542DC908BFEEFFB6C" author="Balss, H." box="[220,279,1232,1256]" pageId="14" pageNumber="265" pagination="104 - 140" refId="ref5997" refString="Balss, H. (1922) Ostasiatische Decapoden. Ill. Die Dromiaceen, Oxystomen und Parthenopiden. Archiv fur Naturgeschichte, Series A, 88, 104 - 140." type="journal article" year="1922">1922</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB543229094FE1CFB6C" author="Stephensen, K." box="[290,484,1231,1256]" pageId="14" pageNumber="265" pagination="57 - 237" refId="ref7298" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, 4, 57 - 237." type="journal article" year="1946">Stephensen 1946</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB543EE9094FCACFB6C" author="Tyndale-Biscoe, M. &amp; George, R. W." box="[494,852,1231,1256]" pageId="14" pageNumber="265" pagination="65 - 96" refId="ref7362" refString="Tyndale-Biscoe, M. &amp; George, R. W. (1962) The Oxystomata and Gymnopleura (Crustacea, Brachyura) of Western Australia with descriptions of two new species from Western Australia and one from India. Journal of the Royal Society of Western Australia, 45, 65 - 96." type="journal article" year="1962">Tyndale-Biscoe &amp; George 1962</bibRefCitation>
;
<bibRefCitation id="7E49F489B24AFFB5415F9094FB49FB6C" author="Campbell, B. M. &amp; Stephenson, W." box="[863,1201,1231,1256]" pageId="14" pageNumber="265" pagination="235 - 301" refId="ref6089" refString="Campbell, B. M. &amp; Stephenson, W. (1970) The sublittoral Brachyura (Crustacea: Decapoda) of Moreton Bay. Memoirs of the Queensland Museum, 15, 235 - 301, pl. 22." type="journal article" year="1970">Campbell &amp; Stephenson 1970</bibRefCitation>
).
<bibRefCitation id="7E49F489B24AFFB546C4908AFF19FA88" author="Komatsu, H. &amp; Takeda, M." pageId="14" pageNumber="265" pagination="267 - 283" refId="ref6441" refString="Komatsu, H. &amp; Takeda, M. (2000) Leucosiid crabs (Crustacea: Decapoda: Brachyura) from the Osumi Islands, southwest Japan, with description of a new species of Cryptocnemus. Species Diversity, 5, 267 - 283. https: // doi. org / 10.12782 / specdiv. 5.267" type="journal article" year="2000">Komatsu &amp; Takeda (2000)</bibRefCitation>
presented a redescription of the
<typeStatus id="C56337DAB24AFFB5404C90AFFD49FA89" box="[588,689,1268,1293]" pageId="14" pageNumber="265" type="holotype">holotype</typeStatus>
, and clarified that previous records of
<taxonomicName id="DDD8F2FBB24AFFB5465C90AEFB1DFA88" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[1116,1253,1268,1292]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5465C90AEFB1DFA88" box="[1116,1253,1268,1292]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
by
<bibRefCitation id="7E49F489B24AFFB5470E90AFFA60FA88" author="Sakai, T." box="[1294,1432,1268,1293]" pageId="14" pageNumber="265" pagination="67 - 192" refId="ref7103" refString="Sakai, T. (1937) Studies on the crabs of Japan. 11. Oxystomata. Science Reports of the Tokyo Bunrika Daigaku, (B) 3, Supplement, 67 - 192, pls. 10 - 19." type="book chapter" year="1937">Sakai (1937</bibRefCitation>
,
<bibRefCitation id="7E49F489B24AFFB54297914CFF28FAB4" author="Sakai, T." box="[151,208,1303,1328]" pageId="14" pageNumber="265" refId="ref7146" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, xxix + 773 pp., 3 maps (English volume); 461 pp. (Japanese volume); 16 pp., 251 pls. (pl. volume)." type="book" year="1976">1976</bibRefCitation>
) actually represent
<taxonomicName id="DDD8F2FBB24AFFB543B49142FDB3FAB4" ID-CoL="4P79V" authorityName="Ihle" authorityYear="1918" box="[436,587,1304,1328]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="28AC556AB24AFFB543B49142FDB3FAB4" box="[436,587,1304,1328]" italics="true" pageId="14" pageNumber="265">P. intermedia</emphasis>
</taxonomicName>
instead. It is likely that those records outside
<collectingCountry id="62CFC9E8B24AFFB546639142FB5BFAB4" box="[1123,1187,1305,1328]" name="Japan" pageId="14" pageNumber="265">Japan</collectingCountry>
contain species other than
<taxonomicName id="DDD8F2FBB24AFFB542CD9166FEAFFAD0" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[205,343,1340,1364]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB542CD9166FEAFFAD0" box="[205,343,1340,1364]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
. For example, the male first gonopods illustrated by
<bibRefCitation id="7E49F489B24AFFB541A49167FB8DFAD1" author="Stephensen, K." box="[932,1141,1340,1365]" pageId="14" pageNumber="265" pagination="57 - 237" refId="ref7298" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, 4, 57 - 237." type="journal article" year="1946">Stephensen (1946)</bibRefCitation>
from the Persian Gulf and
<bibRefCitation id="7E49F489B24AFFB542979104FDECFAFC" author="Tyndale-Biscoe, M. &amp; George, R. W." box="[151,532,1375,1400]" pageId="14" pageNumber="265" pagination="65 - 96" refId="ref7362" refString="Tyndale-Biscoe, M. &amp; George, R. W. (1962) The Oxystomata and Gymnopleura (Crustacea, Brachyura) of Western Australia with descriptions of two new species from Western Australia and one from India. Journal of the Royal Society of Western Australia, 45, 65 - 96." type="journal article" year="1962">Tyndale-Biscoe &amp; George (1962)</bibRefCitation>
from
<collectingCountry id="62CFC9E8B24AFFB5405C9104FD3BFAFC" box="[604,707,1375,1400]" name="Australia" pageId="14" pageNumber="265">Australia</collectingCountry>
appear to be different:
<bibRefCitation id="7E49F489B24AFFB541D29104FAE1FAFC" author="Stephensen, K." box="[978,1305,1375,1400]" pageId="14" pageNumber="265" pagination="57 - 237" refId="ref7298" refString="Stephensen, K. (1946) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, 4, 57 - 237." type="journal article" year="1946">Stephensens (1946: fig. 9D)</bibRefCitation>
illustration shows a simple distal part, whereas
<bibRefCitation id="7E49F489B24AFFB5404191DFFBA2FA19" author="Tyndale-Biscoe, M. &amp; George, R. W." box="[577,1114,1412,1437]" pageId="14" pageNumber="265" pagination="65 - 96" refId="ref7362" refString="Tyndale-Biscoe, M. &amp; George, R. W. (1962) The Oxystomata and Gymnopleura (Crustacea, Brachyura) of Western Australia with descriptions of two new species from Western Australia and one from India. Journal of the Royal Society of Western Australia, 45, 65 - 96." type="journal article" year="1962">Tyndale-Biscoe &amp; George (1962: 87, fig. 7.6)</bibRefCitation>
specifically mentioned that “Distal third divided into two by deep cleft”. We examined Japanese specimens undoubtedly identified with
<taxonomicName id="DDD8F2FBB24AFFB5478791F2FEFDFA60" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5478791F2FEFDFA60" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
, as well as the redescription of the
<typeStatus id="C56337DAB24AFFB540939197FD0DFA61" box="[659,757,1484,1509]" pageId="14" pageNumber="265" type="holotype">holotype</typeStatus>
by
<bibRefCitation id="7E49F489B24AFFB5411F9196FBB6FA60" author="Komatsu, H. &amp; Takeda, M." box="[799,1102,1484,1509]" pageId="14" pageNumber="265" pagination="267 - 283" refId="ref6441" refString="Komatsu, H. &amp; Takeda, M. (2000) Leucosiid crabs (Crustacea: Decapoda: Brachyura) from the Osumi Islands, southwest Japan, with description of a new species of Cryptocnemus. Species Diversity, 5, 267 - 283. https: // doi. org / 10.12782 / specdiv. 5.267" type="journal article" year="2000">Komatsu &amp; Takeda (2000)</bibRefCitation>
, for comparison with the new species and
<taxonomicName id="DDD8F2FBB24AFFB5431F91AAFE4BF98C" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[287,435,1520,1544]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB5431F91AAFE4BF98C" box="[287,435,1520,1544]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
. Unfortunately, no adult male specimens of
<taxonomicName id="DDD8F2FBB24AFFB541A691AAFBC8F98C" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[934,1072,1520,1544]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB541A691AAFBC8F98C" box="[934,1072,1520,1544]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
were available for examination, but in one immature male specimen, although the gonopod 1 was not fully developed, the distal part was already bifurcate. It can thus be assumed that
<taxonomicName id="DDD8F2FBB24AFFB5403F9262FD32F9D4" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[575,714,1592,1616]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5403F9262FD32F9D4" box="[575,714,1592,1616]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
also has a distally bifurcate gonopod 1, like
<taxonomicName id="DDD8F2FBB24AFFB546C19262FADDF9D4" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[1217,1317,1592,1616]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB546C19262FADDF9D4" box="[1217,1317,1592,1616]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5472D926CFA94F9D4" bold="true" box="[1325,1388,1591,1616]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5472D926CFA94F9D4" box="[1325,1388,1591,1616]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="DDD8F2FBB24AFFB542979206FED2F9F0" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[151,298,1628,1652]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB542979206FED2F9F0" box="[151,298,1628,1652]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
. Nevertheless,
<taxonomicName id="DDD8F2FBB24AFFB543D69206FD98F9F0" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[470,608,1628,1652]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB543D69206FD98F9F0" box="[470,608,1628,1652]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
differs from the latter two species in many features: the frontal part of the carapace is more strongly produced with a proportionately narrower frontal margin in
<taxonomicName id="DDD8F2FBB24AFFB5466E92DAFB02F91C" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[1134,1274,1664,1688]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5466E92DAFB02F91C" box="[1134,1274,1664,1688]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5470A9224FA6CF91C" box="[1290,1428,1663,1688]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10A, C</figureCitation>
) than in the latter two species (
<figureCitation id="82E395FDB24AFFB543F392FFFDABF938" box="[499,595,1700,1725]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Figs 3A</figureCitation>
,
<figureCitation id="82E395FDB24AFFB5406192FFFD7EF939" box="[609,646,1700,1725]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">6A</figureCitation>
); the lateral lobes on the frontal margin are distinctly produced in
<taxonomicName id="DDD8F2FBB24AFFB5478792FEFEFEF964" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5478792FEFEFEF964" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB54314929CFE82F964" box="[276,378,1735,1760]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10D</figureCitation>
) (not produced in
<taxonomicName id="DDD8F2FBB24AFFB540489292FD23F964" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[584,731,1736,1760]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB540489292FD23F964" box="[584,731,1736,1760]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
and
<taxonomicName id="DDD8F2FBB24AFFB541119292FC8DF964" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[785,885,1736,1760]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB541119292FC8DF964" box="[785,885,1736,1760]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5417C929CFC42F964" bold="true" box="[892,954,1735,1760]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5417C929CFC42F964" box="[892,954,1735,1760]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="82E395FDB24AFFB541C9929CFBD2F964" box="[969,1066,1735,1760]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Figs. 3B</figureCitation>
,
<figureCitation id="82E395FDB24AFFB54636929CFBA1F964" box="[1078,1113,1735,1760]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">6B</figureCitation>
); the carapace dorsal surface is relatively smoother and less punctate in
<taxonomicName id="DDD8F2FBB24AFFB5407C92B6FCFFF880" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[636,775,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5407C92B6FCFFF880" box="[636,775,1772,1796]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5411792B7FCA4F880" box="[791,860,1772,1797]" captionStart="FIGURE 9" captionStartId="12.[151,250,1974,1996]" captionTargetBox="[151,1435,769,1953]" captionTargetId="figure@12.[151,1435,769,1953]" captionTargetPageId="12" captionText="FIGURE 9. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm), Okinawa Islands, Ryukyu Islands, Japan, carapace in dorsal view, stained with methylene blue to show details of dorsal ornamentation." httpUri="https://zenodo.org/record/2625351/files/figure.png" pageId="14" pageNumber="265">Fig. 9</figureCitation>
) than in
<taxonomicName id="DDD8F2FBB24AFFB541C392B6FBA0F880" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[963,1112,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB541C392B6FBA0F880" box="[963,1112,1772,1796]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5466892B7FB54F880" box="[1128,1196,1772,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1946,1968]" captionTargetBox="[151,1435,913,1925]" captionTargetId="figure@4.[151,1435,913,1925]" captionTargetPageId="4" captionText="FIGURE 2. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand, habitus in dorsal view (left pereopod 3 and right pereopods 3 and 5 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625335/files/figure.png" pageId="14" pageNumber="265">Fig. 2</figureCitation>
) and
<taxonomicName id="DDD8F2FBB24AFFB546ED92B6FAABF880" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[1261,1363,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB546ED92B6FAABF880" box="[1261,1363,1772,1796]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB5475C92B7FA64F881" bold="true" box="[1372,1436,1772,1797]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB5475C92B7FA64F881" box="[1372,1436,1772,1797]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82E395FDB24AFFB5429F9354FF1CF8AC" box="[159,228,1807,1832]" captionStart="FIGURE 5" captionStartId="8.[151,250,1941,1963]" captionTargetBox="[151,1435,828,1919]" captionTargetId="figure@8.[151,1435,828,1919]" captionTargetPageId="8" captionText="FIGURE 5. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan, habitus in dorsal view (left pereopod 2 missing), stained with methylene blue to show details of dorsal ornamentation of carapace." httpUri="https://zenodo.org/record/2625341/files/figure.png" pageId="14" pageNumber="265">Fig. 5</figureCitation>
); the mesogastric median ridge and hepatic elevations on the carapace are more prominent in
<taxonomicName id="DDD8F2FBB24AFFB54711934AFA63F8AC" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[1297,1435,1808,1832]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB54711934AFA63F8AC" box="[1297,1435,1808,1832]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5429F936FFED1F8C8" box="[159,297,1844,1869]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10A, C</figureCitation>
) than in
<taxonomicName id="DDD8F2FBB24AFFB54390936EFDDDF8C8" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[400,549,1844,1868]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB54390936EFDDDF8C8" box="[400,549,1844,1868]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB54036936FFD68F8C8" box="[566,656,1844,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3A</figureCitation>
) and
<taxonomicName id="DDD8F2FBB24AFFB540D1936EFCCFF8C8" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[721,823,1844,1868]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB540D1936EFCCFF8C8" box="[721,823,1844,1868]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB54140936FFC79F8C9" bold="true" box="[832,897,1844,1869]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB54140936FFC79F8C9" box="[832,897,1844,1869]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82E395FDB24AFFB54192936FFC15F8C9" box="[914,1005,1844,1869]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6A</figureCitation>
); the lateral margin of the carapace is nearly smooth or microscopically granular in
<taxonomicName id="DDD8F2FBB24AFFB540C49302FCAEF8F4" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[708,854,1880,1904]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB540C49302FCAEF8F4" box="[708,854,1880,1904]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB5416D930CFC23F8F4" box="[877,987,1879,1904]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10B</figureCitation>
), rather than moderately granular (
<taxonomicName id="DDD8F2FBB24AFFB547879302FEF7F810" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB547879302FEF7F810" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
;
<figureCitation id="82E395FDB24AFFB5431E9327FE8BF810" box="[286,371,1916,1941]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3C</figureCitation>
) or coarsely granular (
<taxonomicName id="DDD8F2FBB24AFFB540769326FD24F810" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[630,732,1916,1940]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB540769326FD24F810" box="[630,732,1916,1940]" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB540E49327FCDBF811" bold="true" box="[740,803,1916,1941]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB540E49327FCDBF811" box="[740,803,1916,1941]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<figureCitation id="82E395FDB24AFFB541339327FC73F811" box="[819,907,1916,1941]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6C</figureCitation>
); the male thoracic sternite 4 is finely granular anterolaterally in
<taxonomicName id="DDD8F2FBB24AFFB5436193FAFE14F83C" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[353,492,1952,1976]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5436193FAFE14F83C" box="[353,492,1952,1976]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB543FE93FBFD9DF83C" box="[510,613,1952,1977]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10E</figureCitation>
), whereas it is more coarsely granular in
<taxonomicName id="DDD8F2FBB24AFFB5464493FAFB21F83C" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[1092,1241,1952,1976]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB5464493FAFB21F83C" box="[1092,1241,1952,1976]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB546EB93FBFABCF83C" box="[1259,1348,1952,1977]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="14" pageNumber="265">Fig. 3D</figureCitation>
) and
<taxonomicName id="DDD8F2FBB24AFFB5478793FAFF18F858" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24AFFB5478793FAFF18F858" italics="true" pageId="14" pageNumber="265">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24AFFB542E9939FFED2F859" bold="true" box="[233,298,1988,2013]" pageId="14" pageNumber="265">
<taxonomicNameLabel id="339FE811B24AFFB542E9939FFED2F859" box="[233,298,1988,2013]" pageId="14" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82E395FDB24AFFB5433B939FFE6FF859" box="[315,407,1988,2013]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="14" pageNumber="265">Fig. 6D</figureCitation>
); and the cheliped palm is more robust in
<taxonomicName id="DDD8F2FBB24AFFB5417D939EFBF1F858" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[893,1033,1988,2012]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24AFFB5417D939EFBF1F858" box="[893,1033,1988,2012]" italics="true" pageId="14" pageNumber="265">P. tridentata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24AFFB54619939FFB66F858" box="[1049,1182,1988,2013]" captionStart="FIGURE 10" captionStartId="13.[151,250,1904,1926]" captionTargetBox="[229,1373,222,1867]" captionTargetId="figure@13.[175,1412,193,1887]" captionTargetPageId="13" captionText="FIGURE 10. Pseudophilyra tridentata Miers, 1879, CBM-ZC 7846, male (6.1×5.3 mm) (A, B, DF, H), female (8.1×7.2 mm) (C, G), Okinawa Islands, Ryukyu Islands, Japan. A, C, carapace, dorsal view (dorsal ornamentation omitted); B, left pterygostomial margin, dorsal view; D, front, dorsal view; E, thoracic sternites 3 and 4, including episternites 4; F, left cheliped, dorsal view; G, right chela, dorsal view; H, left gonopod 1, ventral view." httpUri="https://zenodo.org/record/2625353/files/figure.png" pageId="14" pageNumber="265">Fig. 10F, G</figureCitation>
) than in
<taxonomicName id="DDD8F2FBB24AFFB54707939EFA64F858" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[1287,1436,1988,2012]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24AFFB54707939EFA64F858" box="[1287,1436,1988,2012]" italics="true" pageId="14" pageNumber="265">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24BFFB4429F94CCFEE7FF34" box="[159,287,151,176]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="15" pageNumber="266">Figs. 3F, G</figureCitation>
,
<figureCitation id="82E395FDB24BFFB4432C94C3FEB5FF34" box="[300,333,152,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1588,1610]" captionTargetBox="[187,1409,226,1539]" captionTargetId="figure@6.[150,1436,193,1567]" captionTargetPageId="6" captionText="FIGURE 4. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 7053, female (8.2×7.4 mm), Ishigaki Island, Yaeyama Islands, Ryukyu Islands. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon and telson, ventral view." httpUri="https://zenodo.org/record/2625339/files/figure.png" pageId="15" pageNumber="266">4B</figureCitation>
) and
<taxonomicName id="DDD8F2FBB24BFFB4438F94C2FE0DFF34" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[399,501,152,176]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24BFFB4438F94C2FE0DFF34" box="[399,501,152,176]" italics="true" pageId="15" pageNumber="266">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24BFFB443FE94CCFDC7FF34" bold="true" box="[510,575,151,176]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="339FE811B24BFFB443FE94CCFDC7FF34" box="[510,575,151,176]" pageId="15" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82E395FDB24BFFB4405094CCFD29FF2B" box="[592,721,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="15" pageNumber="266">Figs. 6F, B</figureCitation>
,
<figureCitation id="82E395FDB24BFFB440DE94C3FCF8FF34" box="[734,768,152,176]" captionStart="FIGURE 7" captionStartId="10.[151,250,1575,1597]" captionTargetBox="[188,1389,280,1509]" captionTargetId="figure@10.[151,1435,215,1543]" captionTargetPageId="10" captionText="FIGURE 7. Pseudophilyra parilis n. sp., CBM-ZC 14486, paratype, female (7.9×7.3 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, sternopleonal cavity, ventral view; C, right vulva, ventral view; D, right chela, dorsal view; E, pleon, ventral view (telson broken off)." httpUri="https://zenodo.org/record/2625345/files/figure.png" pageId="15" pageNumber="266">7B</figureCitation>
) (1.8 times as long as wide in
<taxonomicName id="DDD8F2FBB24BFFB4466394C2FB16FF34" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[1123,1262,152,176]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24BFFB4466394C2FB16FF34" box="[1123,1262,152,176]" italics="true" pageId="15" pageNumber="266">P. tridentata</emphasis>
</taxonomicName>
versus 2.02.2 times as long as wide in the latter two species). Furthermore,
<taxonomicName id="DDD8F2FBB24BFFB4414A94E6FC2CFF50" ID-CoL="78H9R" authorityName="Miers" authorityYear="1879" box="[842,980,188,212]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="tridentata">
<emphasis id="28AC556AB24BFFB4414A94E6FC2CFF50" box="[842,980,188,212]" italics="true" pageId="15" pageNumber="266">P. tridentata</emphasis>
</taxonomicName>
seems to be restricted to subtidal depths below
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, whereas
<taxonomicName id="DDD8F2FBB24BFFB4438C94BAFDE7FF7C" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[396,543,224,248]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24BFFB4438C94BAFDE7FF7C" box="[396,543,224,248]" italics="true" pageId="15" pageNumber="266">P. punctulata</emphasis>
</taxonomicName>
and
<taxonomicName id="DDD8F2FBB24BFFB4405594BAFD42FF7C" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[597,698,224,248]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24BFFB4405594BAFD42FF7C" box="[597,698,224,248]" italics="true" pageId="15" pageNumber="266">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24BFFB440C19484FCF8FF7C" bold="true" box="[705,768,223,248]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="339FE811B24BFFB440C19484FCF8FF7C" box="[705,768,223,248]" pageId="15" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
occur in intertidal to shallow subtidal depths.
</paragraph>
<paragraph id="1A678978B24BFFB442C7955EFDE3FE0C" blockId="15.[151,1437,151,464]" pageId="15" pageNumber="266">
It may be interesting to mention that the episternites 4 are partially fused to the thoracic sternite
<quantity id="DD20249DB24BFFB44707955FFACBFE99" box="[1287,1331,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="266" unit="in" value="4.0">4 in</quantity>
the adult male of
<taxonomicName id="DDD8F2FBB24BFFB442F19572FE7CFEC4" ID-CoL="8YYN7" authorityName="Chen &amp; Ng" authorityYear="2003" box="[241,388,296,320]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24BFFB442F19572FE7CFEC4" box="[241,388,296,320]" italics="true" pageId="15" pageNumber="266">P. punctulata</emphasis>
</taxonomicName>
(
<figureCitation id="82E395FDB24BFFB44393957CFE13FEC4" box="[403,491,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1869,1891]" captionTargetBox="[223,1347,233,1832]" captionTargetId="figure@5.[204,1387,193,1853]" captionTargetPageId="5" captionText="FIGURE 3. Pseudophilyra punctulata Chen &amp; Ng, 2003, CBM-ZC 12912, male (6.7×6.0 mm), Phuket, Thailand. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, pleon and telson, ventral view; I, left gonopod 1, ventral view; J, same, dorsal view; K, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625337/files/figure.png" pageId="15" pageNumber="266">Fig. 3D</figureCitation>
), whereas they are fully separated from the thoracic sternite
<quantity id="DD20249DB24BFFB4468B9573FB4DFEC4" box="[1163,1205,295,320]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="266" unit="in" value="4.0">4 in</quantity>
the adult males of
<taxonomicName id="DDD8F2FBB24BFFB447879572FF18FEE0" ID-CoL="84G7B" authority="Komai &amp; Shimetsugu &amp; Ng, 2019" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="parilis" status="sp. nov.">
<emphasis id="28AC556AB24BFFB447879572FF18FEE0" italics="true" pageId="15" pageNumber="266">P. parilis</emphasis>
</taxonomicName>
<emphasis id="28AC556AB24BFFB442E89517FED1FEE1" bold="true" box="[232,297,332,357]" pageId="15" pageNumber="266">
<taxonomicNameLabel id="339FE811B24BFFB442E89517FED1FEE1" box="[232,297,332,357]" pageId="15" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82E395FDB24BFFB443399517FE6DFEE1" box="[313,405,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1899]" captionTargetBox="[214,1360,231,1827]" captionTargetId="figure@9.[196,1379,193,1858]" captionTargetPageId="9" captionText="FIGURE 6. Pseudophilyra parilis n. sp., CBM-ZC 14485, holotype, male (7.2×6.2 mm), Mie Prefecture, Japan. A, carapace, dorsal view (dorsal ornamentation omitted); B, front of carapace and eyes, dorsal view; C, left pterygostomial margin of carapace, dorsal view; D, thoracic sternites 3 and 4, including episternites 4, ventral view; E, left maxilliped 3, outer view (setae omitted); F, right cheliped, dorsal view; G, same, ventral view; H, right pereopod 5, dorsal view; I, pleon and telson, ventral view; J, left gonopod 1, ventral view; K, same, dorsal view; L, left gonopod 2, lateral view." httpUri="https://zenodo.org/record/2625343/files/figure.png" pageId="15" pageNumber="266">Fig. 6D</figureCitation>
). The significance of this character should be reassessed when more adult male specimens become available for examination.
</paragraph>
</subSubSection>
<subSubSection id="52C2DAF3B24BFFB442C795CFFE38FE4B" pageId="15" pageNumber="266" type="etymology">
<paragraph id="1A678978B24BFFB442C795CFFE38FE4B" blockId="15.[151,1437,151,464]" pageId="15" pageNumber="266">
<emphasis id="28AC556AB24BFFB442C795CFFEB6FE29" bold="true" box="[199,334,404,429]" pageId="15" pageNumber="266">Etymology.</emphasis>
From the Latin “
<taxonomicName id="DDD8F2FBB24BFFB4401695CFFDA7FE28" authorityName="Komai &amp; Shimetsugu &amp; Ng" authorityYear="2019" box="[534,607,404,428]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="parilis">
<emphasis id="28AC556AB24BFFB4401695CFFDA7FE28" box="[534,607,404,428]" italics="true" pageId="15" pageNumber="266">parilis</emphasis>
</taxonomicName>
” (adj., similar), in reference to the close similarity of the new species to
<taxonomicName id="DDD8F2FBB24BFFB4429795E3FE41FE54" authorityName="Chen &amp; Ng" authorityYear="2003" box="[151,441,440,464]" class="Malacostraca" family="Leucosiidae" genus="Pseudophilyra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="punctulata">
<emphasis id="28AC556AB24BFFB4429795E3FE41FE54" box="[151,441,440,464]" italics="true" pageId="15" pageNumber="266">Pseudophilyra punctulata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>