treatments-xml/data/03/BA/87/03BA87C78169FFD16EFCFDC56FABFEAB.xml

<document id="DA64E15DAE1F08AA4BBA7190ED8BD449" ID-DOI="10.11646/phytotaxa.385.2.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13725833" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725674717613" checkinUser="felipe" docAuthor="Mamontov, Yuriy S., Vilnet, Anna A. &amp; Potemkin, Alexey D." docDate="2018" docId="03BA87C78169FFD16EFCFDC56FABFEAB" docLanguage="en" docName="phytotaxa.385.2.1.pdf" docOrigin="Phytotaxa 385 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.385.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Scapania marsupelloides Potemkin, Vilnet et Mamontov 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="62" masterDocId="FF83FFBF816DFFD66E74FFBE6F56FFE2" masterDocTitle="Scapania marsupelloides sp. nov. (Scapaniaceae, Marchantiophyta), a remarkable new species near the base of Scapania phylogeny" masterLastPageNumber="66" masterPageNumber="55" pageNumber="59" updateTime="1725687061590" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ECD5C8E64798908323C8D27E1A8ACAB6">Scapania marsupelloides sp. nov. (Scapaniaceae, Marchantiophyta), a remarkable new species near the base of Scapania phylogeny</mods:title>
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<mods:namePart id="0FB47AFA9A451C3B0CAF28F84ED2D767">Mamontov, Yuriy S.</mods:namePart>
<mods:affiliation id="83B736CFAF44C7825663DC914776595C">Botanical Garden-Institute, Russian Academy of Sciences, Makovskogo 142, Vladivostok, 690024, Russia; &amp; Tsitsin Main Botanical Garden, Russian Academy of Sciences, Botanicheskaya 4, Moscow 127276, Russia;</mods:affiliation>
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<mods:namePart id="EFF155A2C31C29122F6789ED0C8DAEE1">Vilnet, Anna A.</mods:namePart>
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<mods:namePart id="78A24575F54AD0E5D33531F1B67A7D1A">Potemkin, Alexey D.</mods:namePart>
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<mods:date id="CE1B29D2B0E7769C190014F70780F52D">2018</mods:date>
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<treatment id="03BA87C78169FFD16EFCFDC56FABFEAB" LSID="urn:lsid:plazi:treatment:03BA87C78169FFD16EFCFDC56FABFEAB" httpUri="http://treatment.plazi.org/id/03BA87C78169FFD16EFCFDC56FABFEAB" lastPageId="7" lastPageNumber="62" pageId="4" pageNumber="59">
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<heading id="D0E481BD8169FFD26EFCFDC56B72FD74" bold="true" box="[136,1060,635,662]" fontSize="11" level="1" pageId="4" pageNumber="59" reason="1">
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<taxonomicName id="4C134D528169FFD26EFCFDC56C49FD74" ID-CoL="79RXF" authority="Potemkin, Vilnet et Mamontov" authorityName="Potemkin, Vilnet et Mamontov" authorityYear="2018" box="[136,799,635,662]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="4" pageNumber="59" phylum="Marchantiophyta" rank="species" species="marsupelloides" status="sp. nov.">
<emphasis id="B967EAC38169FFD26EFCFDC56EFEFD74" bold="true" box="[136,424,635,662]" italics="true" pageId="4" pageNumber="59">Scapania marsupelloides</emphasis>
Potemkin, Vilnet et Mamontov
</taxonomicName>
, 
<emphasis id="B967EAC38169FFD26D5FFDC26CD4FD74" bold="true" box="[811,898,636,662]" italics="true" pageId="4" pageNumber="59">
<taxonomicNameLabel id="A25457B88169FFD26D5FFDC26CD4FD74" box="[811,898,636,662]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="13282A548169FFD26DFDFDC36C87FD74" box="[905,977,637,662]" captionStart="FIGURE 2" captionStartId="3.[136,229,1370,1391]" captionTargetBox="[142,1443,189,1330]" captionTargetId="figure-14@3.[138,1448,187,1342]" captionTargetPageId="3" captionText="FIGURE 2. Line drawing of Scapania marsupelloides sp. nov. 1, 2, 4—Shoots. 3—Cells of the leaf middle, showing papillose surface. 5—Stem cross section. 6–11—Leaves. Scale bars: 3 mm for 1; 4 mm for 2; 100 μm for 3; 500 μm for 5; 5 mm for 4; 1 mm for 6–11. All from D. G. Long &amp; J. Shevock 37350 (isotype, MHA)." figureDoi="http://doi.org/10.5281/zenodo.13725837" httpUri="https://zenodo.org/record/13725837/files/figure.png" pageId="4" pageNumber="59">Figs 2</figureCitation>
, 
<figureCitation id="13282A548169FFD26DAAFDC36CBBFD74" box="[990,1005,637,662]" captionStart="FIGURE 3" captionStartId="5.[136,229,1754,1775]" captionTargetBox="[152,1431,196,1720]" captionTargetId="figure-14@5.[151,1436,190,1729]" captionTargetPageId="5" captionText="FIGURE 3. Line drawing of Scapania marsupelloides sp. nov.1, 2, 3, 17—Shoot sectors (1—postical aspect; 2—postical aspect with long decurrent ventral lobe; 3, 17—antical aspect). 4—Stem cross section. 5—Leaf. 6–9, 10–12, 14—Leaf cross sections. 13, 15, 16—Male bracts. Scale bars: 3 mm for 1, 3, 17; 2 mm for 2; 1.5 mm for 5, 13, 15, 16; 500 мm for 4; 300 мm for 6, 9, 10, 12, 14; 250 мm for 7, 8, 11. All from D. G. Long &amp; J. Shevock 37350 (isotype, MHA)." figureDoi="http://doi.org/10.5281/zenodo.13725839" httpUri="https://zenodo.org/record/13725839/files/figure.png" pageId="4" pageNumber="59">3</figureCitation>
, 
<figureCitation id="13282A548169FFD26D8EFDC36B5FFD74" box="[1018,1033,637,662]" captionStart="FIGURE 4" captionStartId="6.[136,229,1885,1906]" captionTargetBox="[158,1430,245,1860]" captionTargetId="figure-58@6.[158,1430,245,1860]" captionTargetPageId="6" captionText="FIGURE 4. Photomicrographs of Scapania marsupelloides sp. nov. 1–Stem cross section. 2, 3, 5, 6—Leaf apices. 4, 11—Marginal leaf cells. 7—Cells of leaf base. 8–10—Cells of leaf middle. Scale bars: 1, 2, 3, 5, 6 = 100 μm. 4, 7, 8, 10 = 50 μm. 9, 11= 25 μm. All from D. G. Long &amp; J. Shevock 37350 (isotype, MHA)." figureDoi="http://doi.org/10.5281/zenodo.13725841" httpUri="https://zenodo.org/record/13725841/files/figure.png" pageId="4" pageNumber="59">4</figureCitation>
, 
<figureCitation id="13282A548169FFD26A62FDC36B72FD74" box="[1046,1060,637,662]" captionStart="FIGURE 5" captionStartId="8.[136,229,1740,1761]" captionTargetBox="[151,1435,245,1715]" captionTargetId="figure-43@8.[151,1436,245,1715]" captionTargetPageId="8" captionText="FIGURE 5. SEM images of Scapania marsupelloides sp. nov. 1–3, 5, 6—Cells of leaf middle showing papillose surface. 4—Cells of leaf base. 7—Cells of stem cross section. Scale bars: 1, 4 = 30 μm. 2 = 10 μm. 3, 5 = 40 μm. 6 = 200 μm. 7 = 20 μm. All from D. G. Long &amp; J. Shevock 37350 (isotype, LE)." figureDoi="http://doi.org/10.5281/zenodo.13725843" httpUri="https://zenodo.org/record/13725843/files/figure.png" pageId="4" pageNumber="59">5</figureCitation>
</emphasis>
</heading>
</paragraph>
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<emphasis id="B967EAC38169FFD26EFCFD1D6F96FD5A" bold="true" box="[136,192,675,696]" pageId="4" pageNumber="59">Type.</emphasis>
— 
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<collectingCountry id="F30476418169FFD26EADFD1D6E71FD5A" box="[217,295,675,696]" name="China" pageId="4" pageNumber="59">CHINA</collectingCountry>
. 
<collectingRegion id="49D7F8338169FFD26F45FD1D6E28FD5A" box="[305,382,675,696]" country="China" name="Yunnan" pageId="4" pageNumber="59">Yunnan</collectingRegion>
: 
<collectingCounty id="62CD4E5D8169FFD26FFCFD1D6D76FD5A" box="[392,544,675,696]" pageId="4" pageNumber="59">Fugong County</collectingCounty>
, 
<collectingMunicipality id="6BC8ACAB8169FFD26C5EFD1D6D83FD5A" box="[554,725,675,696]" pageId="4" pageNumber="59">Lumadeng Xiang</collectingMunicipality>
, 
<location id="8ECC600A8169FFD26CABFD1D6C03FD5A" LSID="urn:lsid:plazi:treatment:03BA87C78169FFD16EFCFDC56FABFEAB:8ECC600A8169FFD26CABFD1D6C03FD5A" box="[735,853,675,696]" country="China" county="Fugong County" latitude="27.200306" longLatPrecision="1" longitude="98.69447" municipality="Lumadeng Xiang" name="Yaping Cun" pageId="4" pageNumber="59" stateProvince="Yunnan">Yaping Cun</location>
, 
<collectionCode id="ED02AE148169FFD26D14FD1D6C39FD5A" box="[864,879,675,696]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="4" pageNumber="59" type="Herbarium">E</collectionCode>
slope of 
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(Nu Jiang catchment), east slope of 
<location id="8ECC600A8169FFD26EACFD796E4EFD3E" LSID="urn:lsid:plazi:treatment:03BA87C78169FFD16EFCFDC56FABFEAB:8ECC600A8169FFD26EACFD796E4EFD3E" box="[216,280,711,732]" country="China" county="Fugong County" latitude="27.200306" longLatPrecision="1" longitude="98.69447" municipality="Lumadeng Xiang" name="Burma" pageId="4" pageNumber="59" stateProvince="Yunnan">Burma</location>
/ 
<collectingRegion id="49D7F8338169FFD26F6BFD796E3FFD3E" box="[287,361,711,732]" country="China" name="Yunnan" pageId="4" pageNumber="59">Yunnan</collectingRegion>
border ridge beside ‘ 
<location id="8ECC600A8169FFD26C47FD796DF9FD3E" LSID="urn:lsid:plazi:treatment:03BA87C78169FFD16EFCFDC56FABFEAB:8ECC600A8169FFD26C47FD796DF9FD3E" box="[563,687,711,732]" country="China" county="Fugong County" latitude="27.200306" longLatPrecision="1" longitude="98.69447" municipality="Lumadeng Xiang" name="Twin Lakes'" pageId="4" pageNumber="59" stateProvince="Yunnan">Twin Lakes’</location>
south of ‘ 
<location id="8ECC600A8169FFD26D79FD796CC7FD3E" LSID="urn:lsid:plazi:treatment:03BA87C78169FFD16EFCFDC56FABFEAB:8ECC600A8169FFD26D79FD796CC7FD3E" box="[781,913,711,732]" country="China" county="Fugong County" latitude="27.200306" longLatPrecision="1" longitude="98.69447" municipality="Lumadeng Xiang" name="Yaping Pass'" pageId="4" pageNumber="59" stateProvince="Yunnan">Yaping Pass’</location>
, ca. 
<elevation id="003ED1E28169FFD26DC9FD796B10FD3E" box="[957,1094,711,732]" metricMagnitude="3" metricUnit="m" metricValue="3.61" pageId="4" pageNumber="59" unit="m" value="3610.0">
<quantity id="4CEB9B348169FFD26DC9FD796B50FD3E" box="[957,1030,711,732]" metricMagnitude="3" metricUnit="m" metricValue="3.61" pageId="4" pageNumber="59" unit="m" value="3610.0">3610 m</quantity>
a. s. l.
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, 
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, 
<geoCoordinate id="EE2750168169FFD26A97FD796A31FD3E" box="[1251,1383,711,732]" degrees="98" direction="east" minutes="41" orientation="longitude" pageId="4" pageNumber="59" precision="1" seconds="40.1" value="98.69447">98°41′40.1″E</geoCoordinate>
, alpine lake shore with marshy slopes, boulders and tall herbs
</materialsCitation>
; 
<materialsCitation id="3B7B3C8C8169FFD26CAFFD556CE5FCC6" collectingDate="2007-10-05" collectionCode="G, J, LE, MHA, KPABG, VBGI" country="China" pageId="4" pageNumber="59" specimenCount="2" typeStatus="holotype">
on flushed slope, 
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<collectingDate id="EFE9E9F98169FFD26DFCFD556B74FCE2" box="[904,1058,747,768]" pageId="4" pageNumber="59" value="2007-10-05">5 October 2007</collectingDate>
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, 
<emphasis id="B967EAC38169FFD26A59FD556A30FCE2" box="[1069,1382,747,768]" italics="true" pageId="4" pageNumber="59">
D. 
<collectionCode id="ED02AE148169FFD26A3FFD556B09FCE2" box="[1099,1119,747,768]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="4" pageNumber="59" type="Herbarium">G</collectionCode>
. Long &amp; 
<collectionCode id="ED02AE148169FFD26AC8FD556B9CFCE2" box="[1212,1226,747,768]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="4" pageNumber="59" type="Herbarium">J</collectionCode>
. Shevock 37350
</emphasis>
det. as 
<taxonomicName id="4C134D528169FFD26EC9FCB16E9BFCC6" authority=", MO" authorityName="MO" box="[189,461,783,804]" class="Jungermanniopsida" family="Gymnomitriaceae" genus="Marsupella" kingdom="Plantae" order="Jungermanniales" pageId="4" pageNumber="59" phylum="Marchantiophyta" rank="species" species="sphacelata">
<emphasis id="B967EAC38169FFD26EC9FCB16ECCFCC6" box="[189,410,783,804]" italics="true" pageId="4" pageNumber="59">Marsupella sphacelata</emphasis>
, MO
</taxonomicName>
(
<typeStatus id="54A888738169FFD26FADFCB16D61FCC6" box="[473,567,783,804]" pageId="4" pageNumber="59" type="holotype">Holotype</typeStatus>
), 
<collectionCode id="ED02AE148169FFD26C3CFCB16D3EFCC6" box="[584,616,783,804]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="4" pageNumber="59">LE</collectionCode>
, 
<collectionCode id="ED02AE148169FFD26C05FCB16DFDFCC6" box="[625,683,783,804]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15585" lsid="urn:lsid:biocol.org:col:15585" name="Main Botanical Garden of the Russian Academy of Sciences" pageId="4" pageNumber="59" type="Herbarium">MHA</collectionCode>
, 
<collectionCode id="ED02AE148169FFD26CC1FCB16C51FCC6" box="[693,775,783,804]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15236" lsid="urn:lsid:biocol.org:col:15236" name="Polar-Alpine Botanical Garden-Institute" pageId="4" pageNumber="59" type="Herbarium">KPABG</collectionCode>
, 
<collectionCode id="ED02AE148169FFD26D64FCB16C1DFCC6" box="[784,843,783,804]" country="Russia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15212" lsid="urn:lsid:biocol.org:col:15212" name="Botanical Garden-Institute of FEB RAS" pageId="4" pageNumber="59" type="Herbarium">VBGI</collectionCode>
(
<typeStatus id="54A888738169FFD26D2CFCB16CFAFCC6" box="[856,940,783,804]" pageId="4" pageNumber="59" type="isotype">isotypes</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309655A8169FFD26EFCFC8D6A07FC1E" pageId="4" pageNumber="59" type="diagnosis">
<paragraph id="8BAC36D18169FFD26EFCFC8D6A07FC1E" blockId="4.[136,1452,635,1020]" pageId="4" pageNumber="59">
<emphasis id="B967EAC38169FFD26EFCFC8D6E46FCAA" bold="true" box="[136,272,819,840]" pageId="4" pageNumber="59">Diagnosis:—</emphasis>
Brownish green to yellowish brown 
<taxonomicName id="4C134D528169FFD26C0BFC8D6C33FCAA" box="[639,869,819,840]" class="Jungermanniopsida" family="Gymnomitriaceae" genus="Marsupella" kingdom="Plantae" order="Jungermanniales" pageId="4" pageNumber="59" phylum="Marchantiophyta" rank="species" species="sphacelata">
<emphasis id="B967EAC38169FFD26C0BFC8D6C33FCAA" box="[639,869,819,840]" italics="true" pageId="4" pageNumber="59">Marsupella sphacelata</emphasis>
</taxonomicName>
-like plants with coarsely papillose leaf surface in median part of lobes. The species furthermore characterized by 1–2-stratose cortex with indistinct and only locally expressed areas of ± larger ± thick-walled to thin-walled cells situated often side by side with smaller cells or in one part of stem (
<figureCitation id="13282A548169FFD26A8BFCC56A13FC72" box="[1279,1349,891,912]" captionStart="FIGURE 2" captionStartId="3.[136,229,1370,1391]" captionTargetBox="[142,1443,189,1330]" captionTargetId="figure-14@3.[138,1448,187,1342]" captionTargetPageId="3" captionText="FIGURE 2. Line drawing of Scapania marsupelloides sp. nov. 1, 2, 4—Shoots. 3—Cells of the leaf middle, showing papillose surface. 5—Stem cross section. 6–11—Leaves. Scale bars: 3 mm for 1; 4 mm for 2; 100 μm for 3; 500 μm for 5; 5 mm for 4; 1 mm for 6–11. All from D. G. Long &amp; J. Shevock 37350 (isotype, MHA)." figureDoi="http://doi.org/10.5281/zenodo.13725837" httpUri="https://zenodo.org/record/13725837/files/figure.png" pageId="4" pageNumber="59">Figs. 2</figureCitation>
: 
<emphasis id="B967EAC38169FFD26B3BFCC56A0DFC72" box="[1359,1371,891,912]" italics="true" pageId="4" pageNumber="59">5</emphasis>
, 3: 
<emphasis id="B967EAC38169FFD26BF4FCC56ADAFC72" box="[1408,1420,891,912]" italics="true" pageId="4" pageNumber="59">4</emphasis>
, 4: 
<emphasis id="B967EAC38169FFD26EC9FC216F9FFC56" box="[189,201,927,948]" italics="true" pageId="4" pageNumber="59">1</emphasis>
). The leaves are sheathing the stem and not keeled, 0.5–0.6 mostly equally bilobed with oval, ovate or lingulate falcate lobes, rounded to obtuse or rarely angulate at apex, spreading from each other, ± arcuately inserted and ± decurrent on both dorsal and ventral sides. The marginal leaf cells usually darker brown pigmented, ± evenly thick-walled, without distinct trigones.
</paragraph>
</subSubSection>
<subSubSection id="C309655A8169FFD16EFCFB936FABFEAB" lastPageId="7" lastPageNumber="62" pageId="4" pageNumber="59" type="description">
<paragraph id="8BAC36D18169FFD26EFCFB936A48F844" blockId="4.[136,1452,1069,1958]" pageId="4" pageNumber="59">
<emphasis id="B967EAC38169FFD26EFCFB936E4AFBA4" bold="true" box="[136,284,1069,1094]" pageId="4" pageNumber="59">Description:</emphasis>
—Plants brownish green to yellowish brown, robust, 
<quantity id="4CEB9B348169FFD26DD5FB936B77FBA4" box="[929,1057,1069,1094]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="4" pageNumber="59" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
long and (1.5–) 
<quantity id="4CEB9B348169FFD26A91FB936A37FBA4" box="[1253,1377,1069,1094]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="59" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
wide, sporadically with innovations from broken shoot tips, rare with microphyllous terminal branches, apparently 
<taxonomicName id="4C134D528169FFD26B25FBEF6AF4FB88" box="[1361,1442,1105,1130]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="4" pageNumber="59" phylum="Marchantiophyta" rank="genus">
<emphasis id="B967EAC38169FFD26B25FBEF6AF4FB88" box="[1361,1442,1105,1130]" italics="true" pageId="4" pageNumber="59">Radula</emphasis>
</taxonomicName>
- 
<typeStatus id="54A888738169FFD26EFCFBCB6FEDFB6C" box="[136,187,1141,1166]" pageId="4" pageNumber="59">type</typeStatus>
, originating from postical shoot sectors not in leaf axils but below upper leaf insertion. Stem 
<quantity id="4CEB9B348169FFD26A81FBCB6ADBFB6C" box="[1269,1421,1141,1166]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="4" pageNumber="59" unit="mm" value="275.0" valueMax="300.0" valueMin="250.0">250–300 mm</quantity>
in diam., deep brown, reddish-brown or brownish green, in upper part sometimes green. Cortex 1–2-stratose, with outer cortical cells partially larger than internal, moderately thick-walled to thin-walled, ± finely papillose when observed in cross section, with finely grooved surface in SEM images; middle lamellae in cortical cells invisible to ± distinct. Leaves 0.5–0.6 bilobed, broadly sheathing the stem, hardly broadened to the base; keel lacking or hardly developed distally, lobes oval, ovate or lingulate falcate, often with subparallel margins, rounded to obtuse or rarely angulate at apex, reaching but usually not extending beyond the farther edge of stem, entire or very rare with single blunt marginal tooth-like projections of subisodiametric cells, slightly arcuately inserted on dorsal and ventral sides, with indistinct short to rather long decurrency below the level of keel insertion; leaf insertion often indistinct on dark stem. Dorsal lobe mostly equal in area the ventral lobe or a bit smaller, identical to it in shape, ca. × 0.6–0.7(–0.8) as wide as long, ca. 625–1125 μm wide × 1000–1625 μm long, similarly falcate or moderately divergent with it, obtuse or rarely angulate at apex, entire or with a few tooth-like marginal angulations of subisodiametric cells. Leaves distinctly bordered with 1(–2) rows of evenly thick-walled mostly subquadrate marginal cells with yellowish or brownish almost always deeply pigmented walls, sometimes slightly elongated perpendicular to margin, (8–)10–15(–20) × 7–12(–15) μm; only some basal marginal cells elongated along margin; marginal area deeper brown pigmented than median part of leaves. Median lobe cells with yellow walls, (12–)17–20(–25) × 20–25 μm or 25–27 × 25–37 μm, usually with acute to sometimes ± bulging trigones, in mod. 
<emphasis id="B967EAC38169FFD26C01F90B6C13F92C" box="[629,837,1717,1742]" italics="true" pageId="4" pageNumber="59">leptoderma -viridis</emphasis>
trigones may be vestigial. Median cells below sinus occasionally strongly elongated, 20–27 × 50–70(–80) μm, with acute to bulging elongated trigones and yellowish walls. Basal cells elongated at median part of leaf base close to the place of its insertion, (15–)17(–20) × (35–)40– 60(–75) μm, thin- to ± evenly thin-walled, with slighter developed papillae. Cell surface irregularly densely coarsely papillose in distal part of leaves with chiefly hemispherical and partly elongated papillae in median part of lobes and often minute papillae near margins; cell surface in proximal part of leaves below sinus moderately papillose, with elongated papillae; leaf margins smooth or sometimes with distinct flattened papillae. Gemmae not seen.
</paragraph>
<caption id="DF6C66598168FFD36EFCF9646DECF8B9" ID-DOI="http://doi.org/10.5281/zenodo.13725839" ID-Zenodo-Dep="13725839" httpUri="https://zenodo.org/record/13725839/files/figure.png" pageId="5" pageNumber="60" startId="5.[136,229,1754,1775]" targetBox="[152,1431,196,1720]" targetPageId="5" targetType="figure">
<paragraph id="8BAC36D18168FFD36EFCF9646DECF8B9" blockId="5.[136,1452,1754,1883]" pageId="5" pageNumber="60">
<emphasis id="B967EAC38168FFD36EFCF9646FAAF90D" bold="true" box="[136,252,1754,1775]" pageId="5" pageNumber="60">FIGURE 3.</emphasis>
Line drawing of 
<emphasis id="B967EAC38168FFD36FEBF9646D8BF90D" box="[415,733,1754,1775]" italics="true" pageId="5" pageNumber="60">
<taxonomicName id="4C134D528168FFD36FEBF9646DC6F90D" authorityName="Mamontov &amp; Vilnet &amp; Potemkin" authorityYear="2018" box="[415,656,1754,1775]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="5" pageNumber="60" phylum="Marchantiophyta" rank="species" species="marsupelloides" status="sp. nov.">Scapania marsupelloides</taxonomicName>
<taxonomicNameLabel id="A25457B88168FFD36CE0F9646D8BF90D" box="[660,733,1754,1775]" pageId="5" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1, 2, 3, 17—Shoot sectors (1—postical aspect; 2—postical aspect with long decurrent ventral lobe; 3, 17—antical aspect). 4—Stem cross section. 5—Leaf. 6–9, 10–12, 14—Leaf cross sections. 13, 15, 16—Male bracts. Scale bars: 3 mm for 1, 3, 17; 2 mm for 2; 1.5 mm for 5, 13, 15, 16; 500 мm for 4; 300 мm for 6, 9, 10, 12, 14; 250 мm for 7, 8, 11. All from 
<emphasis id="B967EAC38168FFD36E95F8F86D45F8B9" box="[225,531,1862,1883]" italics="true" pageId="5" pageNumber="60">D. G. Long &amp; J. Shevock 37350</emphasis>
(isotype, MHA).
</paragraph>
</caption>
<paragraph id="8BAC36D18168FFD36EC9F8356AFAF7F2" blockId="5.[136,1452,1931,2064]" pageId="5" pageNumber="60">Dioicous. Androecia intercalary of 3–5 pairs of male bracts strongly concave basally, with three distinct cavities, broadly ovate when flattened, with maximal width near the base, sometimes partly purplish in the middle; sinuses shallower than in sterile leaves, ca. 0.35–0.5 × the length; lobes broadly rounded or narrowed to the apex, 0.64–0.94 × as wide as long, ca. 700–1050 μm wide and 1100–1175 μm long; marginal cells occasionally bleached in apical area.</paragraph>
<paragraph id="8BAC36D1816BFFD06EFCFF1E6BEEFF3F" blockId="6.[136,1451,160,221]" pageId="6" pageNumber="61">Antheridia in groups in the cavities, with 1(–2)–seriate stalks, 17–20 μm wide, 50–150 μm long, of ± flattened thin-walled cells; mature body oval, ca. 150 × 125 μm. Paraphyses not seen. Female plants not seen.</paragraph>
<caption id="DF6C6659816BFFD06EFCF8E36D36F858" ID-DOI="http://doi.org/10.5281/zenodo.13725841" ID-Zenodo-Dep="13725841" httpUri="https://zenodo.org/record/13725841/files/figure.png" pageId="6" pageNumber="61" startId="6.[136,229,1885,1906]" targetBox="[158,1430,245,1860]" targetPageId="6" targetType="figure">
<paragraph id="8BAC36D1816BFFD06EFCF8E36D36F858" blockId="6.[136,1452,1885,1978]" pageId="6" pageNumber="61">
<emphasis id="B967EAC3816BFFD06EFCF8E36FA8F890" bold="true" box="[136,254,1885,1906]" pageId="6" pageNumber="61">FIGURE 4.</emphasis>
Photomicrographs of 
<emphasis id="B967EAC3816BFFD06FA2F8E36C4CF890" box="[470,794,1885,1906]" italics="true" pageId="6" pageNumber="61">
<taxonomicName id="4C134D52816BFFD06FA2F8E36D9FF890" authorityName="Mamontov &amp; Vilnet &amp; Potemkin" authorityYear="2018" box="[470,713,1885,1906]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="6" pageNumber="61" phylum="Marchantiophyta" rank="species" species="marsupelloides" status="sp. nov.">Scapania marsupelloides</taxonomicName>
<taxonomicNameLabel id="A25457B8816BFFD06CA4F8E36C4CF890" box="[720,794,1885,1906]" pageId="6" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1–Stem cross section. 2, 3, 5, 6—Leaf apices. 4, 11—Marginal leaf cells. 7—Cells of leaf base. 8–10—Cells of leaf middle. Scale bars: 1, 2, 3, 5, 6 = 100 μm. 4, 7, 8, 10 = 50 μm. 9, 11= 25 μm. All from 
<emphasis id="B967EAC3816BFFD06EFCF81B6E2EF858" box="[136,376,1957,1978]" italics="true" pageId="6" pageNumber="61">D. G. Long &amp; J. Shevock</emphasis>
37350 (isotype, MHA).
</paragraph>
</caption>
<paragraph id="8BAC36D1816BFFD06EC9F8556BEBF7CA" blockId="6.[189,1327,2027,2088]" pageId="6" pageNumber="61">
<emphasis id="B967EAC3816BFFD06EC9F8556E01F7E6" bold="true" box="[189,343,2027,2052]" pageId="6" pageNumber="61">Ethymology:</emphasis>
—The species name originates from its habitual resemblance to 
<taxonomicName id="4C134D52816BFFD06A53F8556A7EF7E6" box="[1063,1320,2027,2052]" class="Jungermanniopsida" family="Gymnomitriaceae" genus="Marsupella" kingdom="Plantae" order="Jungermanniales" pageId="6" pageNumber="61" phylum="Marchantiophyta" rank="species" species="sphacelata">
<emphasis id="B967EAC3816BFFD06A53F8556A7EF7E6" box="[1063,1320,2027,2052]" italics="true" pageId="6" pageNumber="61">Marsupella sphacelata</emphasis>
</taxonomicName>
. 
<emphasis id="B967EAC3816BFFD06EC9F7B16E74F7CA" bold="true" box="[189,290,2063,2088]" pageId="6" pageNumber="61">Ecology.</emphasis>
—Alpine lake shore with marshy slopes, boulders and tall herbs; on flushed slope.
</paragraph>
<paragraph id="8BAC36D1816AFFD16EC9FF1E6FABFEAB" blockId="7.[136,1452,160,329]" pageId="7" pageNumber="62">
<emphasis id="B967EAC3816AFFD16EC9FF1E6EC5FF5B" bold="true" box="[189,403,160,185]" pageId="7" pageNumber="62">Differentiation:—</emphasis>
<emphasis id="B967EAC3816AFFD16FE7FF1E6D3CFF5B" box="[403,618,160,185]" italics="true" pageId="7" pageNumber="62">
<taxonomicName id="4C134D52816AFFD16FE7FF1E6D31FF5B" authorityName="Mitten" authorityYear="1860" box="[403,615,160,185]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="contorta">Scapania contorta</taxonomicName>
,
</emphasis>
<taxonomicName id="4C134D52816AFFD16C07FF1E6B65FF5B" authority="Potemkin (2000: 115)" authorityName="Potemkin" authorityPageNumber="115" authorityYear="2000" box="[627,1075,160,185]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="pseudocotorta">
<emphasis id="B967EAC3816AFFD16C07FF1E6C67FF5B" box="[627,817,160,185]" italics="true" pageId="7" pageNumber="62">S. pseudocotorta</emphasis>
<bibRefCitation id="EF824B20816AFFD16D4EFF1E6B65FF5B" author="Potemkin, A. D." box="[826,1075,160,185]" pageId="7" pageNumber="62" pagination="115 - 122" refId="ref6685" refString="Potemkin, A. D. (2000) Three new species of Scapania (Hepaticae) collected by Dr. David G. Long in Nepal. Arctoa 9: 115 - 122. https: // doi. org / 10.15298 / arctoa. 09.12" type="journal article" year="2000">Potemkin (2000: 115)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C134D52816AFFD16A19FF1E6E58FF3F" authority="Stephani ex Muller (1905: 223)" authorityName="Stephani ex Muller" authorityPageNumber="223" authorityYear="1905" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="hians">
<emphasis id="B967EAC3816AFFD16A19FF1E6B9EFF5B" box="[1133,1224,160,185]" italics="true" pageId="7" pageNumber="62">S. hians</emphasis>
Stephani ex 
<bibRefCitation id="EF824B20816AFFD16B2BFF1E6E58FF3F" author="Muller, K." pageId="7" pageNumber="62" pagination="1 - 312" refId="ref6472" refString="Muller, K. (1905) Monographie der Lebermoosgattung Scapania Dum. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum 83: 1 - 312." type="journal article" year="1905">Müller (1905: 223)</bibRefCitation>
</taxonomicName>
of the subgenus 
<taxonomicName id="4C134D52816AFFD16FA2FF7A6D1AFF3F" box="[470,588,196,221]" pageId="7" pageNumber="62" rank="subGenus" subGenus="Ascapania">
<emphasis id="B967EAC3816AFFD16FA2FF7A6D1AFF3F" box="[470,588,196,221]" italics="true" pageId="7" pageNumber="62">Ascapania</emphasis>
</taxonomicName>
as well as 
<taxonomicName id="4C134D52816AFFD16CA5FF7A6CDCFF3F" box="[721,906,196,221]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="cuspiduligera">
<emphasis id="B967EAC3816AFFD16CA5FF7A6CDCFF3F" box="[721,906,196,221]" italics="true" pageId="7" pageNumber="62">S. cuspiduligera</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF824B20816AFFD16DEFFF7A6B1CFF3F" author="Nees, C. G." box="[923,1098,196,221]" pageId="7" pageNumber="62" refId="ref6551" refString="Nees, C. G. (1833) Naturgeschichte der Europaischen Lebermoose, vol. 1. August Rucker, Berlin, 347 pp." type="book" year="1833">Nees1833: 175</bibRefCitation>
, 180, 181) 
<bibRefCitation id="EF824B20816AFFD16AA4FF7A6AFAFF3F" author="Muller, K." box="[1232,1452,196,221]" pageId="7" pageNumber="62" pagination="465 - 528" refId="ref6499" refString="Muller, K. (1915) Die Lebermoose. In: Dr. L. Rabenhorst's Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aufl., 6 Band), 2 Abth., 22 Lieferung. Eduard Kummer, Leipzig, pp. 465 - 528." type="book chapter" year="1915">Müller (1915: 472)</bibRefCitation>
(except for its 
<taxonomicName id="4C134D52816AFFD16F5EFF566DA1FEE3" authority="Schuster (1974: 361)" authorityName="Schuster" authorityPageNumber="361" authorityYear="1974" box="[298,759,232,257]" pageId="7" pageNumber="62" rank="variety" variety="diplophyllopsis">
var. 
<emphasis id="B967EAC3816AFFD16F2EFF566D53FEE3" box="[346,517,232,257]" italics="true" pageId="7" pageNumber="62">diplophyllopsis</emphasis>
<bibRefCitation id="EF824B20816AFFD16C78FF566DA1FEE3" author="Schuster, R. M." box="[524,759,232,257]" pageId="7" pageNumber="62" refId="ref7042" refString="Schuster, R. M. (1974) The Hepaticae and Anthocerotae of North America east of the hundredth meridian. Vol. III. Columbia University Press, New York, London, 880 + ix pp." type="book" year="1974">Schuster (1974: 361)</bibRefCitation>
</taxonomicName>
that has a typical 
<emphasis id="B967EAC3816AFFD16DB6FF566B6BFEE3" box="[962,1085,232,257]" italics="true" pageId="7" pageNumber="62">scapanioid</emphasis>
habit) are morphologically most close to 
<taxonomicName id="4C134D52816AFFD16E97FEB26EFEFEC7" authorityName="Mamontov &amp; Vilnet &amp; Potemkin" authorityYear="2018" box="[227,424,268,293]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="marsupelloides">
<emphasis id="B967EAC3816AFFD16E97FEB26EFEFEC7" box="[227,424,268,293]" italics="true" pageId="7" pageNumber="62">S. marsupelloides</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C134D52816AFFD16FC1FEB26D2DFEC7" authorityName="Mamontov &amp; Vilnet &amp; Potemkin" authorityYear="2018" box="[437,635,268,293]" class="Jungermanniopsida" family="Scapaniaceae" genus="Scapania" kingdom="Plantae" order="Jungermanniales" pageId="7" pageNumber="62" phylum="Marchantiophyta" rank="species" species="marsupelloides">
<emphasis id="B967EAC3816AFFD16FC1FEB26D2DFEC7" box="[437,635,268,293]" italics="true" pageId="7" pageNumber="62">S. marsupelloides</emphasis>
</taxonomicName>
shares with them subequal lobes spreading from each other and sheathing leaf bases.
</paragraph>
</subSubSection>
</treatment>
</document>