Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China
Author
Fortey, Richard A.
Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.
Author
Wernette, Shelly J.
Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. & Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA
Author
Hughes, Nigel C.
text
Zootaxa
2022
2022-07-08
5162
4
301
356
journal article
93239
10.11646/zootaxa.5162.4.1
7d343017-483e-43be-9439-d06e748fc54e
1175-5326
6810290
DD2279FA-E8F1-4951-A5CA-91082E875580
Reedocalymene unicornis
(
Reed, 1917
)
Figs 12.1–6
,
8
,
10
1917
Calymene unicornis
sp. nov.
; Reed p. 52–54, pl. 8, figs 10–13.
1951
Reedocalymene unicornis
(
Reed, 1917
)
; Kobayashi, p. 45, pl. 4, figs 12–14.
1975
Reedocalymene unicornis
(
Reed, 1917
)
; Lu, p. 447.
2002
Reedocalymene unicornis
(
Reed, 1917
)
; Turvey, p. 55.
2005
Reedocalymene unicornis
(
Reed, 1917
) Turvey
, p. 550.
Material.
Lectotype
(selected by
Turvey, 2005
): damaged cephalon,
Figs 12.1-3
(
Reed, 1917
, pl. 8, fig. 10),
GSI
11913. Additional type material: cephalic fragments,
Figs 12.5
/8 and 12.10/6 (
Reed, 1917
, pl. 8, figs 12,11 respectively),
GSI
11912; pygidium,
Fig. 12.4
(
Reed, 1917
, pl. 8, fig. 13),
GSI
11911? All material from the Shihtien Formation (Darriwilian) at Pupiao, western
Yunnan
.
Discussion.
Reed (1917)
described the limited material of this species in some detail, and new cast material does not greatly enhance our knowledge. The
lectotype
is more distorted than Brock’s original drawing suggests, and the upward curvature of the anterior pro/jection shown by
Reed (1917
, pl. 8, fig. 10a, GSI 11913) may have been exaggerated by this. However, the comparatively laterally expanded L1 on the glabella is probably a real feature, which is not so clear on the original drawing. This differs from other species assigned to the genus by
Lu (1975)
that have a uniformly deeply parabolic glabellar outline. In this respect
R
.
unicornis
more closely resembles species that have been assigned to
Calymenesun
and
Vietnamia
(see
Turvey, 2002
). However, it is possible that deformation may have contributed to this distinction. A very incomplete cranidium shows the outline of the posterior fixed cheek. A backwardly displaced free cheek with a narrow border is best displayed on Reed’s pl. 8, fig. 11; we have not been able to cast this specimen, nor the only example of a pygidium (
Reed, 1917
, pl. 8, fig. 13) reproduced here. The imperfect thorax and pygidium from Pupiao figured by
Reed (1917
, pl. 8, fig. 14) is refigured here (
Fig. 12.7
). It is likely to belong to a reedocalymenine, and possibly to
Reedocalymene
rather than
Neseuretinus
. However, the pygidial pleural ribs appear to be more pronounced than is the case with the pygidium assigned with more certainty to
R. unicornis
by
Reed (1917
, pl. 8, fig.13), and its assignment must remain uncertain.