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<document id="79EEDC5E33837A1366C2D91BCE91139C" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F021FFF4FF4877C137702135" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Longicornus longus Xing &amp; Li" docType="treatment" docVersion="2" lastPageNumber="105" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="105" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="C280CF7B887D1B9A9CCAAD3D1D437BC8">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<subSubSection id="C304BDDEF021FFF4FF4877C1345F23F2" box="[151,530,151,178]" pageId="104" pageNumber="105" type="nomenclature">
<paragraph id="8BA1EE55F021FFF4FF4877C1345F23F2" blockId="104.[151,530,151,214]" box="[151,530,151,178]" pageId="104" pageNumber="105">
<heading id="D0E95939F021FFF4FF4877C1345F23F2" bold="true" box="[151,530,151,178]" fontSize="11" level="2" pageId="104" pageNumber="105" reason="4">
<taxonomicName id="4C1E95D6F021FFF4FF4877C1345F23F2" ID-CoL="8RLQX" authority="Xing &amp; Li" authorityName="Xing &amp; Li" box="[151,530,151,178]" class="Insecta" family="Cicadellidae" genus="Longicornus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B96A3247F021FFF4FF4877C1345F23F2" bold="true" box="[151,530,151,178]" pageId="104" pageNumber="105">
<emphasis id="B96A3247F021FFF4FF4877C137C623F1" bold="true" box="[151,395,151,178]" italics="true" pageId="104" pageNumber="105">Longicornus longus</emphasis>
Xing &amp; Li
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF021FFF4FF4877ED37702135" pageId="104" pageNumber="105" type="description">
<paragraph id="8BA1EE55F021FFF4FF4877ED37532395" blockId="104.[151,530,151,214]" box="[151,286,187,214]" pageId="104" pageNumber="105">
<figureCitation id="1325F2D0F021FFF4FF4877ED37532395" box="[151,286,187,214]" captionStart="FIGURES 56" captionStartId="97.[151,264,1860,1884]" captionTargetBox="[156,1417,203,1821]" captionTargetId="figure-17@97.[151,1435,198,1836]" captionTargetPageId="97" captionText="FIGURES 56AL. Longicornus longus (Xing &amp; Li): AB, male habitus, dorsal and lateral views; C, face; D, genital capsule, lateral view; E, pygofer, lateral view; F, valve and subgenital plates, ventral view; G, subgenital plate, lateral view; H, style, dorsal view; I, subgenital plates, styles, connective and aedeagus, dorsal view; J, connective and aedeagus, ventral view; K, style, connective and aedeagus, lateral view; L, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703893" httpUri="https://zenodo.org/record/14703893/files/figure.png" pageId="104" pageNumber="105">Figs 56AL</figureCitation>
</paragraph>
<paragraph id="8BA1EE55F021FFF4FF48765637422275" blockId="104.[151,1436,255,341]" pageId="104" pageNumber="105">
<taxonomicName id="4C1E95D6F021FFF4FF4876563795225B" authority="Xing &amp; Li" authorityName="Xing &amp; Li" box="[151,472,256,280]" class="Insecta" family="Cicadellidae" genus="Longicornus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B96A3247F021FFF4FF487656372E225B" box="[151,355,256,280]" italics="true" pageId="104" pageNumber="105">Longicornus longus</emphasis>
Xing &amp; Li
</taxonomicName>
, in Li
<emphasis id="B96A3247F021FFF4FDC27656341C225B" box="[541,593,256,280]" italics="true" pageId="104" pageNumber="105">et al.</emphasis>
2011: 112, figs 510, 17; Gou &amp; Xing 2022: 438,
<figureCitation id="1325F2D0F021FFF4FBB4765632832254" box="[1131,1230,255,280]" captionStart-0="FIGURES 4" captionStart-1="FIGURES 5" captionStart-2="FIGURES 6" captionStartId-0="45.[151,264,1920,1944]" captionStartId-1="46.[151,264,1828,1852]" captionStartId-2="47.[151,264,1889,1913]" captionTargetBox-0="[157,1430,186,1891]" captionTargetBox-1="[166,1431,205,1799]" captionTargetBox-2="[156,1423,181,1860]" captionTargetId-0="figure-17@45.[151,1435,181,1896]" captionTargetId-1="figure-26@46.[151,1435,181,1804]" captionTargetId-2="figure-17@47.[151,1435,181,1865]" captionTargetPageId-0="45" captionTargetPageId-1="46" captionTargetPageId-2="47" captionText-0="FIGURES 4AP. Monobazus species, habitus, dorsal and lateral view. AD, Monobazus confuscus (Pruthi), male (AB) and female (CD); EF, M. deccanensis sp. nov. holotype male; GJ, M. dissimilis (Distant), male (GH) and female (IJ); KL, M. himalayensis Distant, male; MN, M. longulus sp. nov., holotype male; OP, M. paulus sp. nov., paratype male." captionText-1="FIGURES 5AR. Monobazus group of genera, habitus, dorsal and lateral view. AB, Monobazus rotundus sp. nov., holotype male; CF, M. rubrostriatus sp. nov., holotype male (CD) and paratype female (EF); GJ, M. sahyadricus sp. nov., holotype male (GH) and paratype female (IJ); KN, M. sikkimensis sp. nov., holotype male (KL) and paratype female (MN); OP, M. sudindicus sp. nov., holotype male; QR, Panghoella sinuata gen. et sp. nov., holotype male." captionText-2="FIGURES 6AR. Monobazus group of genera, habitus, dorsal and lateral view. AD, Panghoella maculosa (Distant) male (AB) and female (CD); EF, Phlogotettix indicus Rao, male; GJ, P. subhimalayanus Meshram &amp; Ramamurthy, male (GH), and female (IJ); KL, Phlogothamnus maculiceps Ishihara, male; MN, P. umiamensis sp. nov. male; OR, Wanritettix wanrianus (Matsumura), male (OP), and female (QR)." figureDoi-0="http://doi.org/10.5281/zenodo.14703751" figureDoi-1="http://doi.org/10.5281/zenodo.14703753" figureDoi-2="http://doi.org/10.5281/zenodo.14703761" httpUri-0="https://zenodo.org/record/14703751/files/figure.png" httpUri-1="https://zenodo.org/record/14703753/files/figure.png" httpUri-2="https://zenodo.org/record/14703761/files/figure.png" pageId="104" pageNumber="105">Figs. 46</figureCitation>
;
<figureCitation id="1325F2D0F021FFF4FB0577A933512254" box="[1242,1308,255,279]" captionStart-0="FIGURES 20" captionStart-1="FIGURES 21" captionStart-2="FIGURES 22" captionStart-3="FIGURES 23" captionStart-4="FIGURES 24" captionStart-5="FIGURES 25" captionStart-6="FIGURES 26" captionStartId-0="61.[151,264,1942,1966]" captionStartId-1="62.[151,264,1943,1967]" captionStartId-2="63.[151,264,1816,1840]" captionStartId-3="64.[151,264,1909,1933]" captionStartId-4="65.[151,264,1910,1934]" captionStartId-5="66.[151,264,1914,1938]" captionStartId-6="67.[151,264,1710,1734]" captionTargetBox-0="[156,1430,186,1913]" captionTargetBox-1="[173,1414,181,1915]" captionTargetBox-2="[157,1430,186,1787]" captionTargetBox-3="[180,1408,186,1880]" captionTargetBox-4="[156,1430,186,1881]" captionTargetBox-5="[155,1431,186,1885]" captionTargetBox-6="[156,1430,186,1681]" captionTargetId-0="figure-17@61.[151,1435,181,1918]" captionTargetId-1="figure-26@62.[168,1420,181,1919]" captionTargetId-2="figure-17@63.[151,1435,181,1792]" captionTargetId-3="figure-26@64.[175,1413,181,1884]" captionTargetId-4="figure-17@65.[151,1435,181,1886]" captionTargetId-5="figure-26@66.[151,1435,181,1890]" captionTargetId-6="figure-17@67.[151,1435,181,1686]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionTargetPageId-2="63" captionTargetPageId-3="64" captionTargetPageId-4="65" captionTargetPageId-5="66" captionTargetPageId-6="67" captionText-0="FIGURES 20AJ. Cicurbanus serratus sp. nov., male genitalia: A, pygofer, lateral view; B, pygofer ventral process, lateral view; C, subgenital plate, ventral view; D, connective, dorsal view; E, style, dorsal view; F, aedeagus dorsal view; G, aedeagus anterodorsal view; H, aedeagal shaft apex, dorsal view; IJ, aedeagus, lateral views." captionText-1="FIGURES 21AI. Lankabanus attenuatus sp. nov., habitus and male genitalia: A, habitus, dorsal view; B, Face; C, genital capsule, lateral view; D, pygofer ventral process, lateral view; E, style, valve and subgenital plate, ventral view; F, connective, dorsal view; G, style, dorsal view; H, aedeagus, anteroventral view; I, aedeagus, lateral view." captionText-2="FIGURES 22 AH. Lankabanus bispinosus gen. et sp. nov.: AB, holotype male, habitus dorsal and lateral view; C, face; D, male genital capsule, lateral view; E, pygofer, ventral view showing ventral processes; F, valve, subgenital plate and style, dorsal view; G, style, connective and aedeagus dorsal view; H, aedeagus, and part of connective, lateral view." captionText-3="FIGURES 23AI. Madhyabus biligiris sp. nov., habitus and male genitalia: A, head and thorax, dorsal view; B, Face; C, male genital capsule, lateral view; D, pygofer and ventral processes, ventral view; E, pygofer ventral process, posterolateral view; F, valve and subgenital plate, ventral view; G, style, dorsal view; H, connective and aedeagus anterodorsal view; I, aedeagus, lateral view." captionText-4="FIGURES 24AG. Madhyabus reticulatus sp. nov., male genitalia: A, pygofer and anal segments, lateral view; B, valve and subgenital plate, ventral view; C, connective, dorsal view; D, style, dorsal view; E, aedeagus posterior view; F, aedeagus, dorsoposterior view; G, aedeagus, lateral view." captionText-5="FIGURES 25AF. Mavromoustaca brevispina sp. nov., male genitalia: A, pygofer, lateral view; B, subgenital plate, ventral view; C, connective, dorsal view; D, style, dorsal view; E, aedeagus posterior view; F, aedeagus, lateral view." captionText-6="FIGURES 26AF. Mavromoustaca channabasavanna sp. nov., male genitalia: A, pygofer, lateral view; B, subgenital plate, ventral view; C, style, dorsal view; D, style and aedeagus, dorsal view; E, aedeagus ventral view; F, connective and aedeagus, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.14703799" figureDoi-1="http://doi.org/10.5281/zenodo.14703803" figureDoi-2="http://doi.org/10.5281/zenodo.14703807" figureDoi-3="http://doi.org/10.5281/zenodo.14703813" figureDoi-4="http://doi.org/10.5281/zenodo.14703815" figureDoi-5="http://doi.org/10.5281/zenodo.14703819" figureDoi-6="http://doi.org/10.5281/zenodo.14703821" httpUri-0="https://zenodo.org/record/14703799/files/figure.png" httpUri-1="https://zenodo.org/record/14703803/files/figure.png" httpUri-2="https://zenodo.org/record/14703807/files/figure.png" httpUri-3="https://zenodo.org/record/14703813/files/figure.png" httpUri-4="https://zenodo.org/record/14703815/files/figure.png" httpUri-5="https://zenodo.org/record/14703819/files/figure.png" httpUri-6="https://zenodo.org/record/14703821/files/figure.png" pageId="104" pageNumber="105">2026</figureCitation>
: unavailable names.
</paragraph>
<paragraph id="8BA1EE55F021FFF4FF48766834B42216" blockId="104.[151,1436,255,341]" box="[151,761,317,341]" pageId="104" pageNumber="105">
<taxonomicName id="4C1E95D6F021FFF4FF487668379F2216" authority="Xing &amp; Li" authorityName="Xing &amp; Li" box="[151,466,317,341]" class="Insecta" family="Cicadellidae" genus="Longicornus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B96A3247F021FFF4FF487668372F2216" box="[151,354,317,341]" italics="true" pageId="104" pageNumber="105">Longicornus longus</emphasis>
Xing &amp; Li
</taxonomicName>
<emphasis id="B96A3247F021FFF4FE08766837A62216" box="[471,491,318,341]" italics="true" pageId="104" pageNumber="105">in</emphasis>
Dmitriev
<emphasis id="B96A3247F021FFF4FD8C766834CA2216" box="[595,647,317,341]" italics="true" pageId="104" pageNumber="105">et al.</emphasis>
2024: 275.
</paragraph>
<paragraph id="8BA1EE55F021FFF4FF4876D2349F214A" blockId="104.[151,1437,387,630]" pageId="104" pageNumber="105">
<emphasis id="B96A3247F021FFF4FF4876D2374422DE" box="[151,265,388,413]" italics="true" pageId="104" pageNumber="105">Diagnosis</emphasis>
.
<taxonomicName id="4C1E95D6F021FFF4FEC976D2373222DE" box="[278,383,387,413]" class="Insecta" family="Cicadellidae" genus="Longicornus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B96A3247F021FFF4FEC976D2373222DE" box="[278,383,387,413]" italics="true" pageId="104" pageNumber="105">L. longus</emphasis>
</taxonomicName>
is very similar to
<emphasis id="B96A3247F021FFF4FD9776D2354922DE" box="[584,772,387,413]" italics="true" pageId="104" pageNumber="105">L. flavipunctatus</emphasis>
Li &amp;Song but differs in the male genitalia characters. Male pygofer in
<taxonomicName id="4C1E95D6F021FFF4FECA76FE373A2282" box="[277,375,423,449]" class="Insecta" family="Cicadellidae" genus="Longicornus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="longus">
<emphasis id="B96A3247F021FFF4FECA76FE373A2282" box="[277,375,423,449]" italics="true" pageId="104" pageNumber="105">L longus</emphasis>
</taxonomicName>
is more deeply excavated (
<figureCitation id="1325F2D0F021FFF4FD7076F135592281" box="[687,788,423,450]" captionStart="FIGURES 56" captionStartId="97.[151,264,1860,1884]" captionTargetBox="[156,1417,203,1821]" captionTargetId="figure-17@97.[151,1435,198,1836]" captionTargetPageId="97" captionText="FIGURES 56AL. Longicornus longus (Xing &amp; Li): AB, male habitus, dorsal and lateral views; C, face; D, genital capsule, lateral view; E, pygofer, lateral view; F, valve and subgenital plates, ventral view; G, subgenital plate, lateral view; H, style, dorsal view; I, subgenital plates, styles, connective and aedeagus, dorsal view; J, connective and aedeagus, ventral view; K, style, connective and aedeagus, lateral view; L, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703893" httpUri="https://zenodo.org/record/14703893/files/figure.png" pageId="104" pageNumber="105">Fig. 56E</figureCitation>
) than in
<emphasis id="B96A3247F021FFF4FCA576FE32752282" box="[890,1080,423,449]" italics="true" pageId="104" pageNumber="105">L. flavipunctatus</emphasis>
and the apical processes of the aedeagal shaft are directed more or less in the direction of aedeagal shaft (
<figureCitation id="1325F2D0F021FFF4FC09769D326622A5" box="[982,1067,459,486]" captionStart="FIGURES 56" captionStartId="97.[151,264,1860,1884]" captionTargetBox="[156,1417,203,1821]" captionTargetId="figure-17@97.[151,1435,198,1836]" captionTargetPageId="97" captionText="FIGURES 56AL. Longicornus longus (Xing &amp; Li): AB, male habitus, dorsal and lateral views; C, face; D, genital capsule, lateral view; E, pygofer, lateral view; F, valve and subgenital plates, ventral view; G, subgenital plate, lateral view; H, style, dorsal view; I, subgenital plates, styles, connective and aedeagus, dorsal view; J, connective and aedeagus, ventral view; K, style, connective and aedeagus, lateral view; L, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703893" httpUri="https://zenodo.org/record/14703893/files/figure.png" pageId="104" pageNumber="105">Figs 56</figureCitation>
KL) whereas in
<emphasis id="B96A3247F021FFF4FB3F769A33D022A6" box="[1248,1437,459,485]" italics="true" pageId="104" pageNumber="105">L. flavipunctatus</emphasis>
they are curved ventrally and then directed basally.
</paragraph>
<paragraph id="8BA1EE55F021FFF4FF1875453305216E" blockId="104.[151,1437,387,630]" box="[199,1352,531,558]" pageId="104" pageNumber="105">
<emphasis id="B96A3247F021FFF4FF18754537D6216E" box="[199,411,531,557]" italics="true" pageId="104" pageNumber="105">Material examined</emphasis>
.
<materialsCitation id="3B76E408F021FFF4FE7675453309216E" box="[425,1348,531,558]" collectingDate="1934-05-20" collectionCode="BMNH" collectorName="N. E. Burma &amp; R. Malaise" country="Myanmar" elevation="2134" location="MYANMAR" pageId="104" pageNumber="105" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F309AEC5F021FFF4FE767545340D216E" box="[425,576,531,557]" name="Myanmar" pageId="104" pageNumber="105">MYANMAR</collectingCountry>
:
<specimenCount id="9D1825DCF021FFF4FD9575453422216E" box="[586,623,531,557]" count="1" pageId="104" pageNumber="105" type="male">1♂</specimenCount>
,
<collectorName id="26EB8B83F021FFF4FDA475423548216D" box="[635,773,532,558]" pageId="104" pageNumber="105">N.E. Burma</collectorName>
, Kambaiti,
<quantity id="4CE643B0F021FFF4FC597545359F216E" box="[902,978,531,557]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="104" pageNumber="105" unit="ft" value="7000.0">
<elevation id="00330966F021FFF4FC597545359F216E" box="[902,978,531,557]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="104" pageNumber="105" unit="ft" value="7000.0">7000ft</elevation>
</quantity>
,
<date id="FFA0C895F021FFF4FC0275453203216E" box="[989,1102,531,557]" pageId="104" pageNumber="105" value="1934-05-20">
<collectingDate id="EFE4317DF021FFF4FC0275453203216E" box="[989,1102,531,557]" pageId="104" pageNumber="105" value="1934-05-20">20.v.1934</collectingDate>
</date>
,
<collectorName id="26EB8B83F021FFF4FB867542329E216E" box="[1113,1235,531,558]" pageId="104" pageNumber="105">R. Malaiśe</collectorName>
(
<collectionCode id="ED0F7690F021FFF4FB3D75453371216E" box="[1250,1340,531,557]" pageId="104" pageNumber="105">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1EE55F021FFF4FF18756137702135" blockId="104.[151,1437,387,630]" pageId="104" pageNumber="105">
<emphasis id="B96A3247F021FFF4FF18756137642112" box="[199,297,567,593]" italics="true" pageId="104" pageNumber="105">Remarks</emphasis>
. Fang &amp; Xing (2018) illustrated the habitus and male genitalia of this species and also
<emphasis id="B96A3247F021FFF4FA59756E377B2136" italics="true" pageId="104" pageNumber="105">L. flavipunctatus</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>