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<document id="E7B7D909AC69F8A66961E39EC35962DF" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. &amp; Hughes, Nigel C." docDate="2022" docId="03CB87846E711D2DFF3AC1184E73F4E1" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Reedocalymene unicornis Turvey" docType="treatment" docVersion="6" lastPageNumber="337" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="337" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C70FD5D198E29B5FDC580680FF2664DE">Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:namePart id="BA0FB220791F5A6EF9E0478FEF1DE28E">Fortey, Richard A.</mods:namePart>
<mods:affiliation id="77575A16DEA73BA29878872715F7F07B">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="9EF297D3C098502A796F39B58C3ECF82">Wernette, Shelly J.</mods:namePart>
<mods:affiliation id="752EC6E36880B4E42EE7477F5101C6A4">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. &amp; Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<mods:namePart id="170A726220A65D46729E86A498F4A5BE">Hughes, Nigel C.</mods:namePart>
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<mods:date id="FD536CC934631A1056591B3223B75E95">2022</mods:date>
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<treatment id="03CB87846E711D2DFF3AC1184E73F4E1" ID-DOI="http://doi.org/10.5281/zenodo.6810354" ID-GBIF-Taxon="197214827" ID-Zenodo-Dep="6810354" LSID="urn:lsid:plazi:treatment:03CB87846E711D2DFF3AC1184E73F4E1" httpUri="http://treatment.plazi.org/id/03CB87846E711D2DFF3AC1184E73F4E1" lastPageId="37" lastPageNumber="337" pageId="36" pageNumber="337">
<subSubSection id="C37865196E711D2CFF3AC1184868F03F" box="[151,627,1369,1396]" pageId="36" pageNumber="337" type="nomenclature">
<paragraph id="8BDD36926E711D2CFF3AC1184868F03F" blockId="36.[151,627,1369,1432]" box="[151,627,1369,1396]" pageId="36" pageNumber="337">
<heading id="D09581FE6E711D2CFF3AC1184868F03F" bold="true" box="[151,627,1369,1396]" fontSize="11" level="1" pageId="36" pageNumber="337" reason="1">
<taxonomicName id="4C624D116E711D2CFF3AC1184868F03F" ID-CoL="4RPNF" authority="(Reed, 1917)" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[151,627,1369,1396]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFF3AC1184868F03F" bold="true" box="[151,627,1369,1396]" pageId="36" pageNumber="337">
<emphasis id="B916EA806E711D2CFF3AC1184BCBF03F" bold="true" box="[151,464,1369,1396]" italics="true" pageId="36" pageNumber="337">Reedocalymene unicornis</emphasis>
(
<bibRefCitation id="EFF34B636E711D2CFE72C1184870F03F" author="Reed, F. R. C." box="[479,619,1369,1395]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37865196E711D2CFF3AC13C496BF096" pageId="36" pageNumber="337" type="description">
<paragraph id="8BDD36926E711D2CFF3AC13C4B4FF0DB" blockId="36.[151,627,1369,1432]" box="[151,340,1405,1432]" pageId="36" pageNumber="337">
<figureCitation id="13592A176E711D2CFF3AC13C4B07F0DB" box="[151,284,1405,1432]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="36" pageNumber="337">Figs 12.1–6</figureCitation>
,
<figureCitation id="13592A176E711D2CFE8FC13C4B2BF0DB" box="[290,304,1405,1431]" captionStart="FIGURE 8" captionStartId="22.[151,250,1587,1611]" captionTargetBox="[273,1314,181,1563]" captionTargetId="figure-29@22.[273,1314,181,1563]" captionTargetPageId="22" captionText="FIGURE 8. (1–6) Birmanites yunnanensis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1. Librigena (original of Reed 1917, pl. 7, fig. 3), GSI 11892. 2. Pygidium (original of Reed 1917, pl. 7, fig. 6), GSI 11895. 3. Pygidium (original of Reed 1917, pl. 7, fig. 8), GSI 11897. 4. Librigena (original of Reed 1917, pl. 7, fig. 5), GSI 11894. 5. Articulated specimen (original of Reed 1917, pl. 6, fig. 12), GSI 11887. 6. Cephalon (original of Reed 1917, pl. 6, fig. 14), GSI 11889. (7, 10) Remopleuriidae gen. et sp. indet. (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 7. Cranidium, (original of Reed 1917, pl. 6, fig. 9), GSI 11884. 10. Librigena (original of Reed 1917, pl. 6, fig. 10), GSI 11885. (8) Dolerobasilicus? sp. (Reed), Pygidium (original of Reed 1915, pl. 7, fig. 3), GSI 11535, Lower Naungkangyi Beds, Nawa, northern Shan State, Myanmar. (9) Nileus sp., Hypostome (original of Reed 1917, pl. 8, fig. 5), GSI 11904, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (11, 12) Niobe sp. (Reed). 11a, b. Pygidium (original of Reed 1915, pl. 5, fig. 14), GSI 11516, Hwe Mawng Beds (uppermost Katian), Nati, northern part of Shan State, Myanmar, original and retrodeformation. 12. Pygidium (original of Reed 1917, pl. 6, fig. 11), GSI 11886, Shihtien Formation (Darriwilian), Shidian, Baoshan Prefecture, western Yunnan, China. Scale bars = 5 mm for 1–6, 8, 9; = 2 mm for 7, 10–12)" figureDoi="http://doi.org/10.5281/zenodo.6810312" httpUri="https://zenodo.org/record/6810312/files/figure.png" pageId="36" pageNumber="337">8</figureCitation>
,
<figureCitation id="13592A176E711D2CFE98C13C4B4FF0DB" box="[309,340,1405,1431]" captionStart="FIGURE 10" captionStartId="31.[151,250,1456,1480]" captionTargetBox="[287,1301,181,1432]" captionTargetId="figure-18@31.[287,1301,181,1432]" captionTargetPageId="31" captionText="FIGURE 10. (1, 2) Lonchodamas? sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 1a, b. Pygidium (original of Reed 1915, pl. 5, fig. 6), GSI 11508, original and retrodeformation. 2. Cranidium (refigured from Reed 1915, pl. 5, fig. 5), GSI 11507. (3, 6) Lonchodamas shanensis (Reed) from the Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 3. Pygidium (original of Reed 1915, pl. 5, fig. 3), GSI 11505, Hwe-hok. 6. Cranidium (original of Reed 1915, pl. 5, fig. 2), GSI 11504, Mong Ha. (4) Dionide hybrida (Reed), Cephalon (original of Reed 1915, pl. 5, fig. 7), GSI 11509, Hwe Mawng Beds (uppermost Katian), Hwe-hok, northern Shan State, Myanmar. (5, 8) Ovalocephalus? sp. indet. (Reed) from the Li-Lu Formation (Katian), Mân-ngai, northern part of Shan State, Myanmar. 5. Cranidium (original of Reed 1915, pl. 8, fig. 17), GSI 11563. 8. Thorax (original of Reed 1915, pl. 8, fig. 21), GSI 11567. (7) Pliomerina martellii (Reed), Cranidium (original of Reed 1917, pl. 8, fig. 15), GSI 11915, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. (9) Prionocheilus liluensis (Reed), Cranidium (original of Reed 1915, pl. 8 figs 6 and 7), GSI 11554, Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. (10, 13) Ovalocephalus sp. (Reed) from the Li-Lu Formation (Katian), Lilu, northern Shan State, Myanmar. 10. Pygidium (original of Reed 1915, pl. 8, fig. 9), GSI 11556. 13. Cephalon; not previously figured; no GSI catalogue number. (11, 12) “Calymene” oldhami (Reed) from Lower Naungkangyi Beds (probable Darriwilian), Mâ Shio, northern Shan State, Myanmar. 11. Cephalon and thorax (original of Reed 1915, pl. 8, fig. 11), GSI 11558. 12. Cephalon and partial thorax (original of Reed 1915, pl. 8, fig. 10), GSI 11557. Scale bars = 2 mm for 1–3, 6, 8–13; = 5 mm for 4, 5; = 1 mm for 7." figureDoi="http://doi.org/10.5281/zenodo.6810316" httpUri="https://zenodo.org/record/6810316/files/figure.png" pageId="36" pageNumber="337">10</figureCitation>
</paragraph>
<paragraph id="8BDD36926E711D2CFF3AC183496BF096" blockId="36.[151,1017,1474,1621]" box="[151,880,1474,1498]" pageId="36" pageNumber="337">
1917
<taxonomicName id="4C624D116E711D2CFF5AC1834BD8F095" authority="Fortey &amp; Wernette &amp; Hughes, 2022" authorityName="Fortey &amp; Wernette &amp; Hughes" authorityYear="2022" box="[247,451,1474,1498]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
<emphasis id="B916EA806E711D2CFF5AC1834BD8F095" box="[247,451,1474,1498]" italics="true" pageId="36" pageNumber="337">Calymene unicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22557FB6E711D2CFE64C1834801F095" box="[457,538,1474,1497]" pageId="36" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
; Reed p. 52–54, pl. 8, figs 10–13.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E711D2CFF3AC1A1493DF318" pageId="36" pageNumber="337" type="reference_group">
<paragraph id="8BDD36926E711D2CFF3AC1A149E2F0B5" blockId="36.[151,1017,1474,1621]" box="[151,1017,1504,1529]" pageId="36" pageNumber="337">
1951
<taxonomicName id="4C624D116E711D2CFF5AC1A14891F0B5" ID-CoL="4RPNF" authority="(Reed, 1917)" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[247,650,1504,1529]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFF5AC1A14BE7F0B4" box="[247,508,1504,1528]" italics="true" pageId="36" pageNumber="337">Reedocalymene unicornis</emphasis>
(
<bibRefCitation id="EFF34B636E711D2CFDA7C1A14899F0B5" author="Reed, F. R. C." box="[522,642,1504,1529]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
)
</taxonomicName>
; Kobayashi, p. 45, pl. 4, figs 12–14.
</paragraph>
<paragraph id="8BDD36926E711D2CFF3AC1BE4918F35B" blockId="36.[151,1017,1474,1621]" box="[151,771,1535,1559]" pageId="36" pageNumber="337">
1975
<taxonomicName id="4C624D116E711D2CFF5AC1BE4891F35B" ID-CoL="4RPNF" authority="(Reed, 1917)" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[247,650,1535,1559]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFF5AC1BE4BE7F35B" box="[247,508,1535,1559]" italics="true" pageId="36" pageNumber="337">Reedocalymene unicornis</emphasis>
(
<bibRefCitation id="EFF34B636E711D2CFDA7C1BE4899F35B" author="Reed, F. R. C." box="[522,642,1535,1559]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
)
</taxonomicName>
; Lu, p. 447.
</paragraph>
<paragraph id="8BDD36926E711D2CFF3AC25F493BF37A" blockId="36.[151,1017,1474,1621]" box="[151,800,1566,1590]" pageId="36" pageNumber="337">
2002
<taxonomicName id="4C624D116E711D2CFF5AC25F4891F37A" ID-CoL="4RPNF" authority="(Reed, 1917)" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[247,650,1566,1590]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFF5AC25F4BE7F379" box="[247,508,1566,1590]" italics="true" pageId="36" pageNumber="337">Reedocalymene unicornis</emphasis>
(
<bibRefCitation id="EFF34B636E711D2CFDA7C25F4899F37A" author="Reed, F. R. C." box="[522,642,1566,1590]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
)
</taxonomicName>
; Turvey, p. 55.
</paragraph>
<paragraph id="8BDD36926E711D2CFF3AC27D493DF318" blockId="36.[151,1017,1474,1621]" box="[151,806,1596,1621]" pageId="36" pageNumber="337">
2005
<taxonomicName id="4C624D116E711D2CFF5AC27D48CDF319" ID-CoL="4RPNF" authority="(Reed, 1917) Turvey" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[247,726,1596,1621]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFF5AC27D4BE7F318" box="[247,508,1596,1620]" italics="true" pageId="36" pageNumber="337">Reedocalymene unicornis</emphasis>
(
<bibRefCitation id="EFF34B636E711D2CFDA7C27D489BF319" author="Reed, F. R. C." box="[522,640,1596,1621]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
) Turvey
</taxonomicName>
, p. 550.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E711D2CFF3AC2C348C4F245" pageId="36" pageNumber="337" type="materials_examined">
<paragraph id="8BDD36926E711D2CFF3AC2C348C4F245" blockId="36.[151,1437,1666,2017]" pageId="36" pageNumber="337">
<emphasis id="B916EA806E711D2CFF3AC2C34B1DF3D0" bold="true" box="[151,262,1666,1692]" pageId="36" pageNumber="337">Material.</emphasis>
<typeStatus id="54D988306E711D2CFEA2C2C24B64F3D1" box="[271,383,1667,1693]" pageId="36" pageNumber="337" type="lectotype">Lectotype</typeStatus>
(selected by
<bibRefCitation id="EFF34B636E711D2CFDBAC2C248AAF3D1" author="Turvey, S. T." box="[535,689,1666,1693]" pageId="36" pageNumber="337" pagination="549 - 575" refId="ref48004" refString="Turvey, S. T. (2005) Reedocalymenine trilobites from the Ordovician of central and eastern Asia, and a review of species assigned to Neseuretus. Palaeontology, 48, 549 - 575. https: // doi. org / 10.1111 / j. 1475 - 4983.2005.00469. x" type="journal article" year="2005">Turvey, 2005</bibRefCitation>
): damaged cephalon,
<figureCitation id="13592A176E711D2CFC0AC2C24E33F3D1" box="[935,1064,1667,1693]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="36" pageNumber="337">Figs 12.1-3</figureCitation>
(
<bibRefCitation id="EFF34B636E711D2CFB95C2C24EA6F3D1" author="Reed, F. R. C." box="[1080,1213,1667,1693]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 8, fig. 10),
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11913. Additional type material: cephalic fragments,
<figureCitation id="13592A176E711D2CFD40C2E6494EF38D" box="[749,853,1702,1729]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="36" pageNumber="337">Figs 12.5</figureCitation>
/8 and 12.10/6 (
<bibRefCitation id="EFF34B636E711D2CFBA8C2E64E91F38D" author="Reed, F. R. C." box="[1029,1162,1703,1729]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 8, figs 12,11 respectively),
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11912; pygidium,
<figureCitation id="13592A176E711D2CFE57C28A487BF3A9" box="[506,608,1739,1765]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="36" pageNumber="337">Fig. 12.4</figureCitation>
(
<bibRefCitation id="EFF34B636E711D2CFDDFC28A48E3F3A9" author="Reed, F. R. C." box="[626,760,1739,1765]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 8, fig. 13),
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11911? All material from the Shihtien Formation (Darriwilian) at Pupiao, western
<collectingRegion id="49A6F8706E711D2CFD2FC2AE48C0F245" box="[642,731,1775,1801]" country="China" name="Yunnan" pageId="36" pageNumber="337">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E711D2DFF6AC3534E73F4E1" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337" type="discussion">
<paragraph id="8BDD36926E711D2DFF6AC3534E73F4E1" blockId="36.[151,1437,1666,2017]" lastBlockId="37.[151,1437,151,429]" lastPageId="37" lastPageNumber="338" pageId="36" pageNumber="337">
<emphasis id="B916EA806E711D2CFF6AC3534B57F260" bold="true" box="[199,332,1810,1836]" pageId="36" pageNumber="337">Discussion.</emphasis>
<bibRefCitation id="EFF34B636E711D2CFEF8C3524BF9F261" author="Reed, F. R. C." box="[341,482,1811,1837]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917)</bibRefCitation>
described the limited material of this species in some detail, and new cast material does not greatly enhance our knowledge. The
<typeStatus id="54D988306E711D2CFD09C3764910F21D" box="[676,779,1847,1873]" pageId="36" pageNumber="337" type="lectotype">lectotype</typeStatus>
is more distorted than Brock’s original drawing suggests, and the upward curvature of the anterior pro/jection shown by
<bibRefCitation id="EFF34B636E711D2CFCC8C31A49F0F239" author="Reed, F. R. C." box="[869,1003,1883,1909]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917</bibRefCitation>
, pl. 8, fig. 10a, GSI 11913) may have been exaggerated by this. However, the comparatively laterally expanded L1 on the glabella is probably a real feature, which is not so clear on the original drawing. This differs from other species assigned to the genus by
<bibRefCitation id="EFF34B636E711D2CFA80C3E24F87F2F1" author="Lu, Y. - H." box="[1325,1436,1954,1981]" pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu (1975)</bibRefCitation>
that have a uniformly deeply parabolic glabellar outline. In this respect
<taxonomicName id="4C624D116E711D2CFC1CC3864E22F2AC" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[945,1081,1991,2017]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E711D2CFC1CC38649D9F2AC" box="[945,962,1991,2016]" italics="true" pageId="36" pageNumber="337">R</emphasis>
.
<emphasis id="B916EA806E711D2CFC62C3864E22F2AC" box="[975,1081,1991,2016]" italics="true" pageId="36" pageNumber="337">unicornis</emphasis>
</taxonomicName>
more closely resembles species that have been assigned to
<taxonomicName id="4C624D116E701D2DFE66C4D64879F5FD" authorityName="Kobayashi" authorityYear="1951" box="[459,610,151,177]" class="Trilobita" family="Bathycheilidae" genus="Calymenesun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="37" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E701D2DFE66C4D64879F5FD" box="[459,610,151,177]" italics="true" pageId="37" pageNumber="338">Calymenesun</emphasis>
</taxonomicName>
and
<taxonomicName id="4C624D116E701D2DFD36C4D64917F5FC" authorityName="Kobayashi" authorityYear="1960" box="[667,780,151,176]" class="Magnoliopsida" family="Apocynaceae" genus="Vietnamia" kingdom="Plantae" order="Gentianales" pageId="37" pageNumber="338" phylum="Tracheophyta" rank="genus">
<emphasis id="B916EA806E701D2DFD36C4D64917F5FC" box="[667,780,151,176]" italics="true" pageId="37" pageNumber="338">Vietnamia</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF34B636E701D2DFCE4C4D649FFF5FD" author="Turvey, S. T." box="[841,996,151,177]" pageId="37" pageNumber="338" pagination="53 - 68" refId="ref47933" refString="Turvey, S. T. (2002) Phylogeny of the Reedocalymeninae (Trilobita): implications for Early Ordovician biogeography of Gondwana. In: Crame, A. J. &amp; Owen, A. W. (Eds.), Palaeobiogeography and biodiversity change: the Ordovician and Mesozoic - Cenozoic radiations. Geological Society of London Special Publications, 194, 53 - 68." type="journal article" year="2002">Turvey, 2002</bibRefCitation>
). However, it is possible that deformation may have contributed to this distinction. A very incomplete cranidium shows the outline of the posterior fixed cheek. A backwardly displaced free cheek with a narrow border is best displayed on Reed’s pl. 8, fig. 11; we have not been able to cast this specimen, nor the only example of a pygidium (
<bibRefCitation id="EFF34B636E701D2DFC13C5424E5AF451" author="Reed, F. R. C." box="[958,1089,259,285]" pageId="37" pageNumber="338" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
, pl. 8, fig. 13) reproduced here. The imperfect thorax and pygidium from Pupiao figured by
<bibRefCitation id="EFF34B636E701D2DFC95C56649A6F40D" author="Reed, F. R. C." box="[824,957,295,321]" pageId="37" pageNumber="338" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917</bibRefCitation>
, pl. 8, fig. 14) is refigured here (
<figureCitation id="13592A176E701D2DFA8AC5664F8BF40D" box="[1319,1424,295,321]" captionStart="FIGURE 12" captionStartId="38.[151,250,1593,1617]" captionTargetBox="[277,1311,181,1569]" captionTargetId="figure-29@38.[277,1311,181,1569]" captionTargetPageId="38" captionText="FIGURE 12. (1–6,8,10) Reedocalymene unicornis (Reed) from the Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. 1–3. Cranidium (original and refigured of Reed 1917, pl. 8, fig. 10a,b), GSI 11913. 4. Pygidium (refigured from Reed 1917, pl. 8, fig. 13), GSI 11911?. 5. Partial cranidium (original of Reed 1917, pl. 8, fig. 12); GSI 11912?. 6. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 11), GSI 11912?. 8. Partial cranidium (refigured from Reed 1917, pl. 8, fig. 12), GSI 11912?. 10. Partial cranidium (original of Reed 1917, pl. 8, fig. 11), GSI 11912?. (7) Reedocalymene? sp. (Reed), Thorax and pygidium (original of Reed 1917, pl. 8, fig. 14), GSI 11914, Shihtien Formation (Darriwilian); Pupiao, Baoshan Prefecture, western Yunnan, China. (9,11–15) Neseuretinus aff. birmanicus (Reed) from the Li-Lu and Pupiao formations (Katian). 9. Cranidium (lectotype, original of Reed 1906, pl. 5, fig. 27), GSI 8341, Kunkaw, northern Shan State, Myanmar. 11. Cranidium; CMC IP 91532, Pupiao, Baoshan Prefecture, western Yunnan, China. 12. Pygidium and partial thorax, CMC IP 91533, Pupiao, Baoshan Prefecture, western Yunnan, China. 13. Pygidium, CMC IP 91534, Pupiao, Baoshan Prefecture, western Yunnan, China. 14, 15. Pygidium (original of Reed 1915, pl. 8, fig. 5), GSI 11553, Loikok, northern part of Shan State, Myanmar, dorsal and right lateral views. Scale bars = 2 mm for 1–6, 8–10, 12–15; = 5 mm for 7, 11." figureDoi="http://doi.org/10.5281/zenodo.6810324" httpUri="https://zenodo.org/record/6810324/files/figure.png" pageId="37" pageNumber="338">Fig. 12.7</figureCitation>
). It is likely to belong to a reedocalymenine, and possibly to
<taxonomicName id="4C624D116E701D2DFC9DC50A49FBF429" authorityName="Kobayashi" authorityYear="1951" box="[816,992,331,357]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="37" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E701D2DFC9DC50A49FBF429" box="[816,992,331,357]" italics="true" pageId="37" pageNumber="338">Reedocalymene</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C624D116E701D2DFBCAC50A4EE3F428" authorityName="Dean" authorityYear="1967" box="[1127,1272,331,356]" class="Trilobita" family="Calymenidae" genus="Neseuretinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="37" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E701D2DFBCAC50A4EE3F428" box="[1127,1272,331,356]" italics="true" pageId="37" pageNumber="338">Neseuretinus</emphasis>
</taxonomicName>
. However, the pygidial pleural ribs appear to be more pronounced than is the case with the pygidium assigned with more certainty to
<taxonomicName id="4C624D116E701D2DFF19C5D24B26F4E0" authorityName="Turvey" baseAuthorityName="Reed" baseAuthorityYear="1917" box="[180,317,403,428]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="37" pageNumber="338" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="B916EA806E701D2DFF19C5D24B26F4E0" box="[180,317,403,428]" italics="true" pageId="37" pageNumber="338">R. unicornis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF34B636E701D2DFECAC5D24BF7F4E1" author="Reed, F. R. C." box="[359,492,403,429]" pageId="37" pageNumber="338" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed (1917</bibRefCitation>
, pl. 8, fig.13), and its assignment must remain uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>
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